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<mods:title id="DC5044AB8ED80E7545D01D4BB21DB5BA">Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata) 2539</mods:title>
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<paragraph id="8BF25812CE49EC637EEEFDB2C423CA7A" blockId="38.[151,1436,552,1186]" box="[151,802,552,579]" pageId="38" pageNumber="39">
<heading id="D0BAEF7ECE49EC637EEEFDB2C423CA7A" bold="true" box="[151,802,552,579]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="B9398400CE49EC637EEEFDB2C423CA7A" bold="true" box="[151,802,552,579]" pageId="38" pageNumber="39">
<taxonomicName id="4C4D2391CE49EC637EEEFDB2C557CA7A" ID-CoL="5XZSF" authority="Pilsbry, 1918" authorityName="Pilsbry" authorityYear="1918" box="[151,598,552,579]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="guiraoensis">
<emphasis id="B9398400CE49EC637EEEFDB2C6AFCA7B" bold="true" box="[151,430,553,579]" italics="true" pageId="38" pageNumber="39">Chondrina guiraoensis</emphasis>
Pilsbry, 1918
</taxonomicName>
(pl. 11 figs HJ)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3570B99CE49EC637EEEFDD7C46DCADB" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="8BF25812CE49EC637EEEFDD7C34DCA9B" blockId="38.[151,1436,552,1186]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="AB5D7F3CCE49EC637EEEFDD7C34DCA9B" pageId="38" pageNumber="39">
<emphasis id="B9398400CE49EC637EEEFDD7C7CECA5B" box="[151,207,588,611]" italics="true" pageId="38" pageNumber="39">Pupa</emphasis>
<treatmentCitation id="0AEC7E03CE49EC637EA1FDD7C559CA5B" author="Rossmassler, E. A." box="[216,600,588,611]" page="110" pageId="38" pageNumber="39" year="1859">
<taxonomicName id="4C4D2391CE49EC637EA1FDD7C559CA5B" ID-CoL="5XZSG" authority="Rossmassler, 1859: 110" authorityName="Rossmassler" authorityPageNumber="110" authorityYear="1859" box="[216,600,588,611]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="jumillensis">
<emphasis id="B9398400CE49EC637EA1FDD7C64DCA5B" box="[216,332,588,611]" italics="true" pageId="38" pageNumber="39">jumillensis</emphasis>
<bibRefCitation id="EFDC25E3CE49EC637F2CFDD7C559CA5B" author="Rossmassler, E. A." box="[341,600,588,611]" pageId="38" pageNumber="39" pagination="81 - 140" refId="ref37159" refString="Rossmassler, E. A. (1859) Iconographie der Land- und Susswasser-Mollusken Europa's, mit vorzuglicher Berucksichtigung kritischer und noch nicht abgebildeter Arten, 3 (5 - 6), I - VIII &amp; 81 - 140." type="book chapter" year="1859">Rossmässler, 1859: 110</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54F6E6B0CE49EC637C1CFDD7C5D3CA5B" box="[613,722,588,611]" pageId="38" pageNumber="39" type="lectotype">Lectotype</typeStatus>
, design.
<treatmentCitation id="0AEC7E03CE49EC637D4CFDD7C342CA5B" author="Gittenberger, E." box="[821,1091,588,611]" page="213" pageId="38" pageNumber="39" year="1973">
<bibRefCitation id="EFDC25E3CE49EC637D4CFDD7C342CA5B" author="Gittenberger, E." box="[821,1091,588,611]" pageId="38" pageNumber="39" pagination="1 - 267" refId="ref34929" refString="Gittenberger, E. (1973) Beitrage zur Kenntnis der Pupillacea III. Chondrininae. Zoologische Verhandelingen, 127, 1 - 267." type="journal article" year="1973">Gittenberger (1973: 213)</bibRefCitation>
</treatmentCitation>
(SMF 43677a):
<collectingCountry id="F35A1882CE49EC637A85FDD7C23ACA5B" box="[1276,1339,588,611]" name="Spain" pageId="38" pageNumber="39">Spain</collectingCountry>
,
<collectingRegion id="498996F0CE49EC637B31FDD7C299CA5B" box="[1352,1432,588,611]" country="Spain" name="Murcia" pageId="38" pageNumber="39">Murcia</collectingRegion>
, Jumilla. The shell characters enable the conclusion that the
<typeStatus id="54F6E6B0CE49EC637D2DFDF7C4B7CABB" box="[852,950,620,643]" pageId="38" pageNumber="39" type="lectotype">lectotype</typeStatus>
is not from Orihuela, Alicante, which is the second locality mentioned on the original label (
<treatmentCitation id="0AEC7E03CE49EC637CB0FD17C474CA9B" author="Haas, F." box="[713,885,652,675]" page="304" pageId="38" pageNumber="39" year="1926">
<bibRefCitation id="EFDC25E3CE49EC637CB0FD17C474CA9B" author="Haas, F." box="[713,885,652,675]" pageId="38" pageNumber="39" pagination="267 - 315" refId="ref35163" refString="Haas, F. (1926) Appendix I. The Abidas and Chondrinas of the Pyrenees and the Iberian peninsula. In: H. A. Pilsbry (Ed.) Manual of Conchology (Second Series: Pulmonata). Pupillidae (Orculinae, Pagodulinae, Acanthinulinae, etc.), 27 (108 [part.]), 267 - 315." type="journal article" year="1926">Haas 1926: 304</bibRefCitation>
</treatmentCitation>
; own observations).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF25812CE49EC637EEEFD37C46DCADB" blockId="38.[151,1436,552,1186]" pageId="38" pageNumber="39">
Not
<emphasis id="B9398400CE49EC637EBEFD37C7FECAFB" box="[199,255,684,707]" italics="true" pageId="38" pageNumber="39">Pupa</emphasis>
<taxonomicName id="4C4D2391CE49EC637F71FD37C51ACAFB" ID-CoL="5XZSG" authority="Pfeiffer, 1853" authorityName="L. Pfeiffer" authorityYear="1853" box="[264,539,684,707]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="jumillensis">
<emphasis id="B9398400CE49EC637F71FD37C67DCAFB" box="[264,380,684,707]" italics="true" pageId="38" pageNumber="39">jumillensis</emphasis>
<bibRefCitation id="EFDC25E3CE49EC637FFDFD37C51ACAFB" author="Pfeiffer, L." box="[388,539,684,707]" pageId="38" pageNumber="39" pagination="1 - 711" refId="ref36656" refString="Pfeiffer, L. (1853) Monographia Heliceorum Viventium. sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum, 3, 1 - 711. F. A. Brockhaus, Leipzig, 594 pp." type="book chapter" year="1853">Pfeiffer, 1853</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C4D2391CE49EC637C67FD37C4ACCAFB" ID-CoL="5XZSF" authority="Pilsbry, 1918" authorityName="Pilsbry" authorityYear="1918" box="[542,941,684,707]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="guiraoensis">
<emphasis id="B9398400CE49EC637C67FD37C416CAFB" box="[542,791,684,707]" italics="true" pageId="38" pageNumber="39">Chondrina guiraoensis</emphasis>
Pilsbry, 1918
</taxonomicName>
[March]: 372, pl. 47
<figureCitation id="13764497CE49EC637AE3FD37C3D6CAFB" box="[1178,1239,684,707]" captionStart="FIGURE 9" captionStartId="42.[151,255,1036,1060]" captionTargetBox="[296,1291,424,1002]" captionTargetId="graphics-241@42.[322,1291,440,1001]" captionTargetPageId="42" captionText="FIGURE 9. Distribution of Rupestrella species in Spain and N-Africa. A, Type localities of the taxa recognized by Pilsbry (1918) with the approximate distribution of R. dupotetii in gray; B, Localities of R. philippii in N-Africa based on material in the collection of the RMNH." figureDoi="http://doi.org/10.5281/zenodo.10094219" httpUri="https://zenodo.org/record/10094219/files/figure.png" pageId="38" pageNumber="39">fig. 9</figureCitation>
.
<typeStatus id="54F6E6B0CE49EC637A9AFD37C247CAFB" box="[1251,1350,684,707]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
(ANSP 22790):
<collectingCountry id="F35A1882CE49EC637F59FD57C661CADB" box="[288,352,716,739]" name="Spain" pageId="38" pageNumber="39">Spain</collectingCountry>
,
<collectingRegion id="498996F0CE49EC637F12FD57C6BACADB" box="[363,443,716,739]" country="Spain" name="Murcia" pageId="38" pageNumber="39">Murcia</collectingRegion>
, Jumilla (Pilsbry 1918 [November]: 51).
</paragraph>
</subSubSection>
<subSubSection id="C3570B99CE49EC637EEEFD77C317CC9A" pageId="38" pageNumber="39" type="description">
<paragraph id="8BF25812CE49EC637EEEFD77C3F4CB3B" blockId="38.[151,1436,552,1186]" box="[151,1269,748,771]" pageId="38" pageNumber="39">
<taxonomicName id="4C4D2391CE49EC637EEEFD77C50BCB3B" authority="Pilsbry, 1918" authorityName="Pilsbry" authorityYear="1918" box="[151,522,748,771]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="guiraonis">
<emphasis id="B9398400CE49EC637EEEFD77C676CB3B" box="[151,375,748,771]" italics="true" pageId="38" pageNumber="39">Chondrina guiraonis</emphasis>
Pilsbry, 1918
</taxonomicName>
[November]: 51. Unjustified emendation (n. n.) (ICZN Art. 32.5.1).
</paragraph>
<paragraph id="8BF25812CE49EC637EEEFC97C49BCB1B" blockId="38.[151,1436,552,1186]" box="[151,922,780,803]" pageId="38" pageNumber="39">
<taxonomicName id="4C4D2391CE49EC637EEEFC97C66BCB1B" baseAuthorityName="Des Moulins" baseAuthorityYear="1835" box="[151,362,780,803]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="farinesii">
<emphasis id="B9398400CE49EC637EEEFC97C66BCB1B" box="[151,362,780,803]" italics="true" pageId="38" pageNumber="39">Chondrina farinesii</emphasis>
</taxonomicName>
; Montoya et al., 1999: 131. Not
<bibRefCitation id="EFDC25E3CE49EC637CBEFC97C497CB1B" author="Des Moulins, C." box="[711,918,780,803]" pageId="38" pageNumber="39" pagination="142 - 165" refId="ref34355" refString="Des Moulins, C. (1835) Description de quelques mollusques terrestres et fluviatiles de la France, nouveau ou peu connus. Actes de la Societe Linneenne de Bordeaux, 7, 142 - 165." type="journal article" year="1835">Des Moulins, 1835</bibRefCitation>
.
</paragraph>
<paragraph id="8BF25812CE49EC637EBCFCB7C58CCBD8" blockId="38.[151,1436,552,1186]" pageId="38" pageNumber="39">Description.—Shell rather thick-set, with a short conical apical part and a last whorl which is often hardly broader than the penultimate one, light brownish to yellowish brown; teleoconch conspicuouisly sculptured with rather coarse riblets. Aperture with a more or less prominent angularis and usually a columellar denticle which is hidden behind the columella in frontal view; and only rarely an obsolete parietalis. Apertural lip neither reflected nor rib-like thickened inside.</paragraph>
<paragraph id="8BF25812CE49EC637EBCFC76C317CC9A" blockId="38.[151,1436,552,1186]" pageId="38" pageNumber="39">
Notes.—In view of the revised interpretation of the eastern Iberian
<taxonomicName id="4C4D2391CE49EC637A79FC76C37ECC3E" authorityName="Reichenbach" authorityYear="1828" box="[1024,1151,1005,1030]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">
<emphasis id="B9398400CE49EC637A79FC76C37ECC3E" box="[1024,1151,1005,1030]" italics="true" pageId="38" pageNumber="39">Chondrina</emphasis>
</taxonomicName>
taxa with more or less obsolete apertural teeth,
<taxonomicName id="4C4D2391CE49EC637FC1FB8FC555CC15" box="[440,596,1044,1069]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="guiroensis">
<emphasis id="B9398400CE49EC637FC1FB8FC555CC15" box="[440,596,1044,1069]" italics="true" pageId="38" pageNumber="39">C. guiroensis</emphasis>
</taxonomicName>
is listed here as a species. Quite surprising, the oldest fossil
<taxonomicName id="4C4D2391CE49EC637B59FB8FC29ACC16" authorityName="Reichenbach" authorityYear="1828" box="[1312,1435,1044,1070]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Chondrina</taxonomicName>
shells (pl. 11 fig. J) collected at the Sierra de Quibas (
<bibRefCitation id="EFDC25E3CE49EC637D68FBA1C304CC6C" author="Montoya, P. &amp; Alberdi, M. T. &amp; Barbadillo, L. J. &amp; Made, J. van der &amp; Morales, J. &amp; Murelaga, X. &amp; Penalver, E. &amp; Robles, F. &amp; Bustos, A. R. &amp; Sanchez, A. &amp; Sanchiz, B. &amp; Soria, D. &amp; Szyndlar, Z." box="[785,1029,1082,1108]" pageId="38" pageNumber="39" pagination="387 - 393" refId="ref36235" refString="Montoya, P., Alberdi, M. T., Barbadillo, L. J., Made, J. van der, Morales, J., Murelaga, X., Penalver, E., Robles, F., Bustos, A. R., Sanchez, A., Sanchiz, B., Soria, D. &amp; Szyndlar, Z. (2001) Une faune tres diversifiee du Pleistocene inferieur de la Sierra de Quibas (the province of Murcia, Espagne). Earth and Planetary Sciences, 332, 387 - 393." type="journal article" year="2001">Montoya et al. 2001</bibRefCitation>
), (which is only c.
<quantity id="4CB5F5F7CE49EC637A9EFBA1C230CC6C" box="[1255,1329,1082,1108]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="km" value="25.0">25 km</quantity>
SE from Jumilla) are extremely similar to the recent
<taxonomicName id="4C4D2391CE49EC637CCBFBFAC460CC42" authorityName="Pilsbry" authorityYear="1918" box="[690,865,1121,1146]" class="Gastropoda" family="Chondrinidae" genus="Chondrina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="guiraoensis">
<emphasis id="B9398400CE49EC637CCBFBFAC460CC42" box="[690,865,1121,1146]" italics="true" pageId="38" pageNumber="39">C. guiraoensis</emphasis>
</taxonomicName>
, indicating that there have been no substantial changes in shell morphology in this species during at least 1.3 million years.
</paragraph>
</subSubSection>
</treatment>
</document>