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<document id="DB6E91A8D803F2D5F88D106DFED6DD28" ID-CLB-Dataset="305079" ID-DOI="10.11646/zootaxa.5536.1.1" ID-GBIF-Dataset="52a66b9d-a4c0-45a2-9e5e-7c15685a8f39" ID-ISSN="1175-5326" ID-Zenodo-Dep="14023827" ID-ZooBank="C47EE4D9-D5B5-4B67-8547-856FBAEFE670" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1730455362568" checkinUser="plazi" docAuthor="Wang, Shuxia, Liu, Chen &amp; David, Lees" docDate="2024" docId="03E41827FFA80A4AFD8EFB31FA6FFD03" docLanguage="en" docName="zootaxa.5536.1.1.pdf" docOrigin="Zootaxa 5536 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Promalactis polyptycha Wang &amp; Liu &amp; David 2024, sp. nov." docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="FFDD605FFFB20A51FD19FF9BFFEEFFCE" masterDocTitle="Partial revision of Promalactis Meyrick (Lepidoptera: Oecophoridae), based on specimens deposited in the Natural History Museum, London" masterLastPageNumber="58" masterPageNumber="1" pageNumber="27" updateTime="1732577891658" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9D6F9A90379A3AB9D7C03FBE9E736434">Partial revision of Promalactis Meyrick (Lepidoptera: Oecophoridae), based on specimens deposited in the Natural History Museum, London</mods:title>
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<mods:affiliation id="A776C1FEAA88C5561AC3F81D50AD1110">College of Life Sciences, Nankai University, Tianjin 300071, China</mods:affiliation>
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<mods:namePart id="EA19FE3A63B0A26A2AA5D0BCD4C47046">David, Lees</mods:namePart>
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<mods:affiliation id="EAA853DD20230ED01BDE07191DAAEDDE">Department of Science, Natural History Museum, London, United Kingdom, SW 7 5 BD</mods:affiliation>
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<treatment id="03E41827FFA80A4AFD8EFB31FA6FFD03" ID-DOI="http://doi.org/10.5281/zenodo.14023901" ID-GBIF-Taxon="246064895" ID-Zenodo-Dep="14023901" LSID="urn:lsid:plazi:treatment:03E41827FFA80A4AFD8EFB31FA6FFD03" httpUri="http://treatment.plazi.org/id/03E41827FFA80A4AFD8EFB31FA6FFD03" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" scope_family="Oecophoridae" scope_order="Lepidoptera">
<subSubSection id="C357FABAFFA80A4BFD8EFB31FD80FB0B" box="[151,622,1194,1221]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BF2A931FFA80A4BFD8EFB31FD80FB0B" blockId="26.[151,622,1194,1257]" box="[151,622,1194,1221]" pageId="26" pageNumber="27">
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<emphasis id="B9397523FFA80A4BFD8EFB31FD80FB0B" bold="true" box="[151,622,1194,1221]" pageId="26" pageNumber="27">
<taxonomicName id="4C4DD2B2FFA80A4BFD8EFB31FDEAFB0B" ID-CoL="CN7QC" authority="Wang" authorityName="Wang &amp; Liu &amp; David" authorityYear="2024" box="[151,516,1194,1221]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="polyptycha" status="sp. nov.">
<emphasis id="B9397523FFA80A4BFD8EFB31FE5CFB0B" bold="true" box="[151,434,1194,1221]" italics="true" pageId="26" pageNumber="27">Promalactis polyptycha</emphasis>
Wang
</taxonomicName>
,
<taxonomicNameLabel id="A20AC858FFA80A4BFF08FB30FD80FB0B" box="[529,622,1195,1221]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C357FABAFFA80A4BFD8EFB54FEC9FB27" box="[151,295,1231,1257]" pageId="26" pageNumber="27" type="description">
<paragraph id="8BF2A931FFA80A4BFD8EFB54FEC9FB27" blockId="26.[151,622,1194,1257]" box="[151,295,1231,1257]" pageId="26" pageNumber="27">
(
<figureCitation id="1376B5B4FFA80A4BFD86FB54FF18FB27" box="[159,246,1231,1257]" captionStart="FIGURES 2429" captionStartId="24.[151,264,1590,1614]" captionTargetBox="[174,1416,321,1556]" captionTargetId="figure-49@24.[159,1428,309,1566]" captionTargetPageId="24" captionText="FIGURES 2429. Adults of Promalactis spp. 24. P. magnisaccata sp. nov.; 25. P. minuticornuta sp. nov.; 26. P. parvissima sp. nov. (female and paratype); 27. P. polyptycha sp. nov.; 28. P. indica sp. nov.; 29. P. similidierli sp. nov. 2425, 2729: Males and holotypes; 26: Male and paratype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023839" httpUri="https://zenodo.org/record/14023839/files/figure.png" pageId="26" pageNumber="27">Figs 27</figureCitation>
,
<figureCitation id="1376B5B4FFA80A4BFC1BFB54FEF1FB27" box="[258,287,1231,1257]" captionStart="FIGURES 5863" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[232,1359,193,1907]" captionTargetId="figure-23@36.[220,1368,181,1919]" captionTargetPageId="36" captionText="FIGURES 5863. Male genitalia of Promalactis spp. 58. P. minuticornuta sp. nov.; 59. P. parvissima sp. nov.; 60. P. polyptycha sp. nov.; 61. P. indica sp. nov.; 62. P. similidierli sp. nov.; 63. P. spinellosa sp. nov. All holotypes. Scale bar: 58, 60, 6263 = 0.2 mm; 59, 61 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14023857" httpUri="https://zenodo.org/record/14023857/files/figure.png" pageId="26" pageNumber="27">60</figureCitation>
)
</paragraph>
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<subSubSection id="C357FABAFFA80A4BFD8EFA8CFD88FAB7" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BF2A931FFA80A4BFD8EFA8CFC83FA9B" blockId="26.[151,1437,1302,2013]" pageId="26" pageNumber="27">
<emphasis id="B9397523FFA80A4BFD8EFA8CFEADFAFE" bold="true" box="[151,323,1302,1329]" pageId="26" pageNumber="27">Type material.</emphasis>
<materialsCitation id="3B25A36CFFA80A4BFC51FA8DFC83FA9B" ID-GBIF-Occurrence="4977736575" collectingDate="1991-11-05" collectingDateMax="1991-11-10" collectingDateMin="1991-11-05" collectionCode="NHMUK" collectorName="I. J. Kitching &amp; A. M. Cotton" country="Thailand" elevation="900" location="Phu Khieo WS" pageId="26" pageNumber="27" specimenCode="NHMUK010317019" specimenCount="1" specimenCount-male="1" stateProvince="Chaiyaphum" typeStatus="holotype">
<emphasis id="B9397523FFA80A4BFC51FA8DFE50FAFE" bold="true" box="[328,446,1302,1328]" pageId="26" pageNumber="27">
<collectingCountry id="F35AE9A1FFA80A4BFC51FA8DFE57FAFE" box="[328,441,1302,1328]" name="Thailand" pageId="26" pageNumber="27">Thailand</collectingCountry>
:
</emphasis>
<typeStatus id="54F61793FFA80A4BFCDAFA8CFDC5FAFF" box="[451,555,1303,1329]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4B62B8FFA80A4BFF29FA8CFDA9FAFF" box="[560,583,1303,1329]" count="1" pageId="26" pageNumber="27" type="male"></specimenCount>
,
<locationDeviation id="9A40B013FFA80A4BFF48FA8CFCBAFAFF" box="[593,852,1302,1329]" pageId="26" pageNumber="27">Plateau 500 m N of HQ</locationDeviation>
,
<location id="8E92FFEAFFA80A4BFE44FA8CFBEAFAFF" LSID="urn:lsid:plazi:treatment:03E41827FFA80A4AFD8EFB31FA6FFD03:8E92FFEAFFA80A4BFE44FA8CFBEAFAFF" box="[861,1028,1303,1329]" country="Thailand" name="Phu Khieo WS" pageId="26" pageNumber="27" stateProvince="Chaiyaphum">Phu Khieo WS</location>
,
<collectingRegion id="498967D3FFA80A4BF914FA8CFB71FAFF" box="[1037,1183,1303,1329]" country="Thailand" name="Chaiyaphum" pageId="26" pageNumber="27">Chaiyaphum</collectingRegion>
, 900 m,
<date id="FFF38FF1FFA80A4BF9E3FA8DFA76FAFF" box="[1274,1432,1302,1329]" pageId="26" pageNumber="27" value="1991-11-05" valueMax="1991-11-10" valueMin="1991-11-05">
<collectingDate id="EFB77619FFA80A4BF9E3FA8DFA76FAFF" box="[1274,1432,1302,1329]" pageId="26" pageNumber="27" value="1991-11-05" valueMax="1991-11-10" valueMin="1991-11-05">510.XI.1991</collectingDate>
</date>
, leg.
<collectorName id="26B8CCE7FFA80A4BFDDEFAA0FEBDFA9B" box="[199,339,1339,1365]" pageId="26" pageNumber="27">I.J. Kitching</collectorName>
&amp;
<collectorName id="26B8CCE7FFA80A4BFC6CFAA0FDE8FA9B" box="[373,518,1339,1365]" pageId="26" pageNumber="27">A.M. Cotton</collectorName>
, slide No.
<specimenCode id="DBEB014AFFA80A4BFF64FAA0FC87FA9B" box="[637,873,1339,1365]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="26" pageNumber="27" type="Museum">NHMUK010317019</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF2A931FFA80A4BFDDEFAC4FD88FAB7" blockId="26.[151,1437,1302,2013]" box="[199,614,1375,1401]" pageId="26" pageNumber="27">
<materialsCitation id="3B25A36CFFA80A4BFDDEFAC4FD8DFAB7" ID-GBIF-Occurrence="4977736534" box="[199,611,1375,1401]" collectingDate="1991-11-05" collectingDateMax="1991-11-10" collectingDateMin="1991-11-05" collectionCode="NHMUK" collectorName="Phu Khieo WS &amp; I. J. Kitching &amp; A. M. Cotton &amp; No." country="Thailand" elevation="500" location="Plateau" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Chaiyaphum" typeStatus="paratype">
<typeStatus id="54F61793FFA80A4BFDDEFAC4FEC3FAB7" box="[199,301,1375,1401]" pageId="26" pageNumber="27" type="paratype">Paratype</typeStatus>
.
<specimenCount id="9D4B62B8FFA80A4BFC2EFAC4FEB2FAB7" box="[311,348,1375,1401]" count="1" pageId="26" pageNumber="27" type="male">1♂</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA80A4BFDDEFA19FBD7F9E3" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BF2A931FFA80A4BFDDEFA19FBD7F9E3" blockId="26.[151,1437,1302,2013]" pageId="26" pageNumber="27">
<emphasis id="B9397523FFA80A4BFDDEFA19FEACFA52" bold="true" box="[199,322,1410,1436]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
The new species can be distinguished from other known species of the group by the valva produced into a bird head-shaped process distally and the phallus with many pleats distally. It is similar to
<taxonomicName id="4C4DD2B2FFA80A4BF9E7FA3CFECFFA2B" authority="Wang, 2018" authorityName="Wang" authorityYear="2018" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="apicuncata">
<emphasis id="B9397523FFA80A4BF9E7FA3CFAE1FA0E" box="[1278,1295,1447,1472]" italics="true" pageId="26" pageNumber="27">P</emphasis>
.
<emphasis id="B9397523FFA80A4BF839FA3CFA72FA0E" box="[1312,1436,1447,1472]" italics="true" pageId="26" pageNumber="27">apicuncata</emphasis>
<bibRefCitation id="EFDCD4C0FFA80A4BFD8EFA50FECFFA2B" author="Wang, S. X. &amp; Jia, Y. Y." box="[151,289,1483,1509]" pageId="26" pageNumber="27" pagination="151 - 160" refId="ref37689" refString="Wang, S. X. &amp; Jia, Y. Y. (2018) Review of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) III. The trilineata species group, with descriptions of four new species. Zootaxa, 4433 (1), 151 - 160. https: // doi. org / 10.11646 / zootaxa. 4433.1.9" type="journal article" year="2018">Wang, 2018</bibRefCitation>
</taxonomicName>
, but can be distinguished by the gnathos with a sub-digitate mesial plate, the costa concave beyond the middle, and the juxta without distinct lateral lobes. In
<taxonomicName id="4C4DD2B2FFA80A4BFFF6FA74FC64F9C6" authorityName="Wang" authorityYear="2018" box="[751,906,1519,1545]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="apicuncata">
<emphasis id="B9397523FFA80A4BFFF6FA74FCEEF9C6" box="[751,768,1519,1544]" italics="true" pageId="26" pageNumber="27">P</emphasis>
.
<emphasis id="B9397523FFA80A4BFE17FA74FC64F9C6" box="[782,906,1519,1544]" italics="true" pageId="26" pageNumber="27">apicuncata</emphasis>
</taxonomicName>
, the gnathos has a sub-trapezoidal mesial plate, the costa is concave before middle, and the juxta has elongate slender lateral lobes.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA80A4AFDDEF9ADFE7FFD4B" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BF2A931FFA80A4BFDDEF9ADFC8CF99F" blockId="26.[151,1437,1302,2013]" box="[199,866,1590,1617]" pageId="26" pageNumber="27">
<emphasis id="B9397523FFA80A4BFDDEF9ADFEBFF99E" bold="true" box="[199,337,1590,1616]" pageId="26" pageNumber="27">Description</emphasis>
. Adult (
<figureCitation id="1376B5B4FFA80A4BFCB4F9ACFDECF99F" box="[429,514,1591,1617]" captionStart="FIGURES 2429" captionStartId="24.[151,264,1590,1614]" captionTargetBox="[174,1416,321,1556]" captionTargetId="figure-49@24.[159,1428,309,1566]" captionTargetPageId="24" captionText="FIGURES 2429. Adults of Promalactis spp. 24. P. magnisaccata sp. nov.; 25. P. minuticornuta sp. nov.; 26. P. parvissima sp. nov. (female and paratype); 27. P. polyptycha sp. nov.; 28. P. indica sp. nov.; 29. P. similidierli sp. nov. 2425, 2729: Males and holotypes; 26: Male and paratype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023839" httpUri="https://zenodo.org/record/14023839/files/figure.png" pageId="26" pageNumber="27">Fig. 27</figureCitation>
). Forewing length 4.0
<quantity id="4CB504D4FFA80A4BFE1FF9ADFCB0F99F" box="[774,862,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="26" pageNumber="27" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
<paragraph id="8BF2A931FFA80A4BFDDEF9C0FF19F92F" blockId="26.[151,1437,1302,2013]" pageId="26" pageNumber="27">
<emphasis id="B9397523FFA80A4BFDDEF9C0FEEAF9BB" box="[199,260,1627,1653]" italics="true" pageId="26" pageNumber="27">Head</emphasis>
. Vertex snowy white; frons brown; occiput ochreous brown, with tufted yellow scales laterally. Labial palpus with second segment rust brown on outer surface, orange yellow on inner surface; third segment black except white apically. Antennal scape white dorsally, greyish white ventrally; flagellum dark brown, alternating with white dorsally.
</paragraph>
<paragraph id="8BF2A931FFA80A4AFDDEF970FD9FFED3" blockId="26.[151,1437,1302,2013]" lastBlockId="27.[151,1437,151,717]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="B9397523FFA80A4BFDDEF970FEF6F8CB" box="[199,280,1771,1797]" italics="true" pageId="26" pageNumber="27">Thorax</emphasis>
. Mesonotum and tegula ochreous brown. Forewing orange yellow; markings white edged with black scales: costal spot triangular, from distal 1/3 oblique outward to above posterior angle of cell, sinuate laterally; basal streak from dorsum to above base of fold; antemedian streak from middle of dorsum to basal 2/5 of anterior margin of cell, slightly curved outward, subparallel with basal streak; dorsal streak finely slender, from before end of fold oblique outward and reaching inner margin of tornal spot; tornal spot large, ovate, reaching below posterior angle of cell; apical spot ovate; termen with two small spots below costal spot, interrupted by black scales; fringe yellow. Hindwing and fringe grey. Foreleg black except femur greyish white dorsally, tibia with white spot at base and middle, with a tuft of white scales at apex, tarsus white at apices of basal two tarsomeres; midleg brown ventrally, black dorsally, tibia with a white spot at base, with a tuft of white scales at apex, tarsus white at apices of basal two and apical tarsomeres; hindleg yellowish brown ventrally, black dorsally, tarsus white at apices of basal four tarsomeres and on entire apical tarsomere.
</paragraph>
<paragraph id="8BF2A931FFA90A4AFDDEFEBCFDB5FDAF" blockId="27.[151,1437,151,717]" pageId="27" pageNumber="28">
<emphasis id="B9397523FFA90A4AFDDEFEBCFE84FE8F" box="[199,362,295,321]" italics="true" pageId="27" pageNumber="28">Male genitalia</emphasis>
(
<figureCitation id="1376B5B4FFA90A4AFC6FFEBCFE27FE8F" box="[374,457,295,321]" captionStart="FIGURES 5863" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[232,1359,193,1907]" captionTargetId="figure-23@36.[220,1368,181,1919]" captionTargetPageId="36" captionText="FIGURES 5863. Male genitalia of Promalactis spp. 58. P. minuticornuta sp. nov.; 59. P. parvissima sp. nov.; 60. P. polyptycha sp. nov.; 61. P. indica sp. nov.; 62. P. similidierli sp. nov.; 63. P. spinellosa sp. nov. All holotypes. Scale bar: 58, 60, 6263 = 0.2 mm; 59, 61 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14023857" httpUri="https://zenodo.org/record/14023857/files/figure.png" pageId="27" pageNumber="28">Fig. 60</figureCitation>
). Uncus wide at base, narrowed to distal 1/3, thereafter uniformly slender to apex, heavily sclerotized distally and laterally. Gnathos longer than uncus; mesial plate short, sub-digitate, rounded apically; basal arm widely band-like, longer than mesial plate. Tegumen furcate from near posterior margin; lateral arm triangular, narrowed to anterior end. Valva with basal 2/3 parallel sided, distal 1/3 slightly narrower, produced into a broad bird head-shaped process acute at apex and finely serrate on dorsal margin; costa concave beyond middle; sacculus slightly concave beyond middle dorsally, distal 2/5 setose, produced into a free process narrowed to pointed apex and curved dorsad. Saccus small, triangular, narrowed to rounded apex, shorter than uncus. Juxta shorter than uncus, U-shaped, produced anteriorly. Phallus as long as valva, with dense longitudinal pleats distally; cornutus spine-shaped, with tiny spines around its base.
</paragraph>
<paragraph id="8BF2A931FFA90A4AFDDEFDF0FE7FFD4B" blockId="27.[151,1437,151,717]" box="[199,401,619,645]" pageId="27" pageNumber="28">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA90A4AFDDEFD15FE3CFD67" box="[199,466,654,681]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BF2A931FFA90A4AFDDEFD15FE3CFD67" blockId="27.[151,1437,151,717]" box="[199,466,654,681]" pageId="27" pageNumber="28">
<emphasis id="B9397523FFA90A4AFDDEFD15FE8FFD66" bold="true" box="[199,353,654,680]" pageId="27" pageNumber="28">Distribution.</emphasis>
<collectingCountry id="F35AE9A1FFA90A4AFC7EFD14FE20FD67" box="[359,462,655,681]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA90A4AFDDEFD28FA6FFD03" box="[199,1409,691,717]" pageId="27" pageNumber="28" type="etymology">
<paragraph id="8BF2A931FFA90A4AFDDEFD28FA6FFD03" blockId="27.[151,1437,151,717]" box="[199,1409,691,717]" pageId="27" pageNumber="28">
<emphasis id="B9397523FFA90A4AFDDEFD28FEA0FD03" bold="true" box="[199,334,691,717]" pageId="27" pageNumber="28">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis id="B9397523FFA90A4AFE4FFD28FC32FD03" box="[854,988,691,717]" italics="true" pageId="27" pageNumber="28">polyptychus</emphasis>
, referring to the pleats of the phallus.
</paragraph>
</subSubSection>
</treatment>
</document>