treatments-xml/data/03/CC/87/03CC87A6FF88FFFD3F847656FD9D706D.xml
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<document id="E510E565755B8AE11048FB30B47B18E9" ID-CLB-Dataset="3276" ID-DOI="10.11646/zootaxa.5072.1.9" ID-GBIF-Dataset="92e598fc-f5fe-45c4-9020-cab8bab17668" ID-ISSN="1175-5326" ID-Zenodo-Dep="5729038" ID-ZooBank="9080121B-76CC-44BA-9F85-A9C35329388F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637914303197" checkinUser="plazi" docAuthor="Liang, Feiyang &amp; Liu, Xingyue" docDate="2021" docId="03CC87A6FF88FFFD3F847656FD9D706D" docLanguage="en" docName="zootaxa.5072.1.9.pdf" docOrigin="Zootaxa 5072 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psyllipsocus yangi Liang &amp; Liu 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="86" masterDocId="FFF5FFDEFF89FFF83F137537FFE1712C" masterDocTitle="A new species of Psyllipsocus (Psocodea: Trogiomorpha: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar" masterLastPageNumber="87" masterPageNumber="81" pageNumber="82" updateTime="1699470356438" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2DB5B1D22C18E6029314CC95538F93D8">A new species of Psyllipsocus (Psocodea: Trogiomorpha: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar</mods:title>
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<mods:namePart id="1B19E65881C84F61530EC401CB60FC6D">Liang, Feiyang</mods:namePart>
<mods:affiliation id="EA1A0B96379522A0735C63A7F5DBBA1D">Key Laboratory of Economic Crops Genetic Improvement and Integrated Utilization, School of Life Science, Hunan University of Science and Technology, Xiangtan 411201, China.</mods:affiliation>
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<mods:namePart id="31076B195A50C646B9EBB6E6A27BA014">Liu, Xingyue</mods:namePart>
<mods:affiliation id="68BF7C8DAF62DFCCB56BA2686F03C8DA">Department of Entomology, China Agricultural University, Beijing 100193, China.</mods:affiliation>
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<treatment id="03CC87A6FF88FFFD3F847656FD9D706D" ID-DOI="http://doi.org/10.5281/zenodo.5729040" ID-GBIF-Taxon="190872015" ID-Zenodo-Dep="5729040" LSID="urn:lsid:plazi:treatment:03CC87A6FF88FFFD3F847656FD9D706D" httpUri="http://treatment.plazi.org/id/03CC87A6FF88FFFD3F847656FD9D706D" lastPageId="5" lastPageNumber="86" pageId="1" pageNumber="82">
<subSubSection id="C37F653BFF88FFF93F847656FE5E7257" box="[151,447,865,892]" pageId="1" pageNumber="82" type="nomenclature">
<paragraph id="8BDA36B0FF88FFF93F847656FE5E7257" blockId="1.[151,447,865,928]" box="[151,447,865,892]" pageId="1" pageNumber="82">
<heading id="D09281DCFF88FFF93F847656FE5E7257" bold="true" box="[151,447,865,892]" fontSize="11" level="1" pageId="1" pageNumber="82" reason="1">
<emphasis id="B911EAA2FF88FFF93F847656FE5E7257" bold="true" box="[151,447,865,892]" pageId="1" pageNumber="82">
<taxonomicName id="4C654D33FF88FFF93F847656FE977257" authority="Liang &amp; Liu, 2021" authorityName="Liang &amp; Liu" authorityYear="2021" box="[151,374,865,892]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov.">
<emphasis id="B911EAA2FF88FFF93F847656FE977257" bold="true" box="[151,374,865,892]" italics="true" pageId="1" pageNumber="82">Psyllipsocus yangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257D9FF88FFF93E6E7656FE5E7257" box="[381,447,865,891]" pageId="1" pageNumber="82" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F653BFF88FFF93F8476B2FEF8728C" box="[151,281,901,928]" pageId="1" pageNumber="82" type="description">
<paragraph id="8BDA36B0FF88FFF93F8476B2FEF8728C" blockId="1.[151,447,865,928]" box="[151,281,901,928]" pageId="1" pageNumber="82">
(
<figureCitation id="135E2A35FF88FFF93F8C76B2FEF1728C" box="[159,272,901,928]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 48" captionStart-2="FIGURES 915" captionStartId-0="2.[151,264,1234,1258]" captionStartId-1="3.[151,264,1681,1705]" captionStartId-2="4.[151,264,1612,1636]" captionTargetBox-0="[167,1417,190,1200]" captionTargetBox-1="[198,1386,200,1643]" captionTargetBox-2="[156,1426,191,1579]" captionTargetId-0="figure-24@2.[160,1427,181,1211]" captionTargetId-1="figure-17@3.[193,1395,181,1657]" captionTargetId-2="figure-24@4.[154,1434,181,1588]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 13. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 1. photograph and drawing of habitus (dorsal view), scale bar=0.5 mm; 2. photograph and drawing of habitus (ventral view), scale bar=0.5 mm; 3. fore- and hindwing, scale bar=0.5 mm." captionText-1="FIGURES 48. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 4. head (dorsal view); 5. maxillary palpus; 6. a conical sensillum on mx2; 7. photograph and drawing of terminalia (ventral view); 8. photograph and drawing of habitus (dorsal view). ep: epiproct; pp: paraproct; ev: external valve; sg: subgenital plate." captionText-2="FIGURES 915. Psyllipsocus yangi sp. n., Paratype CAU-BA-LFY-21002. 9. habitus; 10. mouthparts; 11. forewing; 12. hindwing; 13. in-flight wing-coupling structure; 14. labial palps; 15. lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia." figureDoi-0="http://doi.org/10.5281/zenodo.5729042" figureDoi-1="http://doi.org/10.5281/zenodo.5729052" figureDoi-2="http://doi.org/10.5281/zenodo.5729058" httpUri-0="https://zenodo.org/record/5729042/files/figure.png" httpUri-1="https://zenodo.org/record/5729052/files/figure.png" httpUri-2="https://zenodo.org/record/5729058/files/figure.png" pageId="1" pageNumber="82">Figs 115</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F653BFF88FFF93F8476FAFDEB7593" pageId="1" pageNumber="82" type="diagnosis">
<paragraph id="8BDA36B0FF88FFF93F8476FAFDEB7593" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="82">
<emphasis id="B911EAA2FF88FFF93F8476FAFEF372CB" bold="true" box="[151,274,973,999]" pageId="1" pageNumber="82">Diagnosis.</emphasis>
Macropterous. Ocelli present. Antenna filiform, longer than body, antennae 20-segmented at least, flagellum bearing setae, with secondary annulation. Postclypeus gibbous; mx2 with a conical sensillum, mx4 distally broadened and with flat apex; lacinia developed, with three apical teeth; labial palps with two segments, terminal segment rounded. Wings transparent with complete venation. Forewing margin almost glabrous; veins with one row of setae; pterostigma with four sides, but nearly triangular; R
<subScript id="17E134F5FF88FFF93C537147FCA97552" attach="left" box="[832,840,1136,1150]" fontSize="6" pageId="1" pageNumber="82">1</subScript>
twice the length of Sc. Hindwing with a quadrangular basiradial cell and bifurcate A. Female: Genitalia strongly sclerotized; epiproct subtriangular with round apex; paraproct with 78 trichobothria.
</paragraph>
</subSubSection>
<subSubSection id="C37F653BFF88FFFA3FD471FEFEB77727" lastPageId="2" lastPageNumber="83" pageId="1" pageNumber="82" type="description">
<paragraph id="8BDA36B0FF88FFF93FD471FEFC82747C" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="82">
<emphasis id="B911EAA2FF88FFF93FD471FEFDD675CF" bold="true" box="[199,567,1225,1251]" pageId="1" pageNumber="82">
Description of
<typeStatus id="54DE8812FF88FFF93E6771FEFE3A75CF" box="[372,475,1225,1251]" pageId="1" pageNumber="82" type="holotype">holotype</typeStatus>
female.
</emphasis>
CAU-BA-LFY-21001 (
<figureCitation id="135E2A35FF88FFF93C5371FEFC4575CF" box="[832,932,1225,1251]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 48" captionStartId-0="2.[151,264,1234,1258]" captionStartId-1="3.[151,264,1681,1705]" captionTargetBox-0="[167,1417,190,1200]" captionTargetBox-1="[198,1386,200,1643]" captionTargetId-0="figure-24@2.[160,1427,181,1211]" captionTargetId-1="figure-17@3.[193,1395,181,1657]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 13. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 1. photograph and drawing of habitus (dorsal view), scale bar=0.5 mm; 2. photograph and drawing of habitus (ventral view), scale bar=0.5 mm; 3. fore- and hindwing, scale bar=0.5 mm." captionText-1="FIGURES 48. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 4. head (dorsal view); 5. maxillary palpus; 6. a conical sensillum on mx2; 7. photograph and drawing of terminalia (ventral view); 8. photograph and drawing of habitus (dorsal view). ep: epiproct; pp: paraproct; ev: external valve; sg: subgenital plate." figureDoi-0="http://doi.org/10.5281/zenodo.5729042" figureDoi-1="http://doi.org/10.5281/zenodo.5729052" httpUri-0="https://zenodo.org/record/5729042/files/figure.png" httpUri-1="https://zenodo.org/record/5729052/files/figure.png" pageId="1" pageNumber="82">Figs 18</figureCitation>
). Body length
<quantity id="4C9D9B55FF88FFF93B5671FEFB4675C8" box="[1093,1191,1225,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="1" pageNumber="82" unit="mm" value="1.56">1.56 mm</quantity>
(measured from frons to terminalia). IO:
<quantity id="4C9D9B55FF88FFF93E7471DAFE2C7424" box="[359,461,1261,1288]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="82" unit="mm" value="0.21">0.21 mm</quantity>
, d:
<quantity id="4C9D9B55FF88FFF93EE771DAFDB87424" box="[500,601,1261,1288]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="82" unit="mm" value="0.15">0.15 mm</quantity>
, IO/d = 1.4. f1:
<quantity id="4C9D9B55FF88FFF93C1971DAFC8E7424" box="[778,879,1261,1288]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="82" unit="mm" value="0.13">0.13 mm</quantity>
, f2:
<quantity id="4C9D9B55FF88FFF93C8C71DAFBE47424" box="[927,1029,1261,1288]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="82" unit="mm" value="0.15">0.15 mm</quantity>
, f3:
<quantity id="4C9D9B55FF88FFF93B2671DAFB6C7424" box="[1077,1165,1261,1288]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="82" unit="mm" value="0.1">0.1 mm</quantity>
. FWL:
<quantity id="4C9D9B55FF88FFF93BF271DAFAA77424" box="[1249,1350,1261,1288]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="1" pageNumber="82" unit="mm" value="1.56">1.56 mm</quantity>
, FWW:
<quantity id="4C9D9B55FF88FFF93F847026FF1E7400" box="[151,255,1297,1324]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="82" unit="mm" value="0.63">0.63 mm</quantity>
; HWL:
<quantity id="4C9D9B55FF88FFF93E487026FE237400" box="[347,450,1297,1324]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="1" pageNumber="82" unit="mm" value="1.19">1.19 mm</quantity>
, HWW:
<quantity id="4C9D9B55FF88FFF93D357026FD6F7400" box="[550,654,1297,1324]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="82" unit="mm" value="0.32">0.32 mm</quantity>
. ft1:
<quantity id="4C9D9B55FF88FFF93DDB7026FCCE7400" box="[712,815,1297,1324]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="82" unit="mm" value="0.13">0.13 mm</quantity>
, ft2:
<quantity id="4C9D9B55FF88FFF93C787026FC337400" box="[875,978,1297,1324]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="82" unit="mm" value="0.04">0.04 mm</quantity>
, ft3:
<quantity id="4C9D9B55FF88FFF93B1E7026FB947400" box="[1037,1141,1297,1324]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="82" unit="mm" value="0.05">0.05 mm</quantity>
; mt1:
<quantity id="4C9D9B55FF88FFF93BAF7026FAC27400" box="[1212,1315,1297,1324]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="82" unit="mm" value="0.15">0.15 mm</quantity>
, mt2:
<quantity id="4C9D9B55FF88FFF93A787026FF24747C" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="82" unit="mm" value="0.05">0.05 mm</quantity>
, mt3:
<quantity id="4C9D9B55FF88FFF93E187002FE93747C" box="[267,370,1333,1360]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="82" unit="mm" value="0.05">0.05 mm</quantity>
; ht1:
<quantity id="4C9D9B55FF88FFF93EA37002FDF7747C" box="[432,534,1333,1360]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="82" unit="mm" value="0.24">0.24 mm</quantity>
, ht2:
<quantity id="4C9D9B55FF88FFF93D477002FD5B747C" box="[596,698,1333,1360]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="82" unit="mm" value="0.04">0.04 mm</quantity>
, ht3:
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.
</paragraph>
<paragraph id="8BDA36B0FF88FFF93FD4706EFDCA772F" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="82">
Head with frontal and epicranial suture; compound eyes well developed; three ocelli disposed in triangle between compound eyes. Antennae 20-segmented at least, flagellum bearing setae, with secondary annulation. Maxillary palps four-segmented, mx2 as long as mx4 (
<figureCitation id="135E2A35FF88FFF93DC57096FCDC7490" box="[726,829,1441,1468]" captionStart="FIGURES 48" captionStartId="3.[151,264,1681,1705]" captionTargetBox="[198,1386,200,1643]" captionTargetId="figure-17@3.[193,1395,181,1657]" captionTargetPageId="3" captionText="FIGURES 48. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 4. head (dorsal view); 5. maxillary palpus; 6. a conical sensillum on mx2; 7. photograph and drawing of terminalia (ventral view); 8. photograph and drawing of habitus (dorsal view). ep: epiproct; pp: paraproct; ev: external valve; sg: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5729052" httpUri="https://zenodo.org/record/5729052/files/figure.png" pageId="1" pageNumber="82">Figs 45</figureCitation>
), mx2 with a conical sensillum (
<figureCitation id="135E2A35FF88FFF93BA77096FB1B7490" box="[1204,1274,1441,1468]" captionStart="FIGURES 48" captionStartId="3.[151,264,1681,1705]" captionTargetBox="[198,1386,200,1643]" captionTargetId="figure-17@3.[193,1395,181,1657]" captionTargetPageId="3" captionText="FIGURES 48. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 4. head (dorsal view); 5. maxillary palpus; 6. a conical sensillum on mx2; 7. photograph and drawing of terminalia (ventral view); 8. photograph and drawing of habitus (dorsal view). ep: epiproct; pp: paraproct; ev: external valve; sg: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5729052" httpUri="https://zenodo.org/record/5729052/files/figure.png" pageId="1" pageNumber="82">Fig. 6</figureCitation>
). Labial palps two-segmented, terminal segment rounded. Lacinia not visible. Cervical region distinct. Prothorax narrow and short, meso- and metathorax robust.
</paragraph>
<paragraph id="8BDA36B0FF88FFF93FD4733AFD6577F7" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="82">
Wings transparent, venation well developed, but asymmetric (
<figureCitation id="135E2A35FF88FFF93C96733AFC2B770B" box="[901,970,1549,1575]" captionStart="FIGURES 13" captionStartId="2.[151,264,1234,1258]" captionTargetBox="[167,1417,190,1200]" captionTargetId="figure-24@2.[160,1427,181,1211]" captionTargetPageId="2" captionText="FIGURES 13. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 1. photograph and drawing of habitus (dorsal view), scale bar=0.5 mm; 2. photograph and drawing of habitus (ventral view), scale bar=0.5 mm; 3. fore- and hindwing, scale bar=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5729042" httpUri="https://zenodo.org/record/5729042/files/figure.png" pageId="1" pageNumber="82">Fig. 3</figureCitation>
). Forewing with a row of setae on veins, hindwing without setae. Forewing Sc weak and short, ending free in the membrane; pterostigma not thickened, with four sides but similar to triangle; R
<subScript id="17E134F5FF88FFF93D3A735FFDD0775A" attach="left" box="[553,561,1640,1654]" fontSize="6" pageId="1" pageNumber="82">1</subScript>
twice the length of Sc; Cu2 and A meeting on wing margin; in-flight wingcoupling structure simple, composed of separate and almost straight spines; areola postica narrow and elongate; Cu
<subScript id="17E134F5FF88FFF93FAB7387FF297792" attach="left" box="[184,200,1712,1726]" fontSize="6" pageId="1" pageNumber="82">1b</subScript>
much shorter than Cu
<subScript id="17E134F5FF88FFF93EAC7387FE2F7792" attach="left" box="[447,462,1712,1726]" fontSize="6" pageId="1" pageNumber="82">1a</subScript>
; A simple. Hindwing with Sc reduced; basiradial cell quadrangular; Rs and M with two branches, M fork much longer than Rs fork.
</paragraph>
<paragraph id="8BDA36B0FF88FFF93FD473D1FB2677D3" blockId="1.[151,1437,973,2008]" box="[199,1223,1765,1792]" pageId="1" pageNumber="82">Legs. Tibiae with two distal spurs. Tarsus three segmented. Claws with one preapical tooth.</paragraph>
<paragraph id="8BDA36B0FF88FFF93FD4723EFE007640" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="82">
Abdomen broken, intersegment not visible. Genitalia visible (
<figureCitation id="135E2A35FF88FFF93C99723EFC13760F" box="[906,1010,1801,1827]" captionStart="FIGURES 48" captionStartId="3.[151,264,1681,1705]" captionTargetBox="[198,1386,200,1643]" captionTargetId="figure-17@3.[193,1395,181,1657]" captionTargetPageId="3" captionText="FIGURES 48. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 4. head (dorsal view); 5. maxillary palpus; 6. a conical sensillum on mx2; 7. photograph and drawing of terminalia (ventral view); 8. photograph and drawing of habitus (dorsal view). ep: epiproct; pp: paraproct; ev: external valve; sg: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5729052" httpUri="https://zenodo.org/record/5729052/files/figure.png" pageId="1" pageNumber="82">Figs 78</figureCitation>
). Clunium simple. Epiproct subtriangular with round apex; paraproct with trichobothrial field. Subgenital plate (
<figureCitation id="135E2A35FF88FFF93CE1721AFBD77664" box="[1010,1078,1837,1864]" captionStart="FIGURES 48" captionStartId="3.[151,264,1681,1705]" captionTargetBox="[198,1386,200,1643]" captionTargetId="figure-17@3.[193,1395,181,1657]" captionTargetPageId="3" captionText="FIGURES 48. Psyllipsocus yangi sp. n., holotype CAU-BA-LFY-21001, female. 4. head (dorsal view); 5. maxillary palpus; 6. a conical sensillum on mx2; 7. photograph and drawing of terminalia (ventral view); 8. photograph and drawing of habitus (dorsal view). ep: epiproct; pp: paraproct; ev: external valve; sg: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5729052" httpUri="https://zenodo.org/record/5729052/files/figure.png" pageId="1" pageNumber="82">Fig. 7</figureCitation>
) simple. External valve visible, sclerotized and bearing setae.
</paragraph>
<paragraph id="8BDA36B0FF88FFF93FD47242FC5176FB" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="82">
<emphasis id="B911EAA2FF88FFF93FD47242FD2076A3" bold="true" box="[199,705,1909,1935]" pageId="1" pageNumber="82">
Description of
<typeStatus id="54DE8812FF88FFF93E657242FE0276A3" box="[374,483,1909,1935]" pageId="1" pageNumber="82" type="paratype">paratype</typeStatus>
, gender uncertain.
</emphasis>
CAU-BA-LFY-21002 (
<figureCitation id="135E2A35FF88FFF93CD87242FBDE76BC" box="[971,1087,1909,1936]" captionStart="FIGURES 915" captionStartId="4.[151,264,1612,1636]" captionTargetBox="[156,1426,191,1579]" captionTargetId="figure-24@4.[154,1434,181,1588]" captionTargetPageId="4" captionText="FIGURES 915. Psyllipsocus yangi sp. n., Paratype CAU-BA-LFY-21002. 9. habitus; 10. mouthparts; 11. forewing; 12. hindwing; 13. in-flight wing-coupling structure; 14. labial palps; 15. lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia." figureDoi="http://doi.org/10.5281/zenodo.5729058" httpUri="https://zenodo.org/record/5729058/files/figure.png" pageId="1" pageNumber="82">Figs 915</figureCitation>
). f1:
<quantity id="4C9D9B55FF88FFF93B647242FB3C76BC" box="[1143,1245,1909,1936]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="82" unit="mm" value="0.1">0.10 mm</quantity>
, f2:
<quantity id="4C9D9B55FF88FFF93A1E7242FA9376BC" box="[1293,1394,1909,1936]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="82" unit="mm" value="0.08">0.08 mm</quantity>
, f3:
<quantity id="4C9D9B55FF88FFF93F8472AEFF107698" box="[151,241,1945,1972]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="82" unit="mm" value="0.1">0.1 mm</quantity>
. FWL:
<quantity id="4C9D9B55FF88FFF93E5572AEFE4D7698" box="[326,428,1945,1972]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="1" pageNumber="82" unit="mm" value="1.39">1.39 mm</quantity>
, FWW:
<quantity id="4C9D9B55FF88FFF93D1872AEFD857698" box="[523,612,1945,1972]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="82" unit="mm" value="0.5">0.5 mm</quantity>
; HWL:
<quantity id="4C9D9B55FF88FFF93DAC72AEFCC77698" box="[703,806,1945,1972]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="1" pageNumber="82" unit="mm" value="1.09">1.09 mm</quantity>
, HWW:
<quantity id="4C9D9B55FF88FFF93C9A72AEFC117698" box="[905,1008,1945,1972]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="82" unit="mm" value="0.38">0.38 mm</quantity>
. ft1:
<quantity id="4C9D9B55FF88FFF93B3972AEFB717698" box="[1066,1168,1945,1972]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="82" unit="mm" value="0.15">0.15 mm</quantity>
, ft2:
<quantity id="4C9D9B55FF88FFF93BD972AEFAD07698" box="[1226,1329,1945,1972]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="82" unit="mm" value="0.04">0.04 mm</quantity>
, ft3:
<quantity id="4C9D9B55FF88FFF93A7872AEFF2776F4" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="82" unit="mm" value="0.05">0.05 mm</quantity>
; mt1:
<quantity id="4C9D9B55FF88FFF93E1F728AFE9376F4" box="[268,370,1981,2008]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="82" unit="mm" value="0.15">0.15 mm</quantity>
, mt2:
<quantity id="4C9D9B55FF88FFF93EAB728AFDFF76F4" box="[440,542,1981,2008]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="82" unit="mm" value="0.2">0.20 mm</quantity>
, mt3 broken; hind leg tarsi missing.
</paragraph>
<caption id="DF1A6638FF8BFFFA3F8471E5FEFB741E" ID-DOI="http://doi.org/10.5281/zenodo.5729042" ID-Zenodo-Dep="5729042" httpUri="https://zenodo.org/record/5729042/files/figure.png" pageId="2" pageNumber="83" startId="2.[151,264,1234,1258]" targetBox="[167,1417,190,1200]" targetPageId="2">
<paragraph id="8BDA36B0FF8BFFFA3F8471E5FEFB741E" blockId="2.[151,1437,1234,1331]" pageId="2" pageNumber="83">
<emphasis id="B911EAA2FF8BFFFA3F8471E5FED875C7" bold="true" box="[151,313,1234,1259]" pageId="2" pageNumber="83">FIGURES 13.</emphasis>
<taxonomicName id="4C654D33FF8BFFFA3E2D71E4FE1B75C6" authority="Liang &amp; Liu, 2021" authorityName="Liang &amp; Liu" authorityYear="2021" box="[318,506,1235,1259]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov.">
<emphasis id="B911EAA2FF8BFFFA3E2D71E4FE1B75C6" box="[318,506,1235,1259]" italics="true" pageId="2" pageNumber="83">Psyllipsocus yangi</emphasis>
</taxonomicName>
<emphasis id="B911EAA2FF8BFFFA3D1371E4FDD675C7" bold="true" box="[512,567,1235,1259]" pageId="2" pageNumber="83">
<taxonomicNameLabel id="A22257D9FF8BFFFA3D1371E4FDD675C7" box="[512,567,1235,1259]" pageId="2" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype CAU-BA-LFY-21001, female.
<emphasis id="B911EAA2FF8BFFFA3CC871E4FC0F75C7" bold="true" box="[987,1006,1235,1259]" pageId="2" pageNumber="83">1.</emphasis>
photograph and drawing of habitus (dorsal view), scale bar=0.5 mm;
<emphasis id="B911EAA2FF8BFFFA3EB371C0FE527423" bold="true" box="[416,435,1271,1295]" pageId="2" pageNumber="83">2.</emphasis>
photograph and drawing of habitus (ventral view), scale bar=0.5 mm;
<emphasis id="B911EAA2FF8BFFFA3B9271C0FB757423" bold="true" box="[1153,1172,1271,1295]" pageId="2" pageNumber="83">3.</emphasis>
fore- and hindwing, scale bar=0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDA36B0FF8BFFFA3FD47056FEB77727" blockId="2.[151,1437,1377,2016]" pageId="2" pageNumber="83">
Body and wings similar to the CAU-BA-LFY-21001, but dehydrated and broken. Lacinia (
<figureCitation id="135E2A35FF8BFFFA3BC57056FAB87450" box="[1238,1369,1377,1404]" captionStart="FIGURES 915" captionStartId="4.[151,264,1612,1636]" captionTargetBox="[156,1426,191,1579]" captionTargetId="figure-24@4.[154,1434,181,1588]" captionTargetPageId="4" captionText="FIGURES 915. Psyllipsocus yangi sp. n., Paratype CAU-BA-LFY-21002. 9. habitus; 10. mouthparts; 11. forewing; 12. hindwing; 13. in-flight wing-coupling structure; 14. labial palps; 15. lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia." figureDoi="http://doi.org/10.5281/zenodo.5729058" httpUri="https://zenodo.org/record/5729058/files/figure.png" pageId="2" pageNumber="83">Figs 10, 15</figureCitation>
) with three apical teeth. Labial palps (
<figureCitation id="135E2A35FF8BFFFA3EE970B2FD9874B3" box="[506,633,1413,1439]" captionStart="FIGURES 915" captionStartId="4.[151,264,1612,1636]" captionTargetBox="[156,1426,191,1579]" captionTargetId="figure-24@4.[154,1434,181,1588]" captionTargetPageId="4" captionText="FIGURES 915. Psyllipsocus yangi sp. n., Paratype CAU-BA-LFY-21002. 9. habitus; 10. mouthparts; 11. forewing; 12. hindwing; 13. in-flight wing-coupling structure; 14. labial palps; 15. lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia." figureDoi="http://doi.org/10.5281/zenodo.5729058" httpUri="https://zenodo.org/record/5729058/files/figure.png" pageId="2" pageNumber="83">Figs 10, 14</figureCitation>
) with rounded apex. Wings with “normal” venation, left and right venation symmetric. Forewing (
<figureCitation id="135E2A35FF8BFFFA3EDB709EFDF674EF" box="[456,535,1449,1475]" captionStart="FIGURES 915" captionStartId="4.[151,264,1612,1636]" captionTargetBox="[156,1426,191,1579]" captionTargetId="figure-24@4.[154,1434,181,1588]" captionTargetPageId="4" captionText="FIGURES 915. Psyllipsocus yangi sp. n., Paratype CAU-BA-LFY-21002. 9. habitus; 10. mouthparts; 11. forewing; 12. hindwing; 13. in-flight wing-coupling structure; 14. labial palps; 15. lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia." figureDoi="http://doi.org/10.5281/zenodo.5729058" httpUri="https://zenodo.org/record/5729058/files/figure.png" pageId="2" pageNumber="83">Fig. 11</figureCitation>
) Rs with two branches, M with three branches, in-flight wing-coupling structure simple (
<figureCitation id="135E2A35FF8BFFFA3FE770FAFEA874CB" box="[244,329,1485,1511]" captionStart="FIGURES 915" captionStartId="4.[151,264,1612,1636]" captionTargetBox="[156,1426,191,1579]" captionTargetId="figure-24@4.[154,1434,181,1588]" captionTargetPageId="4" captionText="FIGURES 915. Psyllipsocus yangi sp. n., Paratype CAU-BA-LFY-21002. 9. habitus; 10. mouthparts; 11. forewing; 12. hindwing; 13. in-flight wing-coupling structure; 14. labial palps; 15. lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia." figureDoi="http://doi.org/10.5281/zenodo.5729058" httpUri="https://zenodo.org/record/5729058/files/figure.png" pageId="2" pageNumber="83">Fig. 13</figureCitation>
). Hindwing (
<figureCitation id="135E2A35FF8BFFFA3EF170FAFDD474CB" box="[482,565,1485,1511]" captionStart="FIGURES 915" captionStartId="4.[151,264,1612,1636]" captionTargetBox="[156,1426,191,1579]" captionTargetId="figure-24@4.[154,1434,181,1588]" captionTargetPageId="4" captionText="FIGURES 915. Psyllipsocus yangi sp. n., Paratype CAU-BA-LFY-21002. 9. habitus; 10. mouthparts; 11. forewing; 12. hindwing; 13. in-flight wing-coupling structure; 14. labial palps; 15. lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia." figureDoi="http://doi.org/10.5281/zenodo.5729058" httpUri="https://zenodo.org/record/5729058/files/figure.png" pageId="2" pageNumber="83">Fig. 12</figureCitation>
) with M fork not much longer than Rs fork. Abdomen terminal with a sacshaped structure.
</paragraph>
</subSubSection>
<subSubSection id="C37F653BFF8BFFFA3FD47322FF0A775B" pageId="2" pageNumber="83" type="materials_examined">
<paragraph id="8BDA36B0FF8BFFFA3FD47322FF0A775B" blockId="2.[151,1437,1377,2016]" pageId="2" pageNumber="83">
<emphasis id="B911EAA2FF8BFFFA3FD47322FE617703" bold="true" box="[199,384,1557,1583]" pageId="2" pageNumber="83">Type materials.</emphasis>
The type specimens will eventually be deposited in the Entomological Museum,
<collectingCountry id="F3727620FF8BFFFA3A047322FABB7703" box="[1303,1370,1557,1583]" name="China" pageId="2" pageNumber="83">China</collectingCountry>
Agricultural University (
<collectionCode id="ED74AE75FF8BFFFA3E6F730EFE5C777F" box="[380,445,1593,1619]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="2" pageNumber="83">CAU</collectionCode>
).
<materialsCitation id="3B0D3CEDFF8BFFFA3EC2730EFC7E777F" ID-GBIF-Occurrence="3412204302" box="[465,927,1593,1619]" pageId="2" pageNumber="83" specimenCode="CAU-BA-LFY-21001" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54DE8812FF8BFFFA3EC2730EFDDF777F" box="[465,574,1593,1619]" pageId="2" pageNumber="83" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBC39ECBFF8BFFFA3D59730EFCA3777F" box="[586,834,1593,1619]" pageId="2" pageNumber="83">CAU-BA-LFY-21001</specimenCode>
,
<specimenCount id="9D63FD39FF8BFFFA3C5C730EFC7E777F" box="[847,927,1593,1619]" pageId="2" pageNumber="83" type="female">female</specimenCount>
</materialsCitation>
.
<materialsCitation id="3B0D3CEDFF8BFFFA3CB8730EFF06775B" ID-GBIF-Occurrence="3412204301" pageId="2" pageNumber="83" typeStatus="paratype">
<typeStatus id="54DE8812FF8BFFFA3CB8730EFBF0777F" box="[939,1041,1593,1619]" pageId="2" pageNumber="83" type="paratype">Paratype</typeStatus>
: CAU-BA-LFY-21002, gender uncertain
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F653BFF8BFFFA3FD473B6FF277793" pageId="2" pageNumber="83" type="etymology">
<paragraph id="8BDA36B0FF8BFFFA3FD473B6FF277793" blockId="2.[151,1437,1377,2016]" pageId="2" pageNumber="83">
<emphasis id="B911EAA2FF8BFFFA3FD473B6FEAF77B7" bold="true" box="[199,334,1665,1691]" pageId="2" pageNumber="83">Etymology.</emphasis>
The new species is dedicated to Prof. Ding Yang for his support to the study of Chinese
<taxonomicName id="4C654D33FF8BFFFA3A4373B6FF207793" authorityName="Hennig" authorityYear="1966" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="83" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F653BFF8BFFFD3FD473FEFD9D706D" lastPageId="5" lastPageNumber="86" pageId="2" pageNumber="83" type="discussion">
<paragraph id="8BDA36B0FF8BFFFA3FD473FEFD867663" blockId="2.[151,1437,1377,2016]" pageId="2" pageNumber="83">
<emphasis id="B911EAA2FF8BFFFA3FD473FEFEDA77CF" bold="true" box="[199,315,1737,1763]" pageId="2" pageNumber="83">Remarks.</emphasis>
<bibRefCitation id="EFF44B41FF8BFFFA3E5173FEFDF477CF" author="Wang, R. Q. &amp; Li S. &amp; Ren, D. &amp; Yao, Y. Z." box="[322,533,1737,1763]" pageId="2" pageNumber="83" pagination="104178" refId="ref4145" refString="Wang, R. Q., Li S., Ren, D. &amp; Yao, Y. Z. (2019) New genus and species of the Psyllipsocidae (Psocodea: Trogiomorpha) from mid-Cretaceous Burmese amber. Cretaceous Research, 104, 104178." type="journal article" year="2019">
Wang
<emphasis id="B911EAA2FF8BFFFA3E9973FDFE5D77CF" box="[394,444,1737,1763]" italics="true" pageId="2" pageNumber="83">et al</emphasis>
. (2019)
</bibRefCitation>
and Hakim
<emphasis id="B911EAA2FF8BFFFA3DB373FDFD3077CF" box="[672,721,1737,1763]" italics="true" pageId="2" pageNumber="83">et al</emphasis>
. (2020) indicated the venation of
<taxonomicName id="4C654D33FF8BFFFA3B4373FEFB5B77CF" authorityName="Hennig" authorityYear="1966" box="[1104,1210,1737,1763]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="83" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
is variable. The
<typeStatus id="54DE8812FF8BFFFA3A6473FEFF3C762B" pageId="2" pageNumber="83" type="holotype">holotype</typeStatus>
of the new species shows asymmetric venation: left forewing with Rs two-branched; M three-branched, but M3 branched; right forewing with Rs bifurcating into R
<subScript id="17E134F5FF8BFFFA3C027214FCCB761D" attach="left" box="[785,810,1827,1841]" fontSize="6" pageId="2" pageNumber="83">2+3</subScript>
and R
<subScript id="17E134F5FF8BFFFA3C657214FC71761D" attach="left" box="[886,912,1827,1841]" fontSize="6" pageId="2" pageNumber="83">4+5</subScript>
, but R
<subScript id="17E134F5FF8BFFFA3CCE7214FC16761D" attach="left" box="[989,1015,1827,1841]" fontSize="6" pageId="2" pageNumber="83">4+5</subScript>
branched; M three-branched, but the middle branch not reaching wing margin.
</paragraph>
<paragraph id="8BDA36B0FF8BFFFA3FD4726EFCAC76F3" blockId="2.[151,1437,1377,2016]" pageId="2" pageNumber="83">
This new species can be included in
<taxonomicName id="4C654D33FF8BFFFA3D4D726EFD0B765F" authorityName="Selys-Longchamps" authorityYear="1872" box="[606,746,1881,1907]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8BFFFA3D4D726EFD0B765F" box="[606,746,1881,1907]" italics="true" pageId="2" pageNumber="83">Psyllipsocus</emphasis>
</taxonomicName>
by the following characters: 1) lacinia with three apical teeth; 2) forewing Sc short and ending free in membrane; 3) forewing veins with one row of setae; 4) forewing pterostigma with four sides; 5) Cu2 and A meeting at forewing margin, in-flight wing-coupling structure composed of separate and almost straight spines; 6) broad and pilous external valve.
</paragraph>
<caption id="DF1A6638FF8AFFFB3F8473A6FC3177DD" ID-DOI="http://doi.org/10.5281/zenodo.5729052" ID-Zenodo-Dep="5729052" httpUri="https://zenodo.org/record/5729052/files/figure.png" pageId="3" pageNumber="84" startId="3.[151,264,1681,1705]" targetBox="[198,1386,200,1643]" targetPageId="3">
<paragraph id="8BDA36B0FF8AFFFB3F8473A6FC3177DD" blockId="3.[151,1436,1681,1778]" pageId="3" pageNumber="84">
<emphasis id="B911EAA2FF8AFFFB3F8473A6FEDA7785" bold="true" box="[151,315,1681,1705]" pageId="3" pageNumber="84">FIGURES 48.</emphasis>
<taxonomicName id="4C654D33FF8AFFFB3E5173A6FDE17785" authority="Liang &amp; Liu, 2021" authorityName="Liang &amp; Liu" authorityYear="2021" box="[322,512,1681,1705]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov.">
<emphasis id="B911EAA2FF8AFFFB3E5173A6FDE17785" box="[322,512,1681,1705]" italics="true" pageId="3" pageNumber="84">Psyllipsocus yangi</emphasis>
</taxonomicName>
<emphasis id="B911EAA2FF8AFFFB3D1473A6FDA07785" bold="true" box="[519,577,1681,1705]" pageId="3" pageNumber="84">
<taxonomicNameLabel id="A22257D9FF8AFFFB3D1473A6FDA07785" box="[519,577,1681,1705]" pageId="3" pageNumber="84" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype CAU-BA-LFY-21001, female.
<emphasis id="B911EAA2FF8AFFFB3CF873A6FC1F7785" bold="true" box="[1003,1022,1681,1705]" pageId="3" pageNumber="84">4.</emphasis>
head (dorsal view);
<emphasis id="B911EAA2FF8AFFFB3BC273A6FB057785" bold="true" box="[1233,1252,1681,1705]" pageId="3" pageNumber="84">5.</emphasis>
maxillary palpus;
<emphasis id="B911EAA2FF8AFFFB3F847382FF4B77E1" bold="true" box="[151,170,1717,1741]" pageId="3" pageNumber="84">6.</emphasis>
a conical sensillum on mx2;
<emphasis id="B911EAA2FF8AFFFB3ECA7382FE0A77E1" bold="true" box="[473,491,1717,1741]" pageId="3" pageNumber="84">7.</emphasis>
photograph and drawing of terminalia (ventral view);
<emphasis id="B911EAA2FF8AFFFB3B0F7382FBCE77E1" bold="true" box="[1052,1071,1717,1741]" pageId="3" pageNumber="84">8.</emphasis>
photograph and drawing of habitus (dorsal view). ep: epiproct; pp: paraproct; ev: external valve; sg: subgenital plate.
</paragraph>
</caption>
<paragraph id="8BDA36B0FF8AFFFB3FD47228FB4576C1" blockId="3.[151,1437,1823,2030]" pageId="3" pageNumber="84">
<taxonomicName id="4C654D33FF8AFFFB3FD47228FE7D7615" authorityName="Liang &amp; Liu" authorityYear="2021" box="[199,412,1823,1849]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="B911EAA2FF8AFFFB3FD47228FE7D7615" box="[199,412,1823,1849]" italics="true" pageId="3" pageNumber="84">Psyllipsocus yangi</emphasis>
</taxonomicName>
differs from the genera
<taxonomicName id="4C654D33FF8AFFFB3DAC7217FBAB7615" authority="Wang et al., 2019" authorityName="Wang" authorityYear="2019" box="[703,1098,1823,1850]" class="Insecta" family="Psyllipsocidae" genus="Concavapsocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8AFFFB3DAC7217FC8E7616" box="[703,879,1824,1850]" italics="true" pageId="3" pageNumber="84">Concavapsocus</emphasis>
<bibRefCitation id="EFF44B41FF8AFFFB3C6A7217FBAB7615" author="Wang, R. Q. &amp; Li S. &amp; Ren, D. &amp; Yao, Y. Z." box="[889,1098,1823,1850]" pageId="3" pageNumber="84" pagination="104178" refId="ref4145" refString="Wang, R. Q., Li S., Ren, D. &amp; Yao, Y. Z. (2019) New genus and species of the Psyllipsocidae (Psocodea: Trogiomorpha) from mid-Cretaceous Burmese amber. Cretaceous Research, 104, 104178." type="journal article" year="2019">
Wang
<emphasis id="B911EAA2FF8AFFFB3CD67217FC1B7615" box="[965,1018,1823,1849]" italics="true" pageId="3" pageNumber="84">et al</emphasis>
., 2019
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C654D33FF8AFFFB3B9A7228FF2E7672" authority="Vishniakova, 1975" authorityName="Vishniakova" authorityYear="1975" class="Insecta" family="Psyllipsocidae" genus="Khatangia" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8AFFFB3B9A7228FB1E7615" box="[1161,1279,1823,1849]" italics="true" pageId="3" pageNumber="84">Khatangia</emphasis>
<bibRefCitation id="EFF44B41FF8AFFFB3A1A7228FF2E7672" author="Vishniakova, V. N." pageId="3" pageNumber="84" pagination="63 - 75" refId="ref4022" refString="Vishniakova, V. N. (1975) Psocoptera in Late-Cretaceous insect-bearing resins from the Taimyr. Entomological Review, 54, 63 - 75." type="journal article" year="1975">Vishniakova, 1975</bibRefCitation>
</taxonomicName>
by the presence of nodulus; differs from the genus
<taxonomicName id="4C654D33FF8AFFFB3C087274FB257671" authority="Azar &amp; Nel, 2011" authorityName="Azar &amp; Nel" authorityYear="2011" box="[795,1220,1859,1885]" class="Insecta" family="Pachytroctidae" genus="Libanopsyllipsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8AFFFB3C087274FC127671" box="[795,1011,1859,1885]" italics="true" pageId="3" pageNumber="84">Libanopsyllipsocus</emphasis>
<bibRefCitation id="EFF44B41FF8AFFFB3CEA7274FB257671" author="Azar, D. &amp; Nel, A." box="[1017,1220,1859,1885]" pageId="3" pageNumber="84" pagination="153 - 165" refId="ref2927" refString="Azar, D. &amp; Nel, A. (2011) The oldest psyllipsocid booklice, in Lower Cretaceous amber from Lebanon (Psocodea, Trogiomorpha, Psocathropetae, Psyllipsocidae). ZooKeys, 130, 153 - 165. https: // doi. org / 10.3897 / zookeys. 130.1430" type="journal article" year="2011">Azar &amp; Nel, 2011</bibRefCitation>
</taxonomicName>
by the presence of pterostigma; and differs from the genus
<taxonomicName id="4C654D33FF8AFFFB3D4E7250FBED76AD" authority="Hakim et al., 2018" authorityName="Hakim" authorityYear="2018" box="[605,1036,1895,1922]" class="Insecta" family="Psyllipsocidae" genus="Annulipsyllipsocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8AFFFB3D4E7250FCD076AD" box="[605,817,1895,1921]" italics="true" pageId="3" pageNumber="84">Annulipsyllipsocus</emphasis>
<bibRefCitation id="EFF44B41FF8AFFFB3C2B7250FBED76AD" author="Hakim, M. &amp; Azar, S. &amp; Maksoud, S. &amp; Huang, D. Y. &amp; Azar, D." box="[824,1036,1895,1922]" pageId="3" pageNumber="84" pagination="389 - 400" refId="ref2980" refString="Hakim, M., Azar, S., Maksoud, S., Huang, D. Y. &amp; Azar, D. (2018) New polymorphic psyllipsocids from Burmese amber (Psocodea: Psyllipsocidae). Cretaceous Research, 84, 389 - 400. https: // doi. org / 10.1016 / j. cretres. 2017.11.027" type="journal article" year="2018">
Hakim
<emphasis id="B911EAA2FF8AFFFB3C9F725FFC5F76AD" box="[908,958,1895,1921]" italics="true" pageId="3" pageNumber="84">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
by the pterostigma with four sides. This new species is similar to
<taxonomicName id="4C654D33FF8AFFFB3EFC72BCFCD47689" box="[495,821,1931,1957]" class="Insecta" family="Ptiloneuridae" genus="Sinopsyllipsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="fushunensis">
<emphasis id="B911EAA2FF8AFFFB3EFC72BCFCD47689" box="[495,821,1931,1957]" italics="true" pageId="3" pageNumber="84">Sinopsyllipsocus fushunensis</emphasis>
</taxonomicName>
in the subtriangular pterostigma, the annulated flagellomeres and the presence of a conical sensillum in mx2, but it differs from this taxon by the shorter forewing, the completely transparent wings, the quadrangular basiradial cell and bifurcated A in hindwing.
</paragraph>
<caption id="DF1A6638FF8DFFFC3F84737BFE157780" ID-DOI="http://doi.org/10.5281/zenodo.5729058" ID-Zenodo-Dep="5729058" httpUri="https://zenodo.org/record/5729058/files/figure.png" pageId="4" pageNumber="85" startId="4.[151,264,1612,1636]" targetBox="[156,1426,191,1579]" targetPageId="4">
<paragraph id="8BDA36B0FF8DFFFC3F84737BFE157780" blockId="4.[151,1436,1612,1708]" pageId="4" pageNumber="85">
<emphasis id="B911EAA2FF8DFFFC3F84737BFEA67748" bold="true" box="[151,327,1612,1636]" pageId="4" pageNumber="85">FIGURES 915.</emphasis>
<taxonomicName id="4C654D33FF8DFFFC3E5E737BFDEB7748" authority="Liang &amp; Liu, 2021" authorityName="Liang &amp; Liu" authorityYear="2021" box="[333,522,1612,1636]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov.">
<emphasis id="B911EAA2FF8DFFFC3E5E737BFDEB7748" box="[333,522,1612,1636]" italics="true" pageId="4" pageNumber="85">Psyllipsocus yangi</emphasis>
</taxonomicName>
<emphasis id="B911EAA2FF8DFFFC3D03737AFDA87749" bold="true" box="[528,585,1613,1637]" pageId="4" pageNumber="85">
<taxonomicNameLabel id="A22257D9FF8DFFFC3D03737AFDA87749" box="[528,585,1613,1637]" pageId="4" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, Paratype CAU-BA-LFY-21002.
<emphasis id="B911EAA2FF8DFFFC3C8C737BFC537748" bold="true" box="[927,946,1612,1636]" pageId="4" pageNumber="85">9.</emphasis>
habitus;
<emphasis id="B911EAA2FF8DFFFC3B1C737BFBCE7748" bold="true" box="[1039,1071,1612,1636]" pageId="4" pageNumber="85">10.</emphasis>
mouthparts;
<emphasis id="B911EAA2FF8DFFFC3BA7737BFB337748" bold="true" box="[1204,1234,1612,1636]" pageId="4" pageNumber="85">11.</emphasis>
forewing;
<emphasis id="B911EAA2FF8DFFFC3A52737BFA807748" bold="true" box="[1345,1377,1612,1636]" pageId="4" pageNumber="85">12.</emphasis>
hindwing;
<emphasis id="B911EAA2FF8DFFFC3FCB7347FF1977A4" bold="true" box="[216,248,1648,1672]" pageId="4" pageNumber="85">13.</emphasis>
in-flight wing-coupling structure;
<emphasis id="B911EAA2FF8DFFFC3D497347FD9B77A4" bold="true" box="[602,634,1648,1672]" pageId="4" pageNumber="85">14.</emphasis>
labial palps;
<emphasis id="B911EAA2FF8DFFFC3C177347FCC577A4" bold="true" box="[772,804,1648,1672]" pageId="4" pageNumber="85">15.</emphasis>
lacinia. st: pterostigma; ap: areola postica cell; mx: maxillary palpus; lp: labial palps; la: lacinia.
</paragraph>
</caption>
<paragraph id="8BDA36B0FF8DFFFC3FD473ECFDB8764D" blockId="4.[151,1437,1755,2034]" pageId="4" pageNumber="85">
<bibRefCitation id="EFF44B41FF8DFFFC3FD473ECFE4377DA" author="Zhang, Q. &amp; Nel, A. &amp; Azar, D. &amp; Wang, B." box="[199,418,1755,1782]" pageId="4" pageNumber="85" pagination="366 - 372" refId="ref4297" refString="Zhang, Q., Nel, A., Azar, D. &amp; Wang, B. (2016) New Chinese psocids from Eocene Fushun amber (Insecta: Psocodea). Alcheringa: An Australasian Journal of Palaeontology, 40 (3), 366 - 372. https: // doi. org / 10.1080 / 03115518.2016.1144952" type="journal article" year="2016">
Zhang
<emphasis id="B911EAA2FF8DFFFC3E0573EBFEA977D9" box="[278,328,1755,1781]" italics="true" pageId="4" pageNumber="85">et al</emphasis>
. (2016)
</bibRefCitation>
established the genus
<taxonomicName id="4C654D33FF8DFFFC3DB373ECFCBA77D9" box="[672,859,1755,1781]" class="Insecta" family="Ptiloneuridae" genus="Sinopsyllipsocus" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8DFFFC3DB373ECFCBA77D9" box="[672,859,1755,1781]" italics="true" pageId="4" pageNumber="85">Sinopsyllipsocus</emphasis>
</taxonomicName>
based on
<taxonomicName id="4C654D33FF8DFFFC3CDD73ECFB8F77D9" box="[974,1134,1755,1781]" class="Insecta" family="Ptiloneuridae" genus="Sinopsyllipsocus" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="fushunensis">
<emphasis id="B911EAA2FF8DFFFC3CDD73ECFB8F77D9" box="[974,1134,1755,1781]" italics="true" pageId="4" pageNumber="85">S. fushunensis</emphasis>
</taxonomicName>
from Fushun amber (early Eocene), and placed this genus into the family
<taxonomicName id="4C654D33FF8DFFFC3DBA73C8FCAA7635" authorityName="Kolbe" authorityYear="1884" box="[681,843,1791,1817]" class="Insecta" family="Psyllipsocidae" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="85" phylum="Arthropoda" rank="family">Psyllipsocidae</taxonomicName>
because of the forewing venation, and distinguished
<taxonomicName id="4C654D33FF8DFFFC3F847214FEB37611" box="[151,338,1827,1853]" class="Insecta" family="Ptiloneuridae" genus="Sinopsyllipsocus" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8DFFFC3F847214FEB37611" box="[151,338,1827,1853]" italics="true" pageId="4" pageNumber="85">Sinopsyllipsocus</emphasis>
</taxonomicName>
from the other genera of
<taxonomicName id="4C654D33FF8DFFFC3D677214FCF77611" authorityName="Kolbe" authorityYear="1884" box="[628,790,1827,1853]" class="Insecta" family="Psyllipsocidae" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="85" phylum="Arthropoda" rank="family">Psyllipsocidae</taxonomicName>
by the presence of secondary annulation on the antennae and the presence of a sensillum on mx2.
</paragraph>
<paragraph id="8BDA36B0FF8DFFFC3FD4725CFEC976E1" blockId="4.[151,1437,1755,2034]" pageId="4" pageNumber="85">
Secondary annulation in the antennal flagellomeres is considered an apomorphy for the suborder
<taxonomicName id="4C654D33FF8DFFFC3BEE725CFEB37685" authority="(Lienhard 1998)" baseAuthorityName="Lienhard" baseAuthorityYear="1998" pageId="4" pageNumber="85" rank="subOrder" subOrder="Troctomorpha">Troctomorpha (Lienhard 1998)</taxonomicName>
, although
<bibRefCitation id="EFF44B41FF8DFFFC3ED872B8FD4F7685" author="Hakim, M. &amp; Azar, S. &amp; Maksoud, S. &amp; Huang, D. Y. &amp; Azar, D." box="[459,686,1935,1961]" pageId="4" pageNumber="85" pagination="389 - 400" refId="ref2980" refString="Hakim, M., Azar, S., Maksoud, S., Huang, D. Y. &amp; Azar, D. (2018) New polymorphic psyllipsocids from Burmese amber (Psocodea: Psyllipsocidae). Cretaceous Research, 84, 389 - 400. https: // doi. org / 10.1016 / j. cretres. 2017.11.027" type="journal article" year="2018">
Hakim
<emphasis id="B911EAA2FF8DFFFC3D3372A7FDB57685" box="[544,596,1935,1961]" italics="true" pageId="4" pageNumber="85">et al</emphasis>
. (2018)
</bibRefCitation>
and
<bibRefCitation id="EFF44B41FF8DFFFC3DFB72B8FC077685" author="Perrichot, V. &amp; Azar, D. &amp; Neraudeau, D. &amp; Nel, A." box="[744,998,1935,1961]" pageId="4" pageNumber="85" pagination="669 - 683" refId="ref3690" refString="Perrichot, V., Azar, D., Neraudeau, D. &amp; Nel, A. (2003) New Psocoptera in the Lower Cretaceous ambers of southwestern France and Lebanon (Insecta: Psocoptera: Trogiomorpha). Geological Magazine, 140 (6), 669 - 683. https: // doi. org / 10.1017 / S 0016756803008355" type="journal article" year="2003">
Perrichot
<emphasis id="B911EAA2FF8DFFFC3C4B72A7FC6A7685" box="[856,907,1935,1961]" italics="true" pageId="4" pageNumber="85">et al</emphasis>
. (2003)
</bibRefCitation>
suggested that this character could be homoplastic.
</paragraph>
<paragraph id="8BDA36B0FF8DFFFD3FD472E0FAAE71D6" blockId="4.[151,1437,1755,2034]" lastBlockId="5.[151,1437,151,321]" lastPageId="5" lastPageNumber="86" pageId="4" pageNumber="85">
<bibRefCitation id="EFF44B41FF8DFFFC3FD472E0FE2F76DE" author="Yoshizawa, K. &amp; Lienhard, C. &amp; Johnson, K. P." box="[199,462,2007,2034]" pageId="4" pageNumber="85" pagination="287 - 299" refId="ref4234" refString="Yoshizawa, K., Lienhard, C. &amp; Johnson, K. P. (2006) Molecular systematics of the suborder Trogiomorpha (Insecta: Psocodea: ' Psocoptera'). Zoological Journal of the Linnaean Society, 146, 287 - 299. https: // doi. org / 10.1111 / j. 1096 - 3642.2006.00207. x" type="journal article" year="2006">
Yoshizawa
<emphasis id="B911EAA2FF8DFFFC3E5472EFFE9776DD" box="[327,374,2007,2033]" italics="true" pageId="4" pageNumber="85">et al</emphasis>
. (2006)
</bibRefCitation>
reconstructed the phylogenetic relationships among extant families of the suborder
<taxonomicName id="4C654D33FF8DFFFD3A7872E0FEF1719D" authorityName="Roesler" authorityYear="1940" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="5" lastPageNumber="86" order="Diptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="subOrder" subOrder="Trogiomorpha">Trogiomorpha</taxonomicName>
based on molecular data, and considered “mx2 without conical sensillum” is a key diagnostic character of
<taxonomicName id="4C654D33FF8CFFFD3FA6758CFEBA71F9" authorityName="Kolbe" authorityYear="1884" box="[181,347,187,213]" class="Insecta" family="Psyllipsocidae" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="86" phylum="Arthropoda" rank="family">Psyllipsocidae</taxonomicName>
. However,
<bibRefCitation id="EFF44B41FF8CFFFD3EC4758CFCF471F9" author="Lienhard, C. &amp; Ferreira L. R." box="[471,789,187,213]" pageId="5" pageNumber="86" pagination="421 - 443" refId="ref3180" refString="Lienhard, C. &amp; Ferreira L. R. (2013) Three new species of Psyllipsocus (Psocodea: ' Psocoptera': Psyllipsocidae) from Brazilian caves with description of a novel structure interpreted as a male accessory genital organ. Revue Suisse de Zoologie, 120 (3), 421 - 443." type="journal article" year="2013">Lienhard and Ferreira (2013</bibRefCitation>
,
<bibRefCitation id="EFF44B41FF8CFFFD3C33758CFCBB71F9" author="Lienhard, C. &amp; Ferreira L. R." box="[800,858,187,213]" pageId="5" pageNumber="86" pagination="211 - 246" refId="ref3237" refString="Lienhard, C. &amp; Ferreira L. R. (2014) New species of Psyllipsocus from Brazilian caves (Psocodea: ' Psocoptera': Psyllipsocidae). Revue Suisse de Zoologie, 121 (2), 211 - 246." type="journal article" year="2014">2014</bibRefCitation>
) described nine species of
<taxonomicName id="4C654D33FF8CFFFD3B96758CFAF071F9" authorityName="Selys-Longchamps" authorityYear="1872" box="[1157,1297,187,213]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8CFFFD3B96758CFAF071F9" box="[1157,1297,187,213]" italics="true" pageId="5" pageNumber="86">Psyllipsocus</emphasis>
</taxonomicName>
with a conical sensillum on mx
<quantity id="4C9D9B55FF8CFFFD3E6A75E8FE4271D5" box="[377,419,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="86" unit="in" value="2.0">2 in</quantity>
Brazilian caves, contradicting the diagnosis of
<taxonomicName id="4C654D33FF8CFFFD3CA475E8FBB871D5" authorityName="Kolbe" authorityYear="1884" box="[951,1113,223,249]" class="Insecta" family="Psyllipsocidae" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="86" phylum="Arthropoda" rank="family">Psyllipsocidae</taxonomicName>
of
<bibRefCitation id="EFF44B41FF8CFFFD3B6E75E8FAAA71D6" author="Yoshizawa, K. &amp; Lienhard, C. &amp; Johnson, K. P." box="[1149,1355,223,250]" pageId="5" pageNumber="86" pagination="287 - 299" refId="ref4234" refString="Yoshizawa, K., Lienhard, C. &amp; Johnson, K. P. (2006) Molecular systematics of the suborder Trogiomorpha (Insecta: Psocodea: ' Psocoptera'). Zoological Journal of the Linnaean Society, 146, 287 - 299. https: // doi. org / 10.1111 / j. 1096 - 3642.2006.00207. x" type="journal article" year="2006">Yoshizawa (2006)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDA36B0FF8CFFFD3FD47434FD9D706D" blockId="5.[151,1437,151,321]" pageId="5" pageNumber="86">
In view of the above,
<taxonomicName id="4C654D33FF8CFFFD3ED27434FD9D7031" box="[449,636,259,285]" class="Insecta" family="Ptiloneuridae" genus="Sinopsyllipsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8CFFFD3ED27434FD9D7031" box="[449,636,259,285]" italics="true" pageId="5" pageNumber="86">Sinopsyllipsocus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C654D33FF8CFFFD3C5E7434FC387031" authorityName="Selys-Longchamps" authorityYear="1872" box="[845,985,259,285]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8CFFFD3C5E7434FC387031" box="[845,985,259,285]" italics="true" pageId="5" pageNumber="86">Psyllipsocus</emphasis>
</taxonomicName>
, and it is possible that
<taxonomicName id="4C654D33FF8CFFFD3BF17434FA7C7031" box="[1250,1437,259,285]" class="Insecta" family="Ptiloneuridae" genus="Sinopsyllipsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8CFFFD3BF17434FA7C7031" box="[1250,1437,259,285]" italics="true" pageId="5" pageNumber="86">Sinopsyllipsocus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C654D33FF8CFFFD3FD47410FEB2706D" authorityName="Selys-Longchamps" authorityYear="1872" box="[199,339,295,321]" class="Insecta" family="Psyllipsocidae" genus="Psyllipsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAA2FF8CFFFD3FD47410FEB2706D" box="[199,339,295,321]" italics="true" pageId="5" pageNumber="86">Psyllipsocus</emphasis>
</taxonomicName>
constitue a single lineage.
</paragraph>
</subSubSection>
</treatment>
</document>