184 lines
20 KiB
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184 lines
20 KiB
XML
<document id="A1F8C769CF8ED7B363C33419C8AF5251" ID-CLB-Dataset="294074" ID-DOI="10.4202/app.00620.2019" ID-GBIF-Dataset="757ce45b-cf0d-4a94-af3a-ab063f71f48b" ID-ISSN="1732-2421" ID-Zenodo-Dep="10980956" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="valdenar" checkinTime="1713284361654" checkinUser="felipe" docAuthor="Pindakiewicz, Maciej, Tałanda, Mateusz, Sulej, Tomasz, Niedźwiedzki, Grzegorz, Sennikov, Andrey G., Bakaev, Alexandr S., Bulanov, Valeriy V., Golubev, Valeriy K. & Minikh, Alla V." docDate="2020" docId="03B1726D8119FFE4FFD94F11FC7BFE5F" docLanguage="en" docName="ActaPalaeontolPol.65.1.71-79.pdf" docOrigin="Acta Palaeontologica Polonica 65 (1)" docSource="http://dx.doi.org/10.4202/app.00620.2019" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Isadia arefievi Minikh 2015" docType="treatment" docVersion="7" lastPageNumber="74" masterDocId="FF880A15811AFFE7FFBB4C4EFF8EFFD8" masterDocTitle="Feeding convergence among ray-finned fishes: Teeth of the herbivorous actinopterygians from the latest Permian of East European Platform, Russia" masterLastPageNumber="79" masterPageNumber="71" pageNumber="74" updateTime="1736273494839" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F15F964BB7E6FC7F24AC495146B2E964">Feeding convergence among ray-finned fishes: Teeth of the herbivorous actinopterygians from the latest Permian of East European Platform, Russia</mods:title>
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<mods:namePart id="99802FCEE826995481D12BE96020AE72">Pindakiewicz, Maciej</mods:namePart>
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<mods:namePart id="6DBAC05F834B13843D6D07F189A0A03B">Sulej, Tomasz</mods:namePart>
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<mods:namePart id="5A0C922675EB295FF29DA97426D0992D">Niedźwiedzki, Grzegorz</mods:namePart>
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<mods:namePart id="493046C85393D04CF2C7021251B3423E">Sennikov, Andrey G.</mods:namePart>
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<mods:namePart id="C41AFD5FC56BB3A718646A8AF68949A8">Bakaev, Alexandr S.</mods:namePart>
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<mods:namePart id="BAB876BC11EF7D91916FFCD6F54669C5">Bulanov, Valeriy V.</mods:namePart>
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<mods:namePart id="35ABAA35F2F0C1CEBA3C6B9515BAE780">Golubev, Valeriy K.</mods:namePart>
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<mods:namePart id="C8F1876794995D9DC945D674B4AED5D8">Minikh, Alla V.</mods:namePart>
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<mods:date id="57E2BCA4D34F93D425129C54AF4994F8">2020</mods:date>
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<treatment id="03B1726D8119FFE4FFD94F11FC7BFE5F" ID-DOI="http://doi.org/10.5281/zenodo.10987183" ID-GBIF-Taxon="225693962" ID-Zenodo-Dep="10987183" LSID="urn:lsid:plazi:treatment:03B1726D8119FFE4FFD94F11FC7BFE5F" httpUri="http://treatment.plazi.org/id/03B1726D8119FFE4FFD94F11FC7BFE5F" lastPageNumber="74" pageId="3" pageNumber="74" scope_class="Actinopterygii" scope_family="Eurynotoidiidae" scope_order="Eurynotoidiformes">
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<subSubSection id="C30290F08119FFE4FFD94F11FE5EFCA3" box="[98,464,863,891]" pageId="3" pageNumber="74" type="nomenclature">
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<paragraph id="8BA7C37B8119FFE4FFD94F11FE5EFCA3" blockId="3.[98,464,863,891]" box="[98,464,863,891]" pageId="3" pageNumber="74">
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<heading id="D0EF74178119FFE4FFD94F11FE5EFCA3" box="[98,464,863,891]" fontSize="12" level="2" pageId="3" pageNumber="74" reason="2">
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<taxonomicName id="4C18B8F88119FFE4FFD94F11FE5EFCA3" authority="Minikh, 2015" authorityName="Minikh" authorityYear="2015" box="[98,464,863,891]" class="Actinopterygii" family="Eurynotoidiidae" genus="Isadia" kingdom="Animalia" order="Eurynotoidiformes" pageId="3" pageNumber="74" phylum="Chordata" rank="species" species="arefievi">
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<emphasis id="B96C1F698119FFE4FFD94F11FE95FCA3" box="[98,283,863,891]" italics="true" pageId="3" pageNumber="74">Isadia arefievi</emphasis>
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<bibRefCitation id="EF89BE8A8119FFE4FE994F11FE5EFCA3" author="Minikh, A. V. & Arefiev, M. P. & Golubev, V. K." box="[290,464,863,891]" pageId="3" pageNumber="74" pagination="615 - 626" refId="ref6931" refString="Minikh, A. V., Arefiev, M. P., and Golubev, V. K. 2015. A new fish species of the genus Isadia (Actinopterygii, Eurynototoidiformes) from the new locality on the Malaya Northern Dvina River (terminal Permian, Vologda Region). Paleontological Journal 49 (6): 615 - 626." type="journal article" year="2015">Minikh, 2015</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C30290F08119FFE4FFD94FC5FC7BFE5F" pageId="3" pageNumber="74" type="description">
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<paragraph id="8BA7C37B8119FFE4FFD94FC5FF26FC78" blockId="3.[98,168,907,928]" box="[98,168,907,928]" pageId="3" pageNumber="74">
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<figureCitation id="1323DFFE8119FFE4FFD94FC5FF2BFC78" box="[98,165,907,928]" captionStart="Fig" captionStartId="4.[151,184,1797,1817]" captionTargetBox="[174,1512,159,1772]" captionTargetId="figure-9@4.[174,1512,159,1772]" captionTargetPageId="4" captionText="Fig. 2. The isolated teeth of actinopterygian fish Isadia from the Sokovka outcrop, Vyazniki, Russia, late Permian (Upper Vyatkian). A–D. Isadia aristoviensis Minikh, 1990, mandibulary teeth. A. ZPAL V.51/1, lingual view. B. ZPAL V.51/2, labial view. C. ZPAL V.51/3, lingual view. D. ZPAL V.51/4, labial view. E–I. Isadia aristoviensis Minikh, 1990, maxillary teeth. E. ZPAL V.51/6, lingual view. F. ZPAL V.51/7, labial view. G. ZPAL V.51/5, lingual view. H. ZPAL V.51/8, lingual view. I. ZPAL V.51/9, labial view. J. Isadia arefievi Minikh, 2015, ZPAL V.51/10, mandibular tooth,?lingual view. K, L. Isadia suchonensis Minikh, 1986, mandibular teeth. K. ZPAL V.51/11, lingual (K1) and lateral (K2) views. L. ZPAL V.51/12, labial view. M. Isadia suchonensis Minikh, 1986, ZPAL V.51/13, maxillary teeth,?labial view. Scale bars 1 mm (A–I), 0.5 mm (J, K, M), 0.2 mm (L)." figureDoi="http://doi.org/10.5281/zenodo.10980958" httpUri="https://zenodo.org/record/10980958/files/figure.png" pageId="3" pageNumber="74">Fig. 2J</figureCitation>
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.
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</paragraph>
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<paragraph id="8BA7C37B8119FFE4FFD94FFDFF56FBF5" blockId="3.[98,771,947,1069]" pageId="3" pageNumber="74">
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<emphasis id="B96C1F698119FFE4FFD94FFDFF48FC14" box="[98,198,947,972]" italics="true" pageId="3" pageNumber="74">Material</emphasis>
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.—Isolated tooth (
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<collectionCode id="ED095BBE8119FFE4FE1E4FFDFE62FC14" box="[421,492,947,972]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="3" pageNumber="74">ZPAL</collectionCode>
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<collectionCode id="ED095BBE8119FFE4FE474FFDFD9EFC14" box="[508,528,947,972]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="74" type="Museum">V</collectionCode>
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.51/10) and two unnumbered mandibular teeth from Sokovka outcrop, Vyazniki in
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<collectingRegion id="49DC0D998119FFE4FF7E4FBDFE1DFBD4" box="[197,403,1011,1036]" country="Russia" name="Vladimir" pageId="3" pageNumber="74">Vladimir Region</collectingRegion>
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,
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<collectingCountry id="F30F83EB8119FFE4FE1D4FBDFE7BFBD4" box="[422,501,1011,1036]" name="Russia" pageId="3" pageNumber="74">Russia</collectingCountry>
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, late Permian (Upper Vyatkian).
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</paragraph>
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<paragraph id="8BA7C37B8119FFE4FFD94871FDDDF9C3" blockId="3.[98,771,1087,1563]" pageId="3" pageNumber="74">
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<emphasis id="B96C1F698119FFE4FFD94871FF64FB80" box="[98,234,1087,1112]" italics="true" pageId="3" pageNumber="74">Description</emphasis>
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.—The base of the crown is labio-lingually curved, short and thick. These are characters unique for
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<taxonomicName id="4C18B8F88119FFE4FFD94831FE84FB40" authorityName="Minikh" authorityYear="2015" box="[98,266,1151,1176]" class="Actinopterygii" family="Eurynotoidiidae" genus="Isadia" kingdom="Animalia" order="Eurynotoidiformes" pageId="3" pageNumber="74" phylum="Chordata" rank="species" species="arefievi">
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<emphasis id="B96C1F698119FFE4FFD94831FE84FB40" box="[98,266,1151,1176]" italics="true" pageId="3" pageNumber="74">Isadia arefievi</emphasis>
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</taxonomicName>
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teeth (
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<bibRefCitation id="EF89BE8A8119FFE4FEE74831FDB4FB40" author="Minikh, A. V. & Arefiev, M. P. & Golubev, V. K." box="[348,570,1151,1176]" pageId="3" pageNumber="74" pagination="615 - 626" refId="ref6931" refString="Minikh, A. V., Arefiev, M. P., and Golubev, V. K. 2015. A new fish species of the genus Isadia (Actinopterygii, Eurynototoidiformes) from the new locality on the Malaya Northern Dvina River (terminal Permian, Vologda Region). Paleontological Journal 49 (6): 615 - 626." type="journal article" year="2015">Minikh et al. 2015</bibRefCitation>
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). In the collected teeth the whole crown is very wide and has three cusps (the cusp number of
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<taxonomicName id="4C18B8F88119FFE4FE9948F1FE1BFB00" authorityName="Minikh" authorityYear="2015" box="[290,405,1215,1240]" class="Actinopterygii" family="Eurynotoidiidae" genus="Isadia" kingdom="Animalia" order="Eurynotoidiformes" pageId="3" pageNumber="74" phylum="Chordata" rank="species" species="arefievi">
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<emphasis id="B96C1F698119FFE4FE9948F1FE1BFB00" box="[290,405,1215,1240]" italics="true" pageId="3" pageNumber="74">I. arefievi</emphasis>
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</taxonomicName>
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range up to 5, what it seen on specimens from Eleonora locality; see
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<bibRefCitation id="EF89BE8A8119FFE4FDA448AEFD79FB21" author="Minikh, A. V. & Arefiev, M. P. & Golubev, V. K." box="[543,759,1248,1273]" pageId="3" pageNumber="74" pagination="615 - 626" refId="ref6931" refString="Minikh, A. V., Arefiev, M. P., and Golubev, V. K. 2015. A new fish species of the genus Isadia (Actinopterygii, Eurynototoidiformes) from the new locality on the Malaya Northern Dvina River (terminal Permian, Vologda Region). Paleontological Journal 49 (6): 615 - 626." type="journal article" year="2015">Minikh et al. 2015</bibRefCitation>
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). The middle cusp is especially wide, making crown shovelshaped. Flat crown is lightly labiolingually curved. The two outer cusplets are very small, thin, and occur on different heights of the tooth. The distal half of the crown is made of acrodine. Width of the crown is between 1.2–
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<quantity id="4CE06E9E8119FFE4FDCD49CFFD32FA42" box="[630,700,1409,1434]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="74" unit="mm" value="2.0">2 mm</quantity>
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, teeth are smaller than in
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<taxonomicName id="4C18B8F88119FFE4FEF349EFFDA9FA62" authorityName="Minikh" authorityYear="1990" box="[328,551,1441,1466]" class="Actinopterygii" family="Eurynotoidiidae" genus="Isadia" kingdom="Animalia" order="Eurynotoidiformes" pageId="3" pageNumber="74" phylum="Chordata" rank="species" species="aristoviensis">
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<emphasis id="B96C1F698119FFE4FEF349EFFDA9FA62" box="[328,551,1441,1466]" italics="true" pageId="3" pageNumber="74">Isadia aristoviensis</emphasis>
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</taxonomicName>
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||
. The length is unknown because collected specimens are not well preserved. They represent mandibular teeth according to the number of cusplets in this species (
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<bibRefCitation id="EF89BE8A8119FFE4FECA4A4CFDC9F9C3" author="Minikh, A. V. & Arefiev, M. P. & Golubev, V. K." box="[369,583,1538,1563]" pageId="3" pageNumber="74" pagination="615 - 626" refId="ref6931" refString="Minikh, A. V., Arefiev, M. P., and Golubev, V. K. 2015. A new fish species of the genus Isadia (Actinopterygii, Eurynototoidiformes) from the new locality on the Malaya Northern Dvina River (terminal Permian, Vologda Region). Paleontological Journal 49 (6): 615 - 626." type="journal article" year="2015">Minikh et al. 2015</bibRefCitation>
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||
).
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</paragraph>
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<paragraph id="8BA7C37B8119FFE4FFD94A62FA94FEC2" blockId="3.[98,771,1580,1960]" lastBlockId="3.[808,1481,160,282]" pageId="3" pageNumber="74">
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<emphasis id="B96C1F698119FFE4FFD94A62FF4AF99D" box="[98,196,1580,1605]" italics="true" pageId="3" pageNumber="74">Remarks</emphasis>
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||
.—The teeth of
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<taxonomicName id="4C18B8F88119FFE4FE384A62FDA3F99D" authorityName="Minikh" authorityYear="2015" box="[387,557,1580,1605]" class="Actinopterygii" family="Eurynotoidiidae" genus="Isadia" kingdom="Animalia" order="Eurynotoidiformes" pageId="3" pageNumber="74" phylum="Chordata" rank="species" species="arefievi">
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<emphasis id="B96C1F698119FFE4FE384A62FDA3F99D" box="[387,557,1580,1605]" italics="true" pageId="3" pageNumber="74">Isadia arefievi</emphasis>
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</taxonomicName>
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are rare in material from Sokovka. The same morphotype was found only in
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<collectingRegion id="49DC0D998119FFE4FF7B4A23FEF2F95E" box="[192,380,1645,1670]" country="Russia" name="Vologda" pageId="3" pageNumber="74">Vologda Region</collectingRegion>
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,
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<collectingCountry id="F30F83EB8119FFE4FE314A23FE56F95E" box="[394,472,1645,1670]" name="Russia" pageId="3" pageNumber="74">Russia</collectingCountry>
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, Eleonora site (Salavyoro Formation) and Nizhny Novgorog Region,
|
||
<collectingCountry id="F30F83EB8119FFE4FDF84AC3FD1FF97E" box="[579,657,1677,1702]" name="Russia" pageId="3" pageNumber="74">Russia</collectingCountry>
|
||
, Lagernyi Ovrag-3 (Obnora Formation), dated as lower part of the uppermost Permian by ichthyozone
|
||
<emphasis id="B96C1F698119FFE4FDB14A80FED8F8DF" italics="true" pageId="3" pageNumber="74">
|
||
<taxonomicName id="4C18B8F88119FFE4FDB14A80FD7AF93F" authorityName="Minikh" authorityYear="2006" box="[522,756,1742,1767]" family="Discordichthyidae" genus="Mutovinia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="74" phylum="Chordata" rank="species" species="sennikovi">Mutovinia sennikovi</taxonomicName>
|
||
– Gnathorhiza otschevi
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EF89BE8A8119FFE4FEDC4AA0FDCCF8DF" author="Minikh, A. V. & Arefiev, M. P. & Golubev, V. K." box="[359,578,1774,1799]" pageId="3" pageNumber="74" pagination="615 - 626" refId="ref6931" refString="Minikh, A. V., Arefiev, M. P., and Golubev, V. K. 2015. A new fish species of the genus Isadia (Actinopterygii, Eurynototoidiformes) from the new locality on the Malaya Northern Dvina River (terminal Permian, Vologda Region). Paleontological Journal 49 (6): 615 - 626." type="journal article" year="2015">Minikh et al. 2015</bibRefCitation>
|
||
). This species of
|
||
<taxonomicName id="4C18B8F88119FFE4FFD94B40FF26F8FF" authorityName="Minikh" authorityYear="1986" box="[98,168,1806,1831]" class="Actinopterygii" family="Eurynotoidiidae" genus="Isadia" kingdom="Animalia" order="Eurynotoidiformes" pageId="3" pageNumber="74" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96C1F698119FFE4FFD94B40FF26F8FF" box="[98,168,1806,1831]" italics="true" pageId="3" pageNumber="74">Isadia</emphasis>
|
||
</taxonomicName>
|
||
could have only one row of teeth, forming a tool for scrapping algae. Enlargement of the medium cusp suggests an adaptation to reduce pressure for each tooth while scrapping algae, creating a large abrasive surface. The same trend might be seen in durophagous hybodont teeth (
|
||
<bibRefCitation id="EF89BE8A8119FFE4FDD54BC1FD79F870" author="Ginter, M." box="[622,759,1935,1960]" pageId="3" pageNumber="74" refId="ref6408" refString="Ginter, M. 2012. Ryby kopalne. 172 pp. Wydawnictwo Uniwersytetu Warszawskiego, Warszawa." type="book" year="2012">Ginter 2012</bibRefCitation>
|
||
). Lack of wide, triangular labial roots of the base is caused probably by some taphonomic factors. The teeth described here suggest that
|
||
<taxonomicName id="4C18B8F88119FFE4FC554CAFFBD3FF22" authorityName="Minikh" authorityYear="2015" box="[1006,1117,225,250]" class="Actinopterygii" family="Eurynotoidiidae" genus="Isadia" kingdom="Animalia" order="Eurynotoidiformes" pageId="3" pageNumber="74" phylum="Chordata" rank="species" species="arefievi">
|
||
<emphasis id="B96C1F698119FFE4FC554CAFFBD3FF22" box="[1006,1117,225,250]" italics="true" pageId="3" pageNumber="74">I. arefievi</emphasis>
|
||
</taxonomicName>
|
||
survived to the terminal part of the late Permian in East European Platform.
|
||
</paragraph>
|
||
<paragraph id="8BA7C37B8119FFE4FC934D62FC7BFE5F" blockId="3.[808,1481,300,391]" pageId="3" pageNumber="74">
|
||
<emphasis id="B96C1F698119FFE4FC934D62FB5FFE9D" box="[808,1233,300,325]" italics="true" pageId="3" pageNumber="74">Stratigraphic and geographic range</emphasis>
|
||
.—Terminal late Permian of the
|
||
<collectingRegion id="49DC0D998119FFE4FC7B4D03FBAEFEBE" box="[960,1056,333,358]" country="Russia" name="Moscow" pageId="3" pageNumber="74">Moscow</collectingRegion>
|
||
Syneclise and southwestern Mezen Syneclise;
|
||
<collectingCountry id="F30F83EB8119FFE4FC194D20FC7FFE5F" box="[930,1009,366,391]" name="Russia" pageId="3" pageNumber="74">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |