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<document id="2F4A7CD86E46FF1252A16AB4D3E9CA65" ID-CLB-Dataset="305079" ID-DOI="10.11646/zootaxa.5536.1.1" ID-GBIF-Dataset="52a66b9d-a4c0-45a2-9e5e-7c15685a8f39" ID-ISSN="1175-5326" ID-Zenodo-Dep="14023827" ID-ZooBank="C47EE4D9-D5B5-4B67-8547-856FBAEFE670" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1730455362568" checkinUser="plazi" docAuthor="Wang, Shuxia, Liu, Chen &amp; David, Lees" docDate="2024" docId="03E41827FFAF0A4CFD8EFF0CFEC2FA2B" docLanguage="en" docName="zootaxa.5536.1.1.pdf" docOrigin="Zootaxa 5536 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Promalactis spinellosa Wang &amp; Liu &amp; David 2024, sp. nov." docType="treatment" docVersion="7" lastPageNumber="30" masterDocId="FFDD605FFFB20A51FD19FF9BFFEEFFCE" masterDocTitle="Partial revision of Promalactis Meyrick (Lepidoptera: Oecophoridae), based on specimens deposited in the Natural History Museum, London" masterLastPageNumber="58" masterPageNumber="1" pageNumber="30" updateTime="1732577891658" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0B918C77B4BD70DB53FD7253F8389A19">Partial revision of Promalactis Meyrick (Lepidoptera: Oecophoridae), based on specimens deposited in the Natural History Museum, London</mods:title>
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<mods:affiliation id="6C7D3E862E94860C83324E1658AC2B07">College of Life Sciences, Nankai University, Tianjin 300071, China</mods:affiliation>
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<mods:namePart id="AE246F0D8213087F23D30909FB0025BC">David, Lees</mods:namePart>
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<mods:affiliation id="9D66CC09B1E8439E4E35FB6772C1786A">Department of Science, Natural History Museum, London, United Kingdom, SW 7 5 BD</mods:affiliation>
<mods:nameIdentifier id="0E9F96330895B05BD21382D83A9B7B84" type="email">david.lees@nhm.ac.uk</mods:nameIdentifier>
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<mods:date id="DBABC205C3F3209B3CBC8B495FD40C96">2024</mods:date>
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<treatment id="03E41827FFAF0A4CFD8EFF0CFEC2FA2B" ID-DOI="http://doi.org/10.5281/zenodo.14023909" ID-GBIF-Taxon="246064913" ID-Zenodo-Dep="14023909" LSID="urn:lsid:plazi:treatment:03E41827FFAF0A4CFD8EFF0CFEC2FA2B" httpUri="http://treatment.plazi.org/id/03E41827FFAF0A4CFD8EFF0CFEC2FA2B" lastPageNumber="30" pageId="29" pageNumber="30" scope_family="Oecophoridae" scope_order="Lepidoptera">
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<heading id="D0BA1E5DFFAF0A4CFD8EFF0CFD8BFF7F" bold="true" box="[151,613,151,178]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B9397523FFAF0A4CFD8EFF0CFD8BFF7F" bold="true" box="[151,613,151,178]" pageId="29" pageNumber="30">
<taxonomicName id="4C4DD2B2FFAF0A4CFD8EFF0CFE14FF7F" ID-CoL="CN7QQ" authority="Wang" authorityName="Wang &amp; Liu &amp; David" authorityYear="2024" box="[151,506,151,178]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="spinellosa" status="sp. nov.">
<emphasis id="B9397523FFAF0A4CFD8EFF0CFE47FF7C" bold="true" box="[151,425,151,178]" italics="true" pageId="29" pageNumber="30">Promalactis spinellosa</emphasis>
Wang
</taxonomicName>
,
<taxonomicNameLabel id="A20AC858FFAF0A4CFF1EFF0CFD8BFF7F" box="[519,613,151,177]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C357FABAFFAF0A4CFD8EFF20FEC9FF1B" box="[151,295,187,213]" pageId="29" pageNumber="30" type="description">
<paragraph id="8BF2A931FFAF0A4CFD8EFF20FEC9FF1B" blockId="29.[151,613,151,213]" box="[151,295,187,213]" pageId="29" pageNumber="30">
(
<figureCitation id="1376B5B4FFAF0A4CFD86FF20FF18FF1B" box="[159,246,187,213]" captionStart="FIGURES 3033" captionStartId="30.[151,264,1046,1070]" captionTargetBox="[174,1413,194,1009]" captionTargetId="figure-23@30.[162,1425,181,1021]" captionTargetPageId="30" captionText="FIGURES 3033. Adults of Promalactis spp. 30. P. spinellosa sp. nov.; 31. P. subfalcata sp. nov.; 32. P. trispinata sp. nov.; 33. P. varilonga sp. nov. 3032: Males and holotypes; 33: Male and paratype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023843" httpUri="https://zenodo.org/record/14023843/files/figure.png" pageId="29" pageNumber="30">Figs 30</figureCitation>
,
<figureCitation id="1376B5B4FFAF0A4CFC1BFF20FEF1FF1B" box="[258,287,187,213]" captionStart="FIGURES 5863" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[232,1359,193,1907]" captionTargetId="figure-23@36.[220,1368,181,1919]" captionTargetPageId="36" captionText="FIGURES 5863. Male genitalia of Promalactis spp. 58. P. minuticornuta sp. nov.; 59. P. parvissima sp. nov.; 60. P. polyptycha sp. nov.; 61. P. indica sp. nov.; 62. P. similidierli sp. nov.; 63. P. spinellosa sp. nov. All holotypes. Scale bar: 58, 60, 6263 = 0.2 mm; 59, 61 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14023857" httpUri="https://zenodo.org/record/14023857/files/figure.png" pageId="29" pageNumber="30">63</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFAF0A4CFD8EFE98FD4BFE44" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BF2A931FFAF0A4CFD8EFE98FD4BFE8F" blockId="29.[151,1437,259,1509]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFD8EFE98FEABFED3" bold="true" box="[151,325,259,285]" pageId="29" pageNumber="30">Type material.</emphasis>
<materialsCitation id="3B25A36CFFAF0A4CFC55FE98FD4BFE8F" ID-GBIF-Occurrence="4977736490" collectingDate="1982-02-27" collectionCode="NHMUK" collectorName="G. S. Robinson" country="Brunei" elevation="75" location="Badas" pageId="29" pageNumber="30" specimenCode="NHMUK010316990" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9397523FFAF0A4CFC55FE98FE49FED3" bold="true" box="[332,423,259,285]" pageId="29" pageNumber="30">
<collectingCountry id="F35AE9A1FFAF0A4CFC55FE98FE4DFED3" box="[332,419,259,285]" name="Brunei" pageId="29" pageNumber="30">Brunei</collectingCountry>
:
</emphasis>
<typeStatus id="54F61793FFAF0A4CFCB7FE98FDF8FED3" box="[430,534,259,285]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4B62B8FFAF0A4CFF04FE9FFDDAFED0" box="[541,564,260,286]" count="1" pageId="29" pageNumber="30" type="male"></specimenCount>
, Agathis swamp for. Secondary vegetation,
<location id="8E92FFEAFFAF0A4CF939FE98FB89FED3" LSID="urn:lsid:plazi:treatment:03E41827FFAF0A4CFD8EFF0CFEC2FA2B:8E92FFEAFFAF0A4CF939FE98FB89FED3" box="[1056,1127,259,285]" country="Brunei" name="Badas" pageId="29" pageNumber="30">Badas</location>
,
<quantity id="4CB504D4FFAF0A4CF96BFE98FB0BFED0" box="[1138,1253,259,286]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="29" pageNumber="30" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
<elevation id="00604E02FFAF0A4CF96BFE98FB0BFED0" box="[1138,1253,259,286]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="29" pageNumber="30" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50100 m</elevation>
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,
<date id="FFF38FF1FFAF0A4CF9E9FE98FA89FED3" box="[1264,1383,259,285]" pageId="29" pageNumber="30" value="1982-02-27">
<collectingDate id="EFB77619FFAF0A4CF9E9FE98FA89FED3" box="[1264,1383,259,285]" pageId="29" pageNumber="30" value="1982-02-27">27.II.1982</collectingDate>
</date>
, leg.
<collectorName id="26B8CCE7FFAF0A4CFD8EFEBCFED0FE8F" box="[151,318,295,321]" pageId="29" pageNumber="30">G.S. Robinson</collectorName>
, slide No.
<specimenCode id="DBEB014AFFAF0A4CFCACFEBCFD4FFE8F" box="[437,673,295,321]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="30" type="Museum">NHMUK010316990</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF2A931FFAF0A4CFDDEFED7FD4BFE44" blockId="29.[151,1437,259,1509]" pageId="29" pageNumber="30">
<materialsCitation id="3B25A36CFFAF0A4CFDDEFED7FD4FFE44" ID-GBIF-Occurrence="4977736472" collectingDate="1993-04-15" collectingDateMax="1993-04-20" collectingDateMin="1993-04-15" collectionCode="NHMUK" collectorName="G. S. Robinson " country="Brunei" county="Lamunin" elevation="60" location="Lamunin" municipality="Disturbed" pageId="29" pageNumber="30" specimenCode="NHMUK010317159" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54F61793FFAF0A4CFDDEFED7FEC3FEA8" box="[199,301,332,358]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
.
<emphasis id="B9397523FFAF0A4CFC21FED0FE7AFEAB" bold="true" box="[312,404,331,357]" pageId="29" pageNumber="30">
<collectingCountry id="F35AE9A1FFAF0A4CFC21FED0FE61FEAB" box="[312,399,331,357]" name="Brunei" pageId="29" pageNumber="30">Brunei</collectingCountry>
:
</emphasis>
<specimenCount id="9D4B62B8FFAF0A4CFC85FED0FE2EFEAB" box="[412,448,331,357]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
, Sg Burong water tanks Disturbed lowland forest,
<collectingCounty id="6293D1BDFFAF0A4CF918FED0FB87FEAB" box="[1025,1129,331,357]" pageId="29" pageNumber="30">Lamunin</collectingCounty>
,
<quantity id="4CB504D4FFAF0A4CF96CFED0FB5FFEA8" box="[1141,1201,331,358]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="m" value="60.0">
<elevation id="00604E02FFAF0A4CF96CFED0FB5FFEA8" box="[1141,1201,331,358]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="m" value="60.0">60 m</elevation>
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,
<date id="FFF38FF1FFAF0A4CF9A7FED0FA89FEA8" box="[1214,1383,331,358]" pageId="29" pageNumber="30" value="1993-04-15" valueMax="1993-04-20" valueMin="1993-04-15">
<collectingDate id="EFB77619FFAF0A4CF9A7FED0FA89FEA8" box="[1214,1383,331,358]" pageId="29" pageNumber="30" value="1993-04-15" valueMax="1993-04-20" valueMin="1993-04-15">1520.IV.1993</collectingDate>
</date>
, leg. G.S. Robinson, slide No.
<specimenCode id="DBEB014AFFAF0A4CFCACFEF4FD4FFE44" box="[437,673,367,394]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="30" type="Museum">NHMUK010317159</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFAF0A4CFDDEFE08FDDBFE3B" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8BF2A931FFAF0A4CFDDEFE08FDDBFE3B" blockId="29.[151,1437,259,1509]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFDDEFE08FEACFE63" bold="true" box="[199,322,403,429]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
The new species can be distinguished by having a large foxtail-like gnathos, the valva with two apical processes formed by the apex deeply concave medially, and the lateral arm of the tegumen angled outward at the anterior 1/
<quantity id="4CB504D4FFAF0A4CFC2DFE40FEB1FE3B" box="[308,351,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="29" pageNumber="30" unit="in" value="3.0">3 in</quantity>
the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFAF0A4CFDDEFE64FE7FFAB7" pageId="29" pageNumber="30" type="description">
<paragraph id="8BF2A931FFAF0A4CFDDEFE64FC8CFDD4" blockId="29.[151,1437,259,1509]" box="[199,866,511,538]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFDDEFE64FEBFFDD7" bold="true" box="[199,337,511,537]" pageId="29" pageNumber="30">Description</emphasis>
. Adult (
<figureCitation id="1376B5B4FFAF0A4CFCB4FE64FDECFDD7" box="[429,514,511,537]" captionStart="FIGURES 3033" captionStartId="30.[151,264,1046,1070]" captionTargetBox="[174,1413,194,1009]" captionTargetId="figure-23@30.[162,1425,181,1021]" captionTargetPageId="30" captionText="FIGURES 3033. Adults of Promalactis spp. 30. P. spinellosa sp. nov.; 31. P. subfalcata sp. nov.; 32. P. trispinata sp. nov.; 33. P. varilonga sp. nov. 3032: Males and holotypes; 33: Male and paratype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023843" httpUri="https://zenodo.org/record/14023843/files/figure.png" pageId="29" pageNumber="30">Fig. 30</figureCitation>
). Forewing length 5.0
<quantity id="4CB504D4FFAF0A4CFE1FFE64FCB0FDD4" box="[774,862,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="29" pageNumber="30" unit="mm" value="5.5">5.5 mm</quantity>
.
</paragraph>
<paragraph id="8BF2A931FFAF0A4CFDDEFDB8FBE1FD4B" blockId="29.[151,1437,259,1509]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFDDEFDB8FEEAFDF3" box="[199,260,547,573]" italics="true" pageId="29" pageNumber="30">Head</emphasis>
. Vertex snowy white, frons silvery grey, occiput rust brown. Labial palpus with second segment brown; third segment black, white at base and apex. Antennal scape white dorsally, pale grey ventrally; flagellum black, alternating with white on dorsal surface except basal three flagellomeres white.
</paragraph>
<paragraph id="8BF2A931FFAF0A4CFDDEFD14FA82FC20" blockId="29.[151,1437,259,1509]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFDDEFD14FEF6FD67" box="[199,280,655,681]" italics="true" pageId="29" pageNumber="30">Thorax</emphasis>
. Mesonotum and tegula ochreous brown, thorax with a white horizontal band apically. Forewing ochreous brown; markings white edged with black scales: costal spot sub-rounded, extending to above posterior angle of cell; basal fascia from dorsum oblique inward to base of costal margin; antemedian fascia from basal 3/5 of dorsum to basal 1/4 of costal margin, parallel with basal fascia, widened anteriorly; dorsal streak from before tornus oblique outward to posterior angle of cell; apical spot ovoid; fringe blackish grey except white on extension of apical spot. Hindwing and fringe grey. Foreleg black except femur grey ventrally, tibia with white spot at base and middle, with a tuft of white scales at apex, tarsus white at apex of basal tarsomere and on entire second tarsomere; midleg brown ventrally, black dorsally, tibia with a white spot at base, with a tuft of white scales at apex, tarsus white at apex of basal tarsomere and on entire second and apical tarsomeres; hindleg yellowish brown ventrally, black dorsally, tarsus white at apices of basal and third tarsomeres, on entire second and distal two tarsomeres.
</paragraph>
<paragraph id="8BF2A931FFAF0A4CFDDEFC6CFC13FA98" blockId="29.[151,1437,259,1509]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFDDEFC6CFE9FFBDF" box="[199,369,1015,1041]" italics="true" pageId="29" pageNumber="30">Male genitalia</emphasis>
(
<figureCitation id="1376B5B4FFAF0A4CFC9FFC6CFE31FBDF" box="[390,479,1015,1041]" captionStart="FIGURES 5863" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[232,1359,193,1907]" captionTargetId="figure-23@36.[220,1368,181,1919]" captionTargetPageId="36" captionText="FIGURES 5863. Male genitalia of Promalactis spp. 58. P. minuticornuta sp. nov.; 59. P. parvissima sp. nov.; 60. P. polyptycha sp. nov.; 61. P. indica sp. nov.; 62. P. similidierli sp. nov.; 63. P. spinellosa sp. nov. All holotypes. Scale bar: 58, 60, 6263 = 0.2 mm; 59, 61 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14023857" httpUri="https://zenodo.org/record/14023857/files/figure.png" pageId="29" pageNumber="30">Fig. 63</figureCitation>
). Uncus relatively small, elongate rhombic, slightly widened medially, shallowly concave at middle apically. Gnathos longer than uncus, foxtail-like, fused with tegumen basally, covered with dense spines. Tegumen cylindrical; lateral arm widened and angled outward at distal 1/3, then narrowed to anterior end. Valva subparallel; apex deeply concave, forming two apical processes: dorsoapical process wider, bilobed distally, dorsal lobe horn-shaped, shorter, ventral lobe digitate, twice as long as dorsal lobe; ventroapical process slender and shorter, narrowed to acute apex, setose; costa slightly concave medially; left sacculus arched dorsally, right sacculus concave dorsally at distal 1/3. Saccus clavate, wide at base, narrowed to middle, then uniformly wide to rounded apex. Juxta elongate sub-trapezoidal, broad at base, slightly narrowed to apex, apex deeply incised in middle, pointed at apex; basal lobe slender, short. Phallus longer than valva; with two cornuti, longer cornutus shorter than 1/2 length of phallus, shorter cornutus less than 1/4 length of longer cornutus.
</paragraph>
<paragraph id="8BF2A931FFAF0A4CFDDEFAC4FE7FFAB7" blockId="29.[151,1437,259,1509]" box="[199,401,1375,1401]" pageId="29" pageNumber="30">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFAF0A4CFDDEFA18FE55FA53" box="[199,443,1411,1437]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BF2A931FFAF0A4CFDDEFA18FE55FA53" blockId="29.[151,1437,259,1509]" box="[199,443,1411,1437]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFDDEFA18FE8FFA53" bold="true" box="[199,353,1411,1437]" pageId="29" pageNumber="30">Distribution.</emphasis>
<collectingCountry id="F35AE9A1FFAF0A4CFC7EFA18FE59FA53" box="[359,439,1411,1437]" name="Brunei" pageId="29" pageNumber="30">Brunei</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFAF0A4CFDDEFA3CFEC2FA2B" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8BF2A931FFAF0A4CFDDEFA3CFEC2FA2B" blockId="29.[151,1437,259,1509]" pageId="29" pageNumber="30">
<emphasis id="B9397523FFAF0A4CFDDEFA3CFEA0FA0F" bold="true" box="[199,334,1447,1473]" pageId="29" pageNumber="30">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis id="B9397523FFAF0A4CFE71FA3CFC0AFA0F" box="[872,996,1447,1473]" italics="true" pageId="29" pageNumber="30">spinellosus</emphasis>
, referring to the gnathos covered with dense spines.
</paragraph>
</subSubSection>
</treatment>
</document>