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<document id="0CF8FBCB3918BEF1FFD2F3BEE2B9315A" ID-CLB-Dataset="295393" ID-DOI="10.5281/zenodo.11105986" ID-GBIF-Dataset="4c7b4d3d-1fbe-4f48-8cb2-002e928eae42" ID-ISSN="0068-547X" ID-Zenodo-Dep="11105986" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1714051164034" checkinUser="felipe" docAuthor="Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Castroviejo-Fisher, Santiago &amp; De, Ignacio" docDate="2020" docId="03B1733AFFE1A145FFFBFC99ED27FAE1" docLanguage="en" docName="ProcCalifAcadSci.66.8.137-230.pdf" docOrigin="Proceedings of the California Academy of Sciences 66 (8)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.1:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="1" docTitle="Hyperolius tuberculatus Photo" docType="treatment" docVersion="4" lastPageNumber="163" masterDocId="FF880B42FFFBA15FFF80FF85EE63FFB7" masterDocTitle="Synopsis of the Amphibians of Equatorial Guinea based upon the Authors Field Work and Spanish Natural History Collections" masterLastPageNumber="230" masterPageNumber="137" pageNumber="163" updateTime="1714671758489" updateUser="ExternalLinkService">
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<mods:title id="2BEF125101FAE805DBD7EFB8B4A77BCE">Synopsis of the Amphibians of Equatorial Guinea based upon the Authors Field Work and Spanish Natural History Collections</mods:title>
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<mods:namePart id="3F523356B4B146CF82B304E62FBB2812">Calvo-Revuelta, Marta</mods:namePart>
<mods:affiliation id="7AF2619D99101E07D96FCFD62B5147BD">Museo Nacional de Ciencias Naturales (MNCN-CSIC), C / José Gutiérrez Abascal, 2. 28006, Madrid, Spain;</mods:affiliation>
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<mods:namePart id="A2D0B15E59CE3AB3AEF7341DC33502BD">Castroviejo-Fisher, Santiago</mods:namePart>
<mods:affiliation id="F17741DADBFF8BD519E3442D8A313B4D">Laboratorio de Sistemática de Vertebrados, Pontifícia Universidade Católica do Rio Grande do Sul (PUCRS). Av / Ipiranga, 6681, Prédio 40, sala 110, 90619 - 900, Porto Alegre, Brazil; * Corresponding author: IDlR (iriva @ mncn. csic. es)</mods:affiliation>
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<mods:namePart id="6C34A4B8C03BC06B44D6120C709D2BFC">De, Ignacio</mods:namePart>
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<mods:date id="71EF6BBB3048DFD1DE205548939C5C2D">2020</mods:date>
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<paragraph id="8BA7C22CFFE1A145FFFBFC99EAECFC83" blockId="26.[123,1174,796,1366]" box="[123,1167,796,821]" pageId="26" pageNumber="163">
<heading id="D0EF7540FFE1A145FFFBFC99EAECFC83" box="[123,1167,796,821]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="163" reason="4">
<emphasis id="B96C1E3EFFE1A145FFFBFC99EAECFC83" bold="true" box="[123,1167,796,821]" pageId="26" pageNumber="163">
<taxonomicName id="4C18B9AFFFE1A145FFFBFC99ED87FC82" ID-CoL="3NR26" authority="(Mocquard, 1897) Photo" authorityName="Photo" baseAuthorityName="Mocquard" baseAuthorityYear="1897" box="[123,996,796,821]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="26" pageNumber="163" phylum="Chordata" rank="species" species="tuberculatus">
<emphasis id="B96C1E3EFFE1A145FFFBFC99EFE6FC83" bold="true" box="[123,389,796,820]" italics="true" pageId="26" pageNumber="163">Hyperolius tuberculatus</emphasis>
(Mocquard, 1897) Photo
</taxonomicName>
figures 17EG
</emphasis>
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<paragraph id="8BA7C22CFFE1A145FF2BFCB9EC3EFCE3" blockId="26.[123,1174,796,1366]" box="[171,605,824,853]" pageId="26" pageNumber="163">
<materialsCitation id="3B70C871FFE1A145FF2BFCB9EC35FCE2" box="[171,598,824,853]" collectionCode="LOCALITY" country="Gabon" location="Gabon" pageId="26" pageNumber="163" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54A37C8EFFE1A145FF2BFCB9EE82FCE3" box="[171,225,828,852]" pageId="26" pageNumber="163">TYPE</typeStatus>
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<collectionCode id="ED095AE9FFE1A145FF68FCBAEF2AFCE4" box="[232,329,831,851]" pageId="26" pageNumber="163">LOCALITY</collectionCode>
</smallCapsWord>
.— “Lambaréné”,
<collectingCountry id="F30F82BCFFE1A145FD8FFCBEEC35FCE2" box="[527,598,827,853]" name="Gabon" pageId="26" pageNumber="163">Gabon</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE1A145FF2BFCD9ECF0FC63" pageId="26" pageNumber="163" type="distribution">
<paragraph id="8BA7C22CFFE1A145FF2BFCD9ECF0FC63" blockId="26.[123,1174,796,1366]" pageId="26" pageNumber="163">
<smallCapsWord id="8D4154F0FFE1A145FF2BFCD9EF5FFCC4" baselines="878,878" box="[171,316,860,884]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="26" pageNumber="163">DISTRIBUTION</smallCapsWord>
.— This species extends over rainforests from southeastern
<collectingCountry id="F30F82BCFFE1A145FC4BFCD9EA78FCC2" box="[971,1051,860,885]" name="Nigeria" pageId="26" pageNumber="163">Nigeria</collectingCountry>
to western
<collectingCountry id="F30F82BCFFE1A145FFFBFCFEEFF3FC22" box="[123,400,890,917]" name="Central African Republic" pageId="26" pageNumber="163">Central African Republic</collectingCountry>
, and southwards to
<collectingCountry id="F30F82BCFFE1A145FDFAFCFEED2DFC22" box="[634,846,890,917]" name="Republic of the Congo" pageId="26" pageNumber="163">Republic of Congo</collectingCountry>
and
<collectingCountry id="F30F82BCFFE1A145FC0BFCF9EEA1FC02" name="Democratic Republic of the Congo" pageId="26" pageNumber="163">Democratic Republic of Congo</collectingCountry>
. In
<collectingCountry id="F30F82BCFFE1A145FF6EFC1EEFD1FC02" box="[238,434,923,949]" name="Equatorial Guinea" pageId="26" pageNumber="163">Equatorial Guinea</collectingCountry>
, it has been recorded from both
<collectingCountry id="F30F82BCFFE1A145FC94FC1DED1EFC03" box="[788,893,920,949]" name="Equatorial Guinea" pageId="26" pageNumber="163">Río Muni</collectingCountry>
(
<bibRefCitation id="EF89BFDDFFE1A145FC0DFC19EA5DFC02" author="DE LA RIVA" box="[909,1086,923,949]" pageId="26" pageNumber="163" pagination="123 - 139" refId="ref29939" refString="DE LA RIVA, I. 1994. Anfibios anuros del Parque Nacional de Monte Alen, Rio Muni, Guinea Ecuatorial. Revista Espanola de Herpetologia 8: 123 - 139." type="journal article" year="1994">De la Riva 1994</bibRefCitation>
;
<bibRefCitation id="EF89BFDDFFE1A145FBCEFC1EEEB2FC62" author="BELL, R. C. &amp; J. L. PARRA &amp; G. BADJEDJEA &amp; M. F. BAREJ &amp; D. C. BLACKBURN &amp; M. BURGER &amp; A. CHANNING &amp; J. M. DEHLING &amp; E. GREENBAUM &amp; V. GVOZDIK &amp; J. KIELGAST &amp; C. KASUMBA &amp; S. LOTTERS &amp; P. J. MCLAUGHLIN &amp; Z. T. NAGY &amp; M. - O. RODEL &amp; D. M. PORTIK &amp; B. L. STUART &amp; J. VANDERWAL &amp; A. G. ZASSI-BOULOU &amp; K. R. ZAMUDIO" pageId="26" pageNumber="163" pagination="5223 - 5244" refId="ref28660" refString="BELL, R. C., J. L. PARRA, G. BADJEDJEA, M. F. BAREJ, D. C. BLACKBURN, M. BURGER, A. CHANNING, J. M. DEHLING, E. GREENBAUM, V. GVOZDIK, J. KIELGAST, C. KASUMBA, S. LOTTERS, P. J. MCLAUGHLIN, Z. T. NAGY, M. - O. RODEL, D. M. PORTIK, B. L. STUART, J. VANDERWAL, A. G. ZASSI-BOULOU, AND K. R. ZAMUDIO. 2017. Idiosyncratic responses to climate-driven forest fragmentation and marine incursions in reed frogs from Central Africa and the Gulf of Guinea Islands. Molecular Ecology 26: 5223 - 5244." type="journal article" year="2017">Bell et al. 2017</bibRefCitation>
) and
<taxonomicName id="4C18B9AFFFE1A145FE8FFC3EEC6CFC62" authority="(Bell et al. 2017)" baseAuthorityName="Bell" baseAuthorityYear="2017" box="[271,527,955,981]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="26" pageNumber="163" phylum="Chordata" rank="species" species="bioko">
Bioko (
<bibRefCitation id="EF89BFDDFFE1A145FEE1FC3EEC66FC62" author="BELL, R. C. &amp; J. L. PARRA &amp; G. BADJEDJEA &amp; M. F. BAREJ &amp; D. C. BLACKBURN &amp; M. BURGER &amp; A. CHANNING &amp; J. M. DEHLING &amp; E. GREENBAUM &amp; V. GVOZDIK &amp; J. KIELGAST &amp; C. KASUMBA &amp; S. LOTTERS &amp; P. J. MCLAUGHLIN &amp; Z. T. NAGY &amp; M. - O. RODEL &amp; D. M. PORTIK &amp; B. L. STUART &amp; J. VANDERWAL &amp; A. G. ZASSI-BOULOU &amp; K. R. ZAMUDIO" box="[353,517,955,981]" pageId="26" pageNumber="163" pagination="5223 - 5244" refId="ref28660" refString="BELL, R. C., J. L. PARRA, G. BADJEDJEA, M. F. BAREJ, D. C. BLACKBURN, M. BURGER, A. CHANNING, J. M. DEHLING, E. GREENBAUM, V. GVOZDIK, J. KIELGAST, C. KASUMBA, S. LOTTERS, P. J. MCLAUGHLIN, Z. T. NAGY, M. - O. RODEL, D. M. PORTIK, B. L. STUART, J. VANDERWAL, A. G. ZASSI-BOULOU, AND K. R. ZAMUDIO. 2017. Idiosyncratic responses to climate-driven forest fragmentation and marine incursions in reed frogs from Central Africa and the Gulf of Guinea Islands. Molecular Ecology 26: 5223 - 5244." type="journal article" year="2017">Bell et al. 2017</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="1323DEA9FFE1A145FD9FFC39ECE7FC62" box="[543,644,956,981]" captionStart="MAPS" captionStartId="83.[155,174,1667,1687]" captionTargetBox="[170,1124,242,1634]" captionTargetId="figure-67@83.[139,1156,216,1654]" captionTargetPageId="83" captionText="MAPS 18AC. Distribution maps for Equatorial Guinean records of (A) Hyperolius phantasticus; (B) Hyperolius platyceps; (C) Hyperolius tuberculatus." figureDoi="http://doi.org/10.5281/zenodo.11106101" httpUri="https://zenodo.org/record/11106101/files/figure.png" pageId="26" pageNumber="163">Map 18C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30291A7FFE1A145FF2BFC59ED27FAE1" pageId="26" pageNumber="163" type="discussion">
<paragraph id="8BA7C22CFFE1A145FF2BFC59EEAAFB62" blockId="26.[123,1174,796,1366]" pageId="26" pageNumber="163">
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.—
<bibRefCitation id="EF89BFDDFFE1A145FEC8FC59EF93FC42" author="MERTENS, R." box="[328,496,987,1013]" pageId="26" pageNumber="163" pagination="14 - 29" refId="ref31892" refString="MERTENS, R. 1965. Die Amphibien von Fernando Poo. Bonner Zoologische Beitrage 16: 14 - 29." type="journal article" year="1965">Mertens (1965)</bibRefCitation>
cited
<taxonomicName id="4C18B9AFFFE1A145FDB5FC58EDB9FC42" authority="(Hallowell, 1844)" baseAuthorityName="Hallowell" baseAuthorityYear="1844" box="[565,986,987,1013]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="26" pageNumber="163" phylum="Chordata" rank="species" species="concolor">
<emphasis id="B96C1E3EFFE1A145FDB5FC58ED71FC42" box="[565,786,989,1013]" italics="true" pageId="26" pageNumber="163">Hyperolius concolor</emphasis>
(Hallowell, 1844)
</taxonomicName>
on Bioko (Moca lake surroundings) but this record likely represents a misidentification with
<taxonomicName id="4C18B9AFFFE1A145FC22FC7BEA24FBA1" authorityName="Photo" baseAuthorityName="Mocquard" baseAuthorityYear="1897" box="[930,1095,1022,1046]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="26" pageNumber="163" phylum="Chordata" rank="species" species="tuberculatus">
<emphasis id="B96C1E3EFFE1A145FC22FC7BEA24FBA1" box="[930,1095,1022,1046]" italics="true" pageId="26" pageNumber="163">H. tuberculatus</emphasis>
</taxonomicName>
, which has a similar dorsal pattern at juvenile stages, and it is a very common species around that locality (surprisingly, no records of
<taxonomicName id="4C18B9AFFFE1A145FE39FBBAEC3EFBE1" authorityName="Photo" baseAuthorityName="Mocquard" baseAuthorityYear="1897" box="[441,605,1086,1110]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="26" pageNumber="163" phylum="Chordata" rank="species" species="tuberculatus">
<emphasis id="B96C1E3EFFE1A145FE39FBBAEC3EFBE1" box="[441,605,1086,1110]" italics="true" pageId="26" pageNumber="163">H. tuberculatus</emphasis>
</taxonomicName>
were provided by
<bibRefCitation id="EF89BFDDFFE1A145FCA2FBB8EDA4FBE2" author="MERTENS, R." box="[802,967,1083,1109]" pageId="26" pageNumber="163" pagination="14 - 29" refId="ref31892" refString="MERTENS, R. 1965. Die Amphibien von Fernando Poo. Bonner Zoologische Beitrage 16: 14 - 29." type="journal article" year="1965">Mertens [1965]</bibRefCitation>
). Thus, we exclude
<taxonomicName id="4C18B9AFFFE1A145FFFBFBDAEE98FBC1" baseAuthorityName="Hallowell" baseAuthorityYear="1844" box="[123,251,1118,1142]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="26" pageNumber="163" phylum="Chordata" rank="species" species="concolor">
<emphasis id="B96C1E3EFFE1A145FFFBFBDAEE98FBC1" box="[123,251,1118,1142]" italics="true" pageId="26" pageNumber="163">H. concolor</emphasis>
</taxonomicName>
—a species distributed from
<collectingCountry id="F30F82BCFFE1A145FDB1FBD9ECD8FBC2" box="[561,699,1116,1141]" name="Sierra Leone" pageId="26" pageNumber="163">Sierra Leona</collectingCountry>
to western
<collectingCountry id="F30F82BCFFE1A145FCB8FBD9EDC4FBC2" box="[824,935,1116,1141]" name="Cameroon" pageId="26" pageNumber="163">Cameroon</collectingCountry>
—from the amphibian list of
<collectingCountry id="F30F82BCFFE1A145FF47FBF9EFEDFB22" box="[199,398,1148,1174]" name="Equatorial Guinea" pageId="26" pageNumber="163">Equatorial Guinea</collectingCountry>
. Phylogeographic patterns of this species were studied by
<bibRefCitation id="EF89BFDDFFE1A145FBA6FBF9EEA1FB01" author="BELL, R. C. &amp; J. L. PARRA &amp; G. BADJEDJEA &amp; M. F. BAREJ &amp; D. C. BLACKBURN &amp; M. BURGER &amp; A. CHANNING &amp; J. M. DEHLING &amp; E. GREENBAUM &amp; V. GVOZDIK &amp; J. KIELGAST &amp; C. KASUMBA &amp; S. LOTTERS &amp; P. J. MCLAUGHLIN &amp; Z. T. NAGY &amp; M. - O. RODEL &amp; D. M. PORTIK &amp; B. L. STUART &amp; J. VANDERWAL &amp; A. G. ZASSI-BOULOU &amp; K. R. ZAMUDIO" pageId="26" pageNumber="163" pagination="5223 - 5244" refId="ref28660" refString="BELL, R. C., J. L. PARRA, G. BADJEDJEA, M. F. BAREJ, D. C. BLACKBURN, M. BURGER, A. CHANNING, J. M. DEHLING, E. GREENBAUM, V. GVOZDIK, J. KIELGAST, C. KASUMBA, S. LOTTERS, P. J. MCLAUGHLIN, Z. T. NAGY, M. - O. RODEL, D. M. PORTIK, B. L. STUART, J. VANDERWAL, A. G. ZASSI-BOULOU, AND K. R. ZAMUDIO. 2017. Idiosyncratic responses to climate-driven forest fragmentation and marine incursions in reed frogs from Central Africa and the Gulf of Guinea Islands. Molecular Ecology 26: 5223 - 5244." type="journal article" year="2017">Bell et al. (2017)</bibRefCitation>
. Descriptions of the advertisement calls from Monte Alén were provided by
<bibRefCitation id="EF89BFDDFFE1A145FB93FB19EEA1FB61" author="BOSCH, J. &amp; I. DE LA RIVA &amp; R. MARQUEZ" pageId="26" pageNumber="163" pagination="246 - 255" refId="ref29223" refString="BOSCH, J., I. DE LA RIVA, AND R. MARQUEZ. 2000. Advertisement calls of seven species of hyperoliid frogs, from Equatorial Guinea. Amphibia-Reptilia 21 (2): 246 - 255." type="journal article" year="2000">Bosch et al. (2000)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA7C22CFFE1A145FF2BFB58ED27FAE1" blockId="26.[123,1174,796,1366]" pageId="26" pageNumber="163">
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.—
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. Asonga (Bata-Niefang),
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(
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),
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</date>
(
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),
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<collectingDate id="EFE21D04FFE1A145FC8DFB78EDC3FAA1" box="[781,928,1276,1302]" pageId="26" pageNumber="163" value="1985-04-24">24 April 1985</collectingDate>
</date>
(
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: 2100621007); Engong (Aconibe-Asoc),
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<collectingDate id="EFE21D04FFE1A145FE0EFA98EC7FFA81" box="[398,540,1309,1334]" pageId="26" pageNumber="163" value="1987-05-01">01 May 1987</collectingDate>
</date>
(
<specimenCode id="DBBE6A57FFE1A145FDACFA98ECC8FA81" box="[556,683,1309,1334]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="26" pageNumber="163">EBD 25072</specimenCode>
)
</materialsCitation>
;
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<location id="8EC794F7FFE1A145FD41FA99ED4DFA81" LSID="urn:lsid:plazi:treatment:03B1733AFFE1A145FFFBFC99ED27FAE1:8EC794F7FFE1A145FD41FA99ED4DFA81" box="[705,814,1308,1334]" municipality="San Joaquin de Ndyiacom" name="Ayamiken" pageId="26" pageNumber="163">Ayamiken</location>
,
<collectingMunicipality id="6BC35856FFE1A145FCBBFA98EA31FA81" box="[827,1106,1305,1334]" pageId="26" pageNumber="163">San Joaquín de Ndyiacom</collectingMunicipality>
, 1990 (
<specimenCount id="9D1E09A5FFE1A145FF04FAB8EF77FAE1" box="[132,276,1341,1366]" count="11" pageId="26" pageNumber="163" type="generic">11 specimens</specimenCount>
, without individual voucher numbers,
<specimenCode id="DBBE6A57FFE1A145FD35FABBED56FAE1" box="[693,821,1341,1366]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="26" pageNumber="163">EBD 31516</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>