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<document id="4ECC0EE8FFFC3870DD7327D4216EF2DA" ID-DOI="10.11646/zootaxa.5569.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734246" ID-ZooBank="6563DBB4-40E9-4FDE-9A8F-6A596369A214" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1737734950194" checkinUser="plazi" docAuthor="Parrinha, Diogo, Marques, Mariana P., Tiutenko, Arthur, Heinicke, Matthew P., Bauer, Aaron M. &amp; Ceríaco, Luis M. P." docDate="2025" docId="0382D211D138FFADB0E221103FC9F9DD" docLanguage="en" docName="zootaxa.5569.3.2.pdf" docOrigin="Zootaxa 5569 (3)" docSource="https://doi.org/10.11646/zootaxa.5569.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhoptropus nivimontanus Parrinha, Marques, Tiutenko, Heinicke, Bauer &amp; Ceríaco, 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="453" masterDocId="FFBBAA69D131FFA3B07522543E49FF87" masterDocTitle="A new species of Namib Day Gecko (Gekkonidae: Rhoptropus Peters, 1869) from the Serra da Neve inselberg, southwestern Angola" masterLastPageNumber="458" masterPageNumber="439" pageNumber="448" updateTime="1738069279369" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="7F0AC6E53B8881755AE7E9EEC50C47D1">A new species of Namib Day Gecko (Gekkonidae: Rhoptropus Peters, 1869) from the Serra da Neve inselberg, southwestern Angola</mods:title>
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<mods:namePart id="F153C6C371784480AAF0EBD0678BFA38">Marques, Mariana P.</mods:namePart>
<mods:affiliation id="C5722BA1B06BFF1F07B89768D540D35D">BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal &amp; CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal &amp; Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal &amp; Section of Amphibians and Reptiles, Carnegie Museum of Natural History, 4400 Forbes Avenue Pittsburgh PA 15213 - 4080, USA</mods:affiliation>
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<mods:namePart id="23A647D5D76B7D5F34E58616595E60D9">Heinicke, Matthew P.</mods:namePart>
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<taxonomicName id="4C2B1884D138FFAAB0E221103F9BFCD8" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[151,466,836,863]" class="Reptilia" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="448" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D138FFAAB0E221103F9BFCD8" bold="true" box="[151,466,836,863]" italics="true" pageId="9" pageNumber="448">Rhoptropus nivimontanus</emphasis>
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="13107F82D138FFAAB0EA21D93F40FC20" box="[159,265,909,935]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStartId-0="10.[152,255,1666,1690]" captionStartId-1="12.[152,255,1112,1136]" captionStartId-2="13.[152,255,1596,1620]" captionStartId-3="14.[152,255,1061,1085]" captionStartId-4="15.[152,255,1123,1147]" captionTargetBox-0="[151,1436,198,1643]" captionTargetBox-1="[189,1398,181,1088]" captionTargetBox-2="[164,1428,194,1559]" captionTargetBox-3="[151,1435,181,1037]" captionTargetBox-4="[151,1436,181,1099]" captionTargetId-0="figure-25@10.[151,1436,181,1645]" captionTargetId-1="figure-25@12.[189,1398,181,1088]" captionTargetId-2="figure-20@13.[151,1436,181,1572]" captionTargetId-3="figure-25@14.[151,1435,181,1037]" captionTargetId-4="figure-20@15.[151,1436,181,1099]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionText-0="FIGURE 4. Dorsal (A) and ventral (B) views of the holotype of Rhoptropus nivimontanus sp. nov. in life (MUHNAC/MB03- 1706). Photos by AT." captionText-1="FIGURE 5. Preserved holotype of Rhoptropus nivimontanus sp. nov. (MUHNAC/MB03-1706). Photos by DP." captionText-2="FIGURE 6. Distribution of Rhoptropus nivimontanus sp. nov. and the boultoni “group” inAngola. Stars represent the respective type localities." captionText-3="FIGURE 7. Habitat of Rhoptropus nivimontanus sp. nov. at the type locality near Catchi village, on the highlands of Serra da Neve, Namibe Province. Photo by LMPC." captionText-4="FIGURE 8. General view of Catchi village and plateau, with riparian gallery and type locality of Rhoptropus nivimontanus sp. nov. in foreground (red arrow). Photo by LMPC." figureDoi-0="http://doi.org/10.5281/zenodo.14734256" figureDoi-1="http://doi.org/10.5281/zenodo.14734258" figureDoi-2="http://doi.org/10.5281/zenodo.14734260" figureDoi-3="http://doi.org/10.5281/zenodo.14734262" figureDoi-4="http://doi.org/10.5281/zenodo.14734266" httpUri-0="https://zenodo.org/record/14734256/files/figure.png" httpUri-1="https://zenodo.org/record/14734258/files/figure.png" httpUri-2="https://zenodo.org/record/14734260/files/figure.png" httpUri-3="https://zenodo.org/record/14734262/files/figure.png" httpUri-4="https://zenodo.org/record/14734266/files/figure.png" pageId="9" pageNumber="448">Figs. 4–8</figureCitation>
)
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aff.
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:
<treatmentCitation id="0A8A4516D138FFAAB1DE219F3C8CFC64" ID-DOI="https://doi.org/10.5281/zenodo.11196676" ID-Zenodo-Dep="11196676" author="Marques, M. P. &amp; Parrinha, D. &amp; Lopes-Lima, M. &amp; Tiutenko, A. &amp; Bauer, A. M. &amp; Ceriaco, L. M. P." box="[427,709,971,995]" httpUri="http://treatment.plazi.org/id/B9641044C3EC5D0DA4D0AEE5249296FE" page="189" pageId="9" pageNumber="448" year="2024">
<bibRefCitation id="EFBA1EF6D138FFAAB1DE219F3C8CFC64" author="Marques, M. P. &amp; Parrinha, D. &amp; Lopes-Lima, M. &amp; Tiutenko, A. &amp; Bauer, A. M. &amp; Ceriaco, L. M. P." box="[427,709,971,995]" pageId="9" pageNumber="448" pagination="167 - 217" refId="ref11774" refString="Marques, M. P., Parrinha, D., Lopes-Lima, M., Tiutenko, A., Bauer, A. M. &amp; Ceriaco, L. M. P. (2024 a) An Island in a sea of sand: a first checklist of the herpetofauna of the Serra da Neve inselberg, southwestern Angola. ZooKeys, 1201, 167 - 217. https: // doi. org / 10.3897 / zookeys. 1201.120750" type="journal article" year="2024">
Marques
<emphasis id="B95FBF15D138FFAAB27F219F3C74FC64" box="[522,573,971,995]" italics="true" pageId="9" pageNumber="448">et al.</emphasis>
(2024a: 189)
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[-13.7618°, 13.2514°,
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</paragraph>
<paragraph id="8B946307D138FFAAB0B226293A2FFB3C" blockId="9.[151,1437,1041,2040]" pageId="9" pageNumber="448">
<materialsCitation id="3B43695AD138FFAAB0B226293A29FB10" box="[199,1120,1149,1175]" collectingDate="2022-10-27" collectionCode="MUHNAC" collectorName="Mariana P. Marques &amp; Luis M. P. Ceriaco &amp; Diogo Parrinha &amp; Arthur Tiutenko" country="Angola" elevation="1614" location="Catchi village" municipality="Serra da Neve" pageId="9" pageNumber="448" specimenCode="MB03-1707" specimenCount="1" stateProvince="Namibe" typeStatus="paratype">
<emphasis id="B95FBF15D138FFAAB0B226293F0FFB10" bold="true" box="[199,326,1149,1175]" pageId="9" pageNumber="448">
<typeStatus id="5490DDA5D138FFAAB0B226293F08FB10" box="[199,321,1149,1175]" pageId="9" pageNumber="448" type="paratype">Paratypes</typeStatus>
.
</emphasis>
An
<specimenCount id="9D2DA88ED138FFAAB10D26293C4DFB10" box="[376,516,1149,1175]" pageId="9" pageNumber="448" type="adult">adult female</specimenCount>
(
<collectionCode id="ED3AFBC2D138FFAAB26126293CDEFB10" box="[532,663,1149,1175]" country="Portugal" httpUri="http://grbio.org/cool/swdk-a9xe" name="Museu Nacional de Historia Natural e de CIência" pageId="9" pageNumber="448" type="Museum">MUHNAC</collectionCode>
/
<specimenCode id="DB8DCB7CD138FFAAB2D526293D6CFB10" box="[672,805,1149,1175]" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="9" pageNumber="448">MB03-1707</specimenCode>
, field number LMPC 3121)
</materialsCitation>
and
<materialsCitation id="3B43695AD138FFAAB4ED26293A2AFB3C" collectingDate="2022-10-27" collectionCode="MUHNAC" collectorName="Mariana P. Marques &amp; Luis M. P. Ceriaco &amp; Diogo Parrinha &amp; Arthur Tiutenko" country="Angola" elevation="1614" location="Catchi village" municipality="Serra da Neve" pageId="9" pageNumber="448" specimenCode="MB03-1708" specimenCount="1" stateProvince="Namibe" typeStatus="paratype">
a juvenile (
<collectionCode id="ED3AFBC2D138FFAAB56326293BDDFB10" box="[1302,1428,1149,1175]" country="Portugal" httpUri="http://grbio.org/cool/swdk-a9xe" name="Museu Nacional de Historia Natural e de CIência" pageId="9" pageNumber="448" type="Museum">MUHNAC</collectionCode>
/
<specimenCode id="DB8DCB7CD138FFAAB0E226F53F6AFB3C" box="[151,291,1185,1211]" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="9" pageNumber="448">MB03-1708</specimenCode>
, field number LMPC 3145), with the same collecting data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331308CD138FFAAB0B226913BCAF97B" pageId="9" pageNumber="448" type="diagnosis">
<paragraph id="8B946307D138FFAAB0B226913ADFFA78" blockId="9.[151,1437,1041,2040]" pageId="9" pageNumber="448">
<emphasis id="B95FBF15D138FFAAB0B226913F0BFB58" bold="true" box="[199,322,1221,1247]" pageId="9" pageNumber="448">Diagnosis.</emphasis>
A medium sized
<taxonomicName id="4C2B1884D138FFAAB27426913CCAFB58" box="[513,643,1221,1247]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="9" pageNumber="448" phylum="Chordata" rank="genus">
<emphasis id="B95FBF15D138FFAAB27426913CCAFB58" box="[513,643,1221,1247]" italics="true" pageId="9" pageNumber="448">Rhoptropus</emphasis>
</taxonomicName>
with a maximum SVL of
<quantity id="4CD3CEE2D138FFAAB3D426913DA6FB58" box="[929,1007,1221,1248]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="448" unit="mm" value="55.0">55 mm</quantity>
(MUHNAC/MB03-1707), body stout, partly regenerated tail about the same length as SVL (
<figureCitation id="13107F82D138FFAAB28D26BD3D01FA84" box="[760,840,1257,1283]" captionStart="FIGURE 4" captionStartId="10.[152,255,1666,1690]" captionTargetBox="[151,1436,198,1643]" captionTargetId="figure-25@10.[151,1436,181,1645]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal (A) and ventral (B) views of the holotype of Rhoptropus nivimontanus sp. nov. in life (MUHNAC/MB03- 1706). Photos by AT." figureDoi="http://doi.org/10.5281/zenodo.14734256" httpUri="https://zenodo.org/record/14734256/files/figure.png" pageId="9" pageNumber="448">Figs. 4</figureCitation>
,
<figureCitation id="13107F82D138FFAAB32026BD3D2CFA83" box="[853,869,1257,1284]" captionStart="FIGURE 5" captionStartId="12.[152,255,1112,1136]" captionTargetBox="[189,1398,181,1088]" captionTargetId="figure-25@12.[189,1398,181,1088]" captionTargetPageId="12" captionText="FIGURE 5. Preserved holotype of Rhoptropus nivimontanus sp. nov. (MUHNAC/MB03-1706). Photos by DP." figureDoi="http://doi.org/10.5281/zenodo.14734258" httpUri="https://zenodo.org/record/14734258/files/figure.png" pageId="9" pageNumber="448">5</figureCitation>
). Head wide, with rounded snout. Nasal swellings separated by 2 INS; SL 9–11; IL 8. A well-defined row of enlarged chin shields behind infralabials. Dorsal scales irregularly granular, 62–65 across midbody. Precloacal scales slightly enlarged, bearing 7 pores in males. Digits relatively stout, with 16–18 subdigital scales and lamellae under fourth finger (8–10 SF4 + 8 LF4) and 19 under fourth toe (10–11 ST4 + 8–9 LT4). Tail distinctly segmented, wider at the base and smoothly tapering. Subcaudals transversely enlarged, disposed in a single row with three scales per tail segment. Dorsum greenish blue with a somewhat irregular, yellowish green pattern covering top of head, becoming darker and more reticulated from shoulders to base of tail, interspersed with scattered black speckles; ventral parts light blue.
</paragraph>
<paragraph id="8B946307D138FFAAB0B2245D3BCAF97B" blockId="9.[151,1437,1041,2040]" pageId="9" pageNumber="448">
<taxonomicName id="4C2B1884D138FFAAB0B2245D3FA0F9A4" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[199,489,1545,1571]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="9" pageNumber="448" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D138FFAAB0B2245D3FA0F9A4" box="[199,489,1545,1571]" italics="true" pageId="9" pageNumber="448">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D138FFAAB184245D3C04F9A4" bold="true" box="[497,589,1545,1571]" pageId="9" pageNumber="448">
<taxonomicNameLabel id="A26C026ED138FFAAB184245D3C04F9A4" box="[497,589,1545,1571]" pageId="9" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all its congeners in its blue and green coloration without any reddish pattern, pale or black spots on dorsum (
<emphasis id="B95FBF15D138FFAAB2CF247A3D4BF9C0" box="[698,770,1582,1607]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
dorsum pale brown to black with reddish or black pattern in other species). It further differs from
<taxonomicName id="4C2B1884D138FFAAB21424063C97F9EC" box="[609,734,1617,1643]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="9" pageNumber="448" phylum="Chordata" rank="species" species="boultoni">
<emphasis id="B95FBF15D138FFAAB21424063C97F9EC" box="[609,734,1617,1643]" italics="true" pageId="9" pageNumber="448">R. boultoni</emphasis>
</taxonomicName>
by having fewer dorsal scales across midbody (62–65
<emphasis id="B95FBF15D138FFAAB52024063BD4F9EC" box="[1365,1437,1618,1643]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
74–87); 2 internasal scales (
<emphasis id="B95FBF15D138FFAAB1A324223C57F908" box="[470,542,1654,1679]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
1), and fewer subdigital scales and lamellae under fourth finger (16–18
<emphasis id="B95FBF15D138FFAAB52024223BD4F908" box="[1365,1437,1654,1679]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
18–23) and fourth toe (19–20
<emphasis id="B95FBF15D138FFAAB19624CE3C62F934" box="[483,555,1690,1715]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
20–27); from
<taxonomicName id="4C2B1884D138FFAAB2BC24CE3D36F934" box="[713,895,1689,1715]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="9" pageNumber="448" phylum="Chordata" rank="species" species="benguellensis">
<emphasis id="B95FBF15D138FFAAB2BC24CE3D36F934" box="[713,895,1689,1715]" italics="true" pageId="9" pageNumber="448">R. benguellensis</emphasis>
</taxonomicName>
by having a blue ventrum (
<emphasis id="B95FBF15D138FFAAB4DA24CE3ABEF934" box="[1199,1271,1690,1715]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
creamy white) and 2 internasal scales (
<emphasis id="B95FBF15D138FFAAB1D924EA3FBDF950" box="[428,500,1726,1751]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
1); and from its sister species,
<taxonomicName id="4C2B1884D138FFAAB31524EA3DA7F950" box="[864,1006,1726,1751]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="9" pageNumber="448" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B95FBF15D138FFAAB31524EA3DA7F950" box="[864,1006,1726,1751]" italics="true" pageId="9" pageNumber="448">R. montanus</emphasis>
</taxonomicName>
, by having more dorsal scales across midbody (62–65
<emphasis id="B95FBF15D138FFAAB12D24B63FE9F97C" box="[344,416,1762,1787]" italics="true" pageId="9" pageNumber="448">versus</emphasis>
54–61). The male also has more precloacal pores than males of
<taxonomicName id="4C2B1884D138FFAAB40524B63AB5F97C" box="[1136,1276,1762,1787]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="9" pageNumber="448" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B95FBF15D138FFAAB40524B63AB5F97C" box="[1136,1276,1762,1787]" italics="true" pageId="9" pageNumber="448">R. montanus</emphasis>
</taxonomicName>
(7 vs. 5–6).
</paragraph>
</subSubSection>
<subSubSection id="C331308CD138FFAFB0B225513B00F908" lastPageId="12" lastPageNumber="451" pageId="9" pageNumber="448" type="description">
<paragraph id="8B946307D138FFA8B0B225513AC7FCBE" blockId="9.[151,1437,1041,2040]" lastBlockId="11.[151,1437,151,825]" lastPageId="11" lastPageNumber="450" pageId="9" pageNumber="448">
<emphasis id="B95FBF15D138FFAAB0B225513C59F898" bold="true" box="[199,528,1797,1823]" pageId="9" pageNumber="448">
Description of the
<typeStatus id="5490DDA5D138FFAAB1D725513C42F898" box="[418,523,1797,1823]" pageId="9" pageNumber="448" type="holotype">holotype</typeStatus>
.
</emphasis>
Mensural and meristic characters for the
<typeStatus id="5490DDA5D138FFAAB3AA25513A08F898" box="[991,1089,1797,1823]" pageId="9" pageNumber="448" type="holotype">holotype</typeStatus>
are presented in
<tableCitation id="C6A956BCD138FFAAB48B25513B1BF898" box="[1278,1362,1797,1823]" captionStart="TABLE 3" captionStartId="11.[152,244,872,896]" captionTargetPageId="11" captionText="TABLE 3. Mensural and meristic data for the type series of Rhoptropus nivimontanus sp. nov.. Abbreviations are the ones listed in the materials and methods." pageId="9" pageNumber="448">Table 3</tableCitation>
. Adult male with a nearly complete, original tail (
<figureCitation id="13107F82D138FFAAB20E257D3CA0F8C3" box="[635,745,1833,1860]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="10.[152,255,1666,1690]" captionStartId-1="12.[152,255,1112,1136]" captionTargetBox-0="[151,1436,198,1643]" captionTargetBox-1="[189,1398,181,1088]" captionTargetId-0="figure-25@10.[151,1436,181,1645]" captionTargetId-1="figure-25@12.[189,1398,181,1088]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionText-0="FIGURE 4. Dorsal (A) and ventral (B) views of the holotype of Rhoptropus nivimontanus sp. nov. in life (MUHNAC/MB03- 1706). Photos by AT." captionText-1="FIGURE 5. Preserved holotype of Rhoptropus nivimontanus sp. nov. (MUHNAC/MB03-1706). Photos by DP." figureDoi-0="http://doi.org/10.5281/zenodo.14734256" figureDoi-1="http://doi.org/10.5281/zenodo.14734258" httpUri-0="https://zenodo.org/record/14734256/files/figure.png" httpUri-1="https://zenodo.org/record/14734258/files/figure.png" pageId="9" pageNumber="448">Figs. 4–5</figureCitation>
). Body robust (AGD/SVL 0.41). Head moderately long (HL/ SVL 0.29), wide (HW/HL 0.73), weakly depressed (HH/HL 0.36), distinct from neck. Snout medium (ES/HL 0.43), rounded, with blunt profile. Canthus rostralis not prominent; shallow depression present immediately behind nasal swellings. Eye small (ED/HL 0.17), its diameter shorter than distance from eye to tip of snout (ED/ES 0.39). Supracilliaries small and slightly pointed; no supraciliary dermal fold present. Ear opening vertically ovoid, situated below eye level, smaller than eye diameter (EH/ED 0.56), with small dermal fold extending from superior margin; distance from eye to ear greater than eye diameter (EE/ED 1.71). Rostral wider than deep, with small backwards prolongation between nasal swellings, in contact with first supralabial and two nasals at each side, and two internasal scales. Nostril situated between three weakly swollen circumnasal scales, oriented dorsally; nasal swellings separated by two INS, followed posteriorly by three smaller scales. Scales on snout larger than those on dorsum and occiput; largest located in front of eye, slightly conical and keeled. Mental greatly elongated and tapering to a point posteriorly, in contact with first infralabial on each side and median chin shield posteriorly. A well-defined row of enlarged chin shields is present, bordering mental and first three infralabials on each side, extending obliquely backwards along gular region. Gular scales irregular in size, those immediately posterior to chin shields slightly larger than those on throat. SL 11/10 (R/L); IL 8/8 (R/L), first two greatly elongated; small intrusive scale present between IL III and IV on left side. Dorsal scales small and granular, rounded, with weak tendency to become larger towards flanks, 65 across midbody. Ventral scales larger than dorsals, smooth, rounded to sub-hexagonal, slightly imbricate on chest. Precloacal scales slightly enlarged, rounded to sub-hexagonal, bearing 7 continuous pores disposed in a chevron. Limbs long and stout (FLL/SVL 0.15, CL/SVL 0.18); scales on limbs smooth and juxtaposed; those on anterior and ventral aspect of hindlimbs distinctly larger, as well as those on anterior aspect of forearm; scales on palms and soles small and rounded. Digits relatively stout, wider on distal portion; covered above with somewhat enlarged scales distally; clawless or with small triangular scale in place of claw. Ventral surface of digits covered with enlarged scales along the midline (non-scansorial), followed by undivided (except for distalmost) transverse lamellae under distal expansion. Fourth finger of right manus with 10 enlarged scales along midline and 8 transverse lamellae on distal expansion; fourth toe of right pes with 11 enlarged scales along midline and 9 transverse lamellae on distal expansion. Relative length of digits III&gt;IV&gt;II&gt;I=V (right manus); III=IV&gt;V&gt;II&gt;I (right pes). Tail slightly shorter than SVL (TAL/SVL 0.84), last
<quantity id="4CD3CEE2D13AFFA8B222203F3CD0FD02" box="[599,665,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="450" unit="mm" value="2.0">2 mm</quantity>
regenerated, weakly depressed, wider at base, tapering to point. Tail segmented, with distinct lateral constrictions between proximal segments, less conspicuous distally; dorsal surface covered with smooth, granular scales disposed in rows, slightly larger than dorsals; subcaudal scales irregularly enlarged in first 3 segments, fourth segment with 3 pairs of enlarged scales on midventral section and following segments with a single row of transversely enlarged scales, disposed in 3 scales per tail segment. Regenerated portion of tail unsegmented, with a single row of enlarged scales along midventral section.
</paragraph>
<caption id="DF54338FD13BFFA9B0ED24D63F24F938" ID-DOI="http://doi.org/10.5281/zenodo.14734256" ID-Zenodo-Dep="14734256" httpUri="https://zenodo.org/record/14734256/files/figure.png" pageId="10" pageNumber="449" startId="10.[152,255,1666,1690]" targetBox="[151,1436,198,1643]" targetPageId="10" targetType="figure">
<paragraph id="8B946307D13BFFA9B0ED24D63F24F938" blockId="10.[151,1437,1666,1727]" pageId="10" pageNumber="449">
<emphasis id="B95FBF15D13BFFA9B0ED24D63F51F91D" bold="true" box="[152,280,1666,1690]" pageId="10" pageNumber="449">FIGURE 4.</emphasis>
Dorsal (
<emphasis id="B95FBF15D13BFFA9B10424D63FCAF91D" bold="true" box="[369,387,1666,1690]" pageId="10" pageNumber="449">A</emphasis>
) and ventral (
<emphasis id="B95FBF15D13BFFA9B26724D73C6AF91C" bold="true" box="[530,547,1667,1691]" pageId="10" pageNumber="449">B</emphasis>
) views of the holotype of
<taxonomicName id="4C2B1884D13BFFA9B35A24D73A7AF91D" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[815,1075,1667,1691]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="10" pageNumber="449" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13BFFA9B35A24D73A7AF91D" box="[815,1075,1667,1691]" italics="true" pageId="10" pageNumber="449">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13BFFA9B44C24D73AC3F91C" bold="true" box="[1081,1162,1667,1691]" pageId="10" pageNumber="449">
<taxonomicNameLabel id="A26C026ED13BFFA9B44C24D73AC3F91C" box="[1081,1162,1667,1691]" pageId="10" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (MUHNAC/MB03- 1706). Photos by AT.
</paragraph>
</caption>
<caption id="DF54338FD13AFFA8B0ED213C3C00FC23" pageId="11" pageNumber="450" startId="11.[152,244,872,896]" targetBox="[163,1397,949,2019]" targetIsTable="true" targetPageId="11" targetType="table">
<paragraph id="8B946307D13AFFA8B0ED213C3C00FC23" blockId="11.[151,1437,872,932]" pageId="11" pageNumber="450">
<emphasis id="B95FBF15D13AFFA8B0ED213C3F59FC07" bold="true" box="[152,272,872,896]" pageId="11" pageNumber="450">TABLE 3.</emphasis>
Mensural and meristic data for the type series of
<taxonomicName id="4C2B1884D13AFFA8B346213C3A0EFC07" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[819,1095,872,896]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="11" pageNumber="450" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13AFFA8B346213C3A0EFC07" box="[819,1095,872,896]" italics="true" pageId="11" pageNumber="450">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13AFFA8B43A213D3AEFFC06" bold="true" box="[1103,1190,873,897]" pageId="11" pageNumber="450">
<taxonomicNameLabel id="A26C026ED13AFFA8B43A213D3AEFFC06" box="[1103,1190,873,897]" pageId="11" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Abbreviations are the ones listed in the materials and methods.
</paragraph>
</caption>
<paragraph id="8B946307D13AFFA8B18A21E13B41F864" pageId="11" pageNumber="450">
<table id="F92B91A7D13A005CB0D621E13B3CF864" box="[163,1397,949,2019]" gridcols="4" gridrows="27" pageId="11" pageNumber="450">
<tr id="351B6145D13A005CB0D621E13B3CFC4A" box="[163,1397,949,973]" gridrow="0" pageId="11" pageNumber="450" rowspan-0="1">
<th id="76CA0839D13A005CB18421E13D4BFC4A" box="[497,770,949,973]" gridcol="1" gridrow="0" pageId="11" pageNumber="450">MUHNAC/MB03-1706</th>
<th id="76CA0839D13A005CB33421E13A7DFC4A" box="[833,1076,949,973]" gridcol="2" gridrow="0" pageId="11" pageNumber="450">MUHNAC/MB03-1707</th>
<th id="76CA0839D13A005CB4F721E13B3CFC4A" box="[1154,1397,949,973]" gridcol="3" gridrow="0" pageId="11" pageNumber="450">MUHNAC/MB03-1708</th>
</tr>
<tr id="351B6145D13A005CB0D621893B3CFC72" box="[163,1397,989,1013]" gridrow="1" pageId="11" pageNumber="450" rowspan-0="1">
<td id="76CA0839D13A005CB18421893D4BFC72" box="[497,770,989,1013]" gridcol="1" gridrow="1" pageId="11" pageNumber="450">Holotype</td>
<td id="76CA0839D13A005CB33421893A7DFC72" box="[833,1076,989,1013]" gridcol="2" gridrow="1" pageId="11" pageNumber="450">Paratype</td>
<td id="76CA0839D13A005CB4F721893B3CFC72" box="[1154,1397,989,1013]" gridcol="3" gridrow="1" pageId="11" pageNumber="450">Paratype</td>
</tr>
<tr id="351B6145D13A005CB0D626513B3CFB9A" box="[163,1397,1029,1053]" gridrow="2" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D626513F4DFB9A" box="[163,260,1029,1053]" gridcol="0" gridrow="2" pageId="11" pageNumber="450">Sex</th>
<td id="76CA0839D13A005CB18426513D4BFB9A" box="[497,770,1029,1053]" gridcol="1" gridrow="2" pageId="11" pageNumber="450">Male</td>
<td id="76CA0839D13A005CB33426513A7DFB9A" box="[833,1076,1029,1053]" gridcol="2" gridrow="2" pageId="11" pageNumber="450">Female</td>
<td id="76CA0839D13A005CB4F726513B3CFB9A" box="[1154,1397,1029,1053]" gridcol="3" gridrow="2" pageId="11" pageNumber="450">Juvenile</td>
</tr>
<tr id="351B6145D13A005CB0D626793B3CFBC1" box="[163,1397,1069,1094]" gridrow="3" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D626793F4DFBC1" box="[163,260,1069,1094]" gridcol="0" gridrow="3" pageId="11" pageNumber="450">SVL</th>
<td id="76CA0839D13A005CB18426793D4BFBC1" box="[497,770,1069,1094]" gridcol="1" gridrow="3" pageId="11" pageNumber="450">52.4</td>
<td id="76CA0839D13A005CB33426793A7DFBC1" box="[833,1076,1069,1094]" gridcol="2" gridrow="3" pageId="11" pageNumber="450">55.3</td>
<td id="76CA0839D13A005CB4F726793B3CFBC1" box="[1154,1397,1069,1094]" gridcol="3" gridrow="3" pageId="11" pageNumber="450">31.7</td>
</tr>
<tr id="351B6145D13A005CB0D626013B3CFBE9" box="[163,1397,1109,1134]" gridrow="4" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D626013F4DFBE9" box="[163,260,1109,1134]" gridcol="0" gridrow="4" pageId="11" pageNumber="450">AGD</th>
<td id="76CA0839D13A005CB18426013D4BFBE9" box="[497,770,1109,1134]" gridcol="1" gridrow="4" pageId="11" pageNumber="450">21.5</td>
<td id="76CA0839D13A005CB33426013A7DFBE9" box="[833,1076,1109,1134]" gridcol="2" gridrow="4" pageId="11" pageNumber="450">21.2</td>
<td id="76CA0839D13A005CB4F726013B3CFBE9" box="[1154,1397,1109,1134]" gridcol="3" gridrow="4" pageId="11" pageNumber="450">12</td>
</tr>
<tr id="351B6145D13A005CB0D6262A3B3CFB11" box="[163,1397,1150,1174]" gridrow="5" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6262A3F4DFB11" box="[163,260,1150,1174]" gridcol="0" gridrow="5" pageId="11" pageNumber="450">BW</th>
<td id="76CA0839D13A005CB184262A3D4BFB11" box="[497,770,1150,1174]" gridcol="1" gridrow="5" pageId="11" pageNumber="450">13.6</td>
<td id="76CA0839D13A005CB334262A3A7DFB11" box="[833,1076,1150,1174]" gridcol="2" gridrow="5" pageId="11" pageNumber="450">15</td>
<td id="76CA0839D13A005CB4F7262A3B3CFB11" box="[1154,1397,1150,1174]" gridcol="3" gridrow="5" pageId="11" pageNumber="450">7.8</td>
</tr>
<tr id="351B6145D13A005CB0D626F23B3CFB39" box="[163,1397,1190,1214]" gridrow="6" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D626F23F4DFB39" box="[163,260,1190,1214]" gridcol="0" gridrow="6" pageId="11" pageNumber="450">TAL</th>
<td id="76CA0839D13A005CB18426F23D4BFB39" box="[497,770,1190,1214]" gridcol="1" gridrow="6" pageId="11" pageNumber="450">44 (last 2 mm regenerated)</td>
<td id="76CA0839D13A005CB33426F23A7DFB39" box="[833,1076,1190,1214]" gridcol="2" gridrow="6" pageId="11" pageNumber="450">-</td>
<td id="76CA0839D13A005CB4F726F23B3CFB39" box="[1154,1397,1190,1214]" gridcol="3" gridrow="6" pageId="11" pageNumber="450">-</td>
</tr>
<tr id="351B6145D13A005CB0D6269A3B3CFB61" box="[163,1397,1230,1254]" gridrow="7" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6269A3F4DFB61" box="[163,260,1230,1254]" gridcol="0" gridrow="7" pageId="11" pageNumber="450">TW</th>
<td id="76CA0839D13A005CB184269A3D4BFB61" box="[497,770,1230,1254]" gridcol="1" gridrow="7" pageId="11" pageNumber="450">6.2</td>
<td id="76CA0839D13A005CB334269A3A7DFB61" box="[833,1076,1230,1254]" gridcol="2" gridrow="7" pageId="11" pageNumber="450">6.3</td>
<td id="76CA0839D13A005CB4F7269A3B3CFB61" box="[1154,1397,1230,1254]" gridcol="3" gridrow="7" pageId="11" pageNumber="450">2.8</td>
</tr>
<tr id="351B6145D13A005CB0D626A33B3CFA88" box="[163,1397,1271,1295]" gridrow="8" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D626A33F4DFA88" box="[163,260,1271,1295]" gridcol="0" gridrow="8" pageId="11" pageNumber="450">FLL</th>
<td id="76CA0839D13A005CB18426A33D4BFA88" box="[497,770,1271,1295]" gridcol="1" gridrow="8" pageId="11" pageNumber="450">7.6</td>
<td id="76CA0839D13A005CB33426A33A7DFA88" box="[833,1076,1271,1295]" gridcol="2" gridrow="8" pageId="11" pageNumber="450">8.1</td>
<td id="76CA0839D13A005CB4F726A33B3CFA88" box="[1154,1397,1271,1295]" gridcol="3" gridrow="8" pageId="11" pageNumber="450">4.7</td>
</tr>
<tr id="351B6145D13A005CB0D6274B3B3CFAB0" box="[163,1397,1311,1335]" gridrow="9" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6274B3F4DFAB0" box="[163,260,1311,1335]" gridcol="0" gridrow="9" pageId="11" pageNumber="450">CL</th>
<td id="76CA0839D13A005CB184274B3D4BFAB0" box="[497,770,1311,1335]" gridcol="1" gridrow="9" pageId="11" pageNumber="450">9.7</td>
<td id="76CA0839D13A005CB334274B3A7DFAB0" box="[833,1076,1311,1335]" gridcol="2" gridrow="9" pageId="11" pageNumber="450">9.9</td>
<td id="76CA0839D13A005CB4F7274B3B3CFAB0" box="[1154,1397,1311,1335]" gridcol="3" gridrow="9" pageId="11" pageNumber="450">5.9</td>
</tr>
<tr id="351B6145D13A005CB0D627133B3CFAD8" box="[163,1397,1351,1375]" gridrow="10" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D627133F4DFAD8" box="[163,260,1351,1375]" gridcol="0" gridrow="10" pageId="11" pageNumber="450">HL</th>
<td id="76CA0839D13A005CB18427133D4BFAD8" box="[497,770,1351,1375]" gridcol="1" gridrow="10" pageId="11" pageNumber="450">15</td>
<td id="76CA0839D13A005CB33427133A7DFAD8" box="[833,1076,1351,1375]" gridcol="2" gridrow="10" pageId="11" pageNumber="450">16.3</td>
<td id="76CA0839D13A005CB4F727133B3CFAD8" box="[1154,1397,1351,1375]" gridcol="3" gridrow="10" pageId="11" pageNumber="450">9.7</td>
</tr>
<tr id="351B6145D13A005CB0D6273B3B3CFA00" box="[163,1397,1391,1415]" gridrow="11" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6273B3F4DFA00" box="[163,260,1391,1415]" gridcol="0" gridrow="11" pageId="11" pageNumber="450">HH</th>
<td id="76CA0839D13A005CB184273B3D4BFA00" box="[497,770,1391,1415]" gridcol="1" gridrow="11" pageId="11" pageNumber="450">5.4</td>
<td id="76CA0839D13A005CB334273B3A7DFA00" box="[833,1076,1391,1415]" gridcol="2" gridrow="11" pageId="11" pageNumber="450">5.4</td>
<td id="76CA0839D13A005CB4F7273B3B3CFA00" box="[1154,1397,1391,1415]" gridcol="3" gridrow="11" pageId="11" pageNumber="450">3.5</td>
</tr>
<tr id="351B6145D13A005CB0D627C33B3CFA37" box="[163,1397,1431,1456]" gridrow="12" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D627C33F4DFA37" box="[163,260,1431,1456]" gridcol="0" gridrow="12" pageId="11" pageNumber="450">HW</th>
<td id="76CA0839D13A005CB18427C33D4BFA37" box="[497,770,1431,1456]" gridcol="1" gridrow="12" pageId="11" pageNumber="450">11</td>
<td id="76CA0839D13A005CB33427C33A7DFA37" box="[833,1076,1431,1456]" gridcol="2" gridrow="12" pageId="11" pageNumber="450">11.6</td>
<td id="76CA0839D13A005CB4F727C33B3CFA37" box="[1154,1397,1431,1456]" gridcol="3" gridrow="12" pageId="11" pageNumber="450">7.1</td>
</tr>
<tr id="351B6145D13A005CB0D627943B3CFA5F" box="[163,1397,1472,1496]" gridrow="13" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D627943F4DFA5F" box="[163,260,1472,1496]" gridcol="0" gridrow="13" pageId="11" pageNumber="450">ED</th>
<td id="76CA0839D13A005CB18427943D4BFA5F" box="[497,770,1472,1496]" gridcol="1" gridrow="13" pageId="11" pageNumber="450">2.5</td>
<td id="76CA0839D13A005CB33427943A7DFA5F" box="[833,1076,1472,1496]" gridcol="2" gridrow="13" pageId="11" pageNumber="450">2.7</td>
<td id="76CA0839D13A005CB4F727943B3CFA5F" box="[1154,1397,1472,1496]" gridcol="3" gridrow="13" pageId="11" pageNumber="450">1.8</td>
</tr>
<tr id="351B6145D13A005CB0D627BC3B3CF987" box="[163,1397,1512,1536]" gridrow="14" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D627BC3F4DF987" box="[163,260,1512,1536]" gridcol="0" gridrow="14" pageId="11" pageNumber="450">EN</th>
<td id="76CA0839D13A005CB18427BC3D4BF987" box="[497,770,1512,1536]" gridcol="1" gridrow="14" pageId="11" pageNumber="450">4.8</td>
<td id="76CA0839D13A005CB33427BC3A7DF987" box="[833,1076,1512,1536]" gridcol="2" gridrow="14" pageId="11" pageNumber="450">5.1</td>
<td id="76CA0839D13A005CB4F727BC3B3CF987" box="[1154,1397,1512,1536]" gridcol="3" gridrow="14" pageId="11" pageNumber="450">2.9</td>
</tr>
<tr id="351B6145D13A005CB0D624443B3CF9AF" box="[163,1397,1552,1576]" gridrow="15" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D624443F4DF9AF" box="[163,260,1552,1576]" gridcol="0" gridrow="15" pageId="11" pageNumber="450">ES</th>
<td id="76CA0839D13A005CB18424443D4BF9AF" box="[497,770,1552,1576]" gridcol="1" gridrow="15" pageId="11" pageNumber="450">6.4</td>
<td id="76CA0839D13A005CB33424443A7DF9AF" box="[833,1076,1552,1576]" gridcol="2" gridrow="15" pageId="11" pageNumber="450">6.6</td>
<td id="76CA0839D13A005CB4F724443B3CF9AF" box="[1154,1397,1552,1576]" gridcol="3" gridrow="15" pageId="11" pageNumber="450">4</td>
</tr>
<tr id="351B6145D13A005CB0D6246D3B3CF9D6" box="[163,1397,1593,1617]" gridrow="16" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6246D3F4DF9D6" box="[163,260,1593,1617]" gridcol="0" gridrow="16" pageId="11" pageNumber="450">EE</th>
<td id="76CA0839D13A005CB184246D3D4BF9D6" box="[497,770,1593,1617]" gridcol="1" gridrow="16" pageId="11" pageNumber="450">4.3</td>
<td id="76CA0839D13A005CB334246D3A7DF9D6" box="[833,1076,1593,1617]" gridcol="2" gridrow="16" pageId="11" pageNumber="450">4.1</td>
<td id="76CA0839D13A005CB4F7246D3B3CF9D6" box="[1154,1397,1593,1617]" gridcol="3" gridrow="16" pageId="11" pageNumber="450">2.4</td>
</tr>
<tr id="351B6145D13A005CB0D624353B3CF9FE" box="[163,1397,1633,1657]" gridrow="17" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D624353F4DF9FE" box="[163,260,1633,1657]" gridcol="0" gridrow="17" pageId="11" pageNumber="450">IN</th>
<td id="76CA0839D13A005CB18424353D4BF9FE" box="[497,770,1633,1657]" gridcol="1" gridrow="17" pageId="11" pageNumber="450">1.4</td>
<td id="76CA0839D13A005CB33424353A7DF9FE" box="[833,1076,1633,1657]" gridcol="2" gridrow="17" pageId="11" pageNumber="450">1.4</td>
<td id="76CA0839D13A005CB4F724353B3CF9FE" box="[1154,1397,1633,1657]" gridcol="3" gridrow="17" pageId="11" pageNumber="450">0.8</td>
</tr>
<tr id="351B6145D13A005CB0D624DD3B3CF926" box="[163,1397,1673,1697]" gridrow="18" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D624DD3F4DF926" box="[163,260,1673,1697]" gridcol="0" gridrow="18" pageId="11" pageNumber="450">IOD</th>
<td id="76CA0839D13A005CB18424DD3D4BF926" box="[497,770,1673,1697]" gridcol="1" gridrow="18" pageId="11" pageNumber="450">1.4</td>
<td id="76CA0839D13A005CB33424DD3A7DF926" box="[833,1076,1673,1697]" gridcol="2" gridrow="18" pageId="11" pageNumber="450">1.7</td>
<td id="76CA0839D13A005CB4F724DD3B3CF926" box="[1154,1397,1673,1697]" gridcol="3" gridrow="18" pageId="11" pageNumber="450">1</td>
</tr>
<tr id="351B6145D13A005CB0D624E53B3CF94E" box="[163,1397,1713,1737]" gridrow="19" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D624E53F4DF94E" box="[163,260,1713,1737]" gridcol="0" gridrow="19" pageId="11" pageNumber="450">PCP</th>
<td id="76CA0839D13A005CB18424E53D4BF94E" box="[497,770,1713,1737]" gridcol="1" gridrow="19" pageId="11" pageNumber="450">7</td>
<td id="76CA0839D13A005CB33424E53A7DF94E" box="[833,1076,1713,1737]" gridcol="2" gridrow="19" pageId="11" pageNumber="450">-</td>
<td id="76CA0839D13A005CB4F724E53B3CF94E" box="[1154,1397,1713,1737]" gridcol="3" gridrow="19" pageId="11" pageNumber="450">-</td>
</tr>
<tr id="351B6145D13A005CB0D6248E3B3CF975" box="[163,1397,1754,1778]" gridrow="20" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6248E3F4DF975" box="[163,260,1754,1778]" gridcol="0" gridrow="20" pageId="11" pageNumber="450">SL</th>
<td id="76CA0839D13A005CB184248E3D4BF975" box="[497,770,1754,1778]" gridcol="1" gridrow="20" pageId="11" pageNumber="450">11</td>
<td id="76CA0839D13A005CB334248E3A7DF975" box="[833,1076,1754,1778]" gridcol="2" gridrow="20" pageId="11" pageNumber="450">11</td>
<td id="76CA0839D13A005CB4F7248E3B3CF975" box="[1154,1397,1754,1778]" gridcol="3" gridrow="20" pageId="11" pageNumber="450">9</td>
</tr>
<tr id="351B6145D13A005CB0D625563B3CF89D" box="[163,1397,1794,1818]" gridrow="21" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D625563F4DF89D" box="[163,260,1794,1818]" gridcol="0" gridrow="21" pageId="11" pageNumber="450">IL</th>
<td id="76CA0839D13A005CB18425563D4BF89D" box="[497,770,1794,1818]" gridcol="1" gridrow="21" pageId="11" pageNumber="450">8</td>
<td id="76CA0839D13A005CB33425563A7DF89D" box="[833,1076,1794,1818]" gridcol="2" gridrow="21" pageId="11" pageNumber="450">8</td>
<td id="76CA0839D13A005CB4F725563B3CF89D" box="[1154,1397,1794,1818]" gridcol="3" gridrow="21" pageId="11" pageNumber="450">8</td>
</tr>
<tr id="351B6145D13A005CB0D6257E3B3CF8C5" box="[163,1397,1834,1858]" gridrow="22" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6257E3F4DF8C5" box="[163,260,1834,1858]" gridcol="0" gridrow="22" pageId="11" pageNumber="450">CN</th>
<td id="76CA0839D13A005CB184257E3D4BF8C5" box="[497,770,1834,1858]" gridcol="1" gridrow="22" pageId="11" pageNumber="450">3</td>
<td id="76CA0839D13A005CB334257E3A7DF8C5" box="[833,1076,1834,1858]" gridcol="2" gridrow="22" pageId="11" pageNumber="450">3</td>
<td id="76CA0839D13A005CB4F7257E3B3CF8C5" box="[1154,1397,1834,1858]" gridcol="3" gridrow="22" pageId="11" pageNumber="450">3</td>
</tr>
<tr id="351B6145D13A005CB0D625063B3CF8ED" box="[163,1397,1874,1898]" gridrow="23" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D625063F4DF8ED" box="[163,260,1874,1898]" gridcol="0" gridrow="23" pageId="11" pageNumber="450">INS</th>
<td id="76CA0839D13A005CB18425063D4BF8ED" box="[497,770,1874,1898]" gridcol="1" gridrow="23" pageId="11" pageNumber="450">2</td>
<td id="76CA0839D13A005CB33425063A7DF8ED" box="[833,1076,1874,1898]" gridcol="2" gridrow="23" pageId="11" pageNumber="450">2</td>
<td id="76CA0839D13A005CB4F725063B3CF8ED" box="[1154,1397,1874,1898]" gridcol="3" gridrow="23" pageId="11" pageNumber="450">2</td>
</tr>
<tr id="351B6145D13A005CB0D6252F3B3CF814" box="[163,1397,1915,1939]" gridrow="24" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6252F3F4DF814" box="[163,260,1915,1939]" gridcol="0" gridrow="24" pageId="11" pageNumber="450">SF4+LF4</th>
<td id="76CA0839D13A005CB184252F3D4BF814" box="[497,770,1915,1939]" gridcol="1" gridrow="24" pageId="11" pageNumber="450">10+8</td>
<td id="76CA0839D13A005CB334252F3A7DF814" box="[833,1076,1915,1939]" gridcol="2" gridrow="24" pageId="11" pageNumber="450">9+8</td>
<td id="76CA0839D13A005CB4F7252F3B3CF814" box="[1154,1397,1915,1939]" gridcol="3" gridrow="24" pageId="11" pageNumber="450">8+8</td>
</tr>
<tr id="351B6145D13A005CB0D625F73B3CF83C" box="[163,1397,1955,1979]" gridrow="25" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D625F73F4DF83C" box="[163,260,1955,1979]" gridcol="0" gridrow="25" pageId="11" pageNumber="450">ST4+LT4</th>
<td id="76CA0839D13A005CB18425F73D4BF83C" box="[497,770,1955,1979]" gridcol="1" gridrow="25" pageId="11" pageNumber="450">11+9</td>
<td id="76CA0839D13A005CB33425F73A7DF83C" box="[833,1076,1955,1979]" gridcol="2" gridrow="25" pageId="11" pageNumber="450">10+9</td>
<td id="76CA0839D13A005CB4F725F73B3CF83C" box="[1154,1397,1955,1979]" gridcol="3" gridrow="25" pageId="11" pageNumber="450">11+8</td>
</tr>
<tr id="351B6145D13A005CB0D6259F3B3CF864" box="[163,1397,1995,2019]" gridrow="26" pageId="11" pageNumber="450">
<th id="76CA0839D13A005CB0D6259F3F4DF864" box="[163,260,1995,2019]" gridcol="0" gridrow="26" pageId="11" pageNumber="450">DSR</th>
<td id="76CA0839D13A005CB184259F3D4BF864" box="[497,770,1995,2019]" gridcol="1" gridrow="26" pageId="11" pageNumber="450">65</td>
<td id="76CA0839D13A005CB334259F3A7DF864" box="[833,1076,1995,2019]" gridcol="2" gridrow="26" pageId="11" pageNumber="450">62</td>
<td id="76CA0839D13A005CB4F7259F3B3CF864" box="[1154,1397,1995,2019]" gridcol="3" gridrow="26" pageId="11" pageNumber="450">63</td>
</tr>
</table>
</paragraph>
<caption id="DF54338FD13DFFAFB0ED260C3B4FFBE8" ID-DOI="http://doi.org/10.5281/zenodo.14734258" ID-Zenodo-Dep="14734258" box="[152,1286,1112,1136]" httpUri="https://zenodo.org/record/14734258/files/figure.png" pageId="12" pageNumber="451" startId="12.[152,255,1112,1136]" targetBox="[189,1398,181,1088]" targetPageId="12" targetType="figure">
<paragraph id="8B946307D13DFFAFB0ED260C3B4FFBE8" blockId="12.[152,1286,1112,1136]" box="[152,1286,1112,1136]" pageId="12" pageNumber="451">
<emphasis id="B95FBF15D13DFFAFB0ED260C3F51FBF7" bold="true" box="[152,280,1112,1136]" pageId="12" pageNumber="451">FIGURE 5.</emphasis>
Preserved holotype of
<taxonomicName id="4C2B1884D13DFFAFB276260C3D41FBF7" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[515,776,1112,1136]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="12" pageNumber="451" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13DFFAFB276260C3D41FBF7" box="[515,776,1112,1136]" italics="true" pageId="12" pageNumber="451">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13DFFAFB37B260C3D17FBF7" bold="true" box="[782,862,1112,1136]" pageId="12" pageNumber="451">
<taxonomicNameLabel id="A26C026ED13DFFAFB37B260C3D17FBF7" box="[782,862,1112,1136]" pageId="12" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MUHNAC/MB03-1706). Photos by DP.
</paragraph>
</caption>
<paragraph id="8B946307D13DFFAFB0B226903AA8FAE9" blockId="12.[151,1437,1220,2039]" pageId="12" pageNumber="451">
<emphasis id="B95FBF15D13DFFAFB0B226903FD9FB59" bold="true" box="[199,400,1220,1246]" pageId="12" pageNumber="451">Coloration in life</emphasis>
(
<figureCitation id="13107F82D13DFFAFB1EA26903FAAFB58" box="[415,483,1220,1247]" captionStart="FIGURE 4" captionStartId="10.[152,255,1666,1690]" captionTargetBox="[151,1436,198,1643]" captionTargetId="figure-25@10.[151,1436,181,1645]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal (A) and ventral (B) views of the holotype of Rhoptropus nivimontanus sp. nov. in life (MUHNAC/MB03- 1706). Photos by AT." figureDoi="http://doi.org/10.5281/zenodo.14734256" httpUri="https://zenodo.org/record/14734256/files/figure.png" pageId="12" pageNumber="451">Fig. 4</figureCitation>
): Dorsum greenish blue; a light yellowish green hue begins behind the nasal swellings and extends through interorbital region to occiput, where it is more distinct; green pattern becomes progressively darker and more reticulated from shoulders towards base of tail, interspersed with scattered black speckles. Limbs mostly homogeneous, with faint and irregular dark shades on hindlimbs; enlarged scales over distal expansion of digits black centered. Ventral parts light blue, slightly darker on precloacal and subcaudal regions.
</paragraph>
<paragraph id="8B946307D13DFFAFB0B2272C3C14FA79" blockId="12.[151,1437,1220,2039]" pageId="12" pageNumber="451">
<emphasis id="B95FBF15D13DFFAFB0B2272C3C48FA15" bold="true" box="[199,513,1400,1426]" pageId="12" pageNumber="451">Coloration in preservative</emphasis>
(
<figureCitation id="13107F82D13DFFAFB264272C3C1FFA14" box="[529,598,1400,1427]" captionStart="FIGURE 5" captionStartId="12.[152,255,1112,1136]" captionTargetBox="[189,1398,181,1088]" captionTargetId="figure-25@12.[189,1398,181,1088]" captionTargetPageId="12" captionText="FIGURE 5. Preserved holotype of Rhoptropus nivimontanus sp. nov. (MUHNAC/MB03-1706). Photos by DP." figureDoi="http://doi.org/10.5281/zenodo.14734258" httpUri="https://zenodo.org/record/14734258/files/figure.png" pageId="12" pageNumber="451">Fig. 5</figureCitation>
): Dorsum dark greyish blue; green pattern on dorsum reduced to dull dark brown; on dorsal aspect of head, light green pattern replaced by a distinct black patch, starting behind internasal granules and between postnasal depressions, extending through forehead and interorbital region above frontal bone and fading at occiput. Ventral parts blue.
</paragraph>
<paragraph id="8B946307D13DFFAFB0B2245C3B00F908" blockId="12.[151,1437,1220,2039]" pageId="12" pageNumber="451">
<emphasis id="B95FBF15D13DFFAFB0B2245C3F76F9A5" bold="true" box="[199,319,1544,1570]" pageId="12" pageNumber="451">Variation.</emphasis>
Mensural and meristic variation among the type series is presented in
<tableCitation id="C6A956BCD13DFFAFB425245C3AECF9A4" box="[1104,1189,1544,1571]" captionStart="TABLE 3" captionStartId="11.[152,244,872,896]" captionTargetPageId="11" captionText="TABLE 3. Mensural and meristic data for the type series of Rhoptropus nivimontanus sp. nov.. Abbreviations are the ones listed in the materials and methods." pageId="12" pageNumber="451">Table 3</tableCitation>
. Both
<typeStatus id="5490DDA5D13DFFAFB49B245D3B10F9A4" box="[1262,1369,1545,1571]" pageId="12" pageNumber="451" type="paratype">paratypes</typeStatus>
agree entirely with the
<typeStatus id="5490DDA5D13DFFAFB12124783FFFF9C1" box="[340,438,1580,1606]" pageId="12" pageNumber="451" type="holotype">holotype</typeStatus>
in terms of coloration. Female
<typeStatus id="5490DDA5D13DFFAFB36624793D3AF9C0" box="[787,883,1581,1607]" pageId="12" pageNumber="451" type="paratype">paratype</typeStatus>
<collectionCode id="ED3AFBC2D13DFFAFB30F24793DB2F9C0" box="[890,1019,1581,1607]" country="Portugal" httpUri="http://grbio.org/cool/swdk-a9xe" name="Museu Nacional de Historia Natural e de CIência" pageId="12" pageNumber="451" type="Museum">MUHNAC</collectionCode>
/
<specimenCode id="DB8DCB7CD13DFFAFB47124793ACEF9C0" box="[1028,1159,1581,1607]" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="12" pageNumber="451">MB03-1707</specimenCode>
lacking precloacal pores or pits; mostly clawless, bearing minute claws on digits I and
<collectionCode id="ED3AFBC2D13DFFAFB32224053D22F9EC" box="[855,875,1617,1643]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="451" type="Museum">V</collectionCode>
of left pes. Tail broken in both
<typeStatus id="5490DDA5D13DFFAFB4A124053B76F9EC" box="[1236,1343,1617,1643]" pageId="12" pageNumber="451" type="paratype">paratypes</typeStatus>
prior to collection, after first (
<collectionCode id="ED3AFBC2D13DFFAFB1FF24213C44F908" box="[394,525,1653,1679]" country="Portugal" httpUri="http://grbio.org/cool/swdk-a9xe" name="Museu Nacional de Historia Natural e de CIência" pageId="12" pageNumber="451" type="Museum">MUHNAC</collectionCode>
/
<specimenCode id="DB8DCB7CD13DFFAFB26024213CD3F908" box="[533,666,1653,1679]" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="12" pageNumber="451">MB03-1708</specimenCode>
) and third (
<collectionCode id="ED3AFBC2D13DFFAFB36924213DD6F908" box="[796,927,1653,1679]" country="Portugal" httpUri="http://grbio.org/cool/swdk-a9xe" name="Museu Nacional de Historia Natural e de CIência" pageId="12" pageNumber="451" type="Museum">MUHNAC</collectionCode>
/
<specimenCode id="DB8DCB7CD13DFFAFB3D224213A65F908" box="[935,1068,1653,1679]" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="12" pageNumber="451">MB03-1707</specimenCode>
) postpygal tail segments.
</paragraph>
</subSubSection>
<subSubSection id="C331308CD13DFFAFB0B224CC3F74F898" pageId="12" pageNumber="451" type="distribution">
<paragraph id="8B946307D13DFFAFB0B224CC3F74F898" blockId="12.[151,1437,1220,2039]" pageId="12" pageNumber="451">
<emphasis id="B95FBF15D13DFFAFB0B224CC3F28F935" bold="true" box="[199,353,1688,1714]" pageId="12" pageNumber="451">Distribution.</emphasis>
<taxonomicName id="4C2B1884D13DFFAFB11324CC3CCCF935" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[358,645,1688,1714]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="12" pageNumber="451" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13DFFAFB11324CC3CCCF935" box="[358,645,1688,1714]" italics="true" pageId="12" pageNumber="451">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13DFFAFB2FE24CD3CAAF934" bold="true" box="[651,739,1689,1715]" pageId="12" pageNumber="451">
<taxonomicNameLabel id="A26C026ED13DFFAFB2FE24CD3CAAF934" box="[651,739,1689,1715]" pageId="12" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the
<typeStatus id="5490DDA5D13DFFAFB38524CD3A69F934" box="[1008,1056,1689,1715]" pageId="12" pageNumber="451">type</typeStatus>
locality near Catchi village, in the highlands of the Serra da Neve inselberg,
<collectingRegion id="49EFADE5D13DFFAFB20724E83D70F951" box="[626,825,1724,1750]" country="Angola" name="Namibe" pageId="12" pageNumber="451">Namibe Province</collectingRegion>
,
<collectingCountry id="F33C2397D13DFFAFB33124E83DDFF951" box="[836,918,1724,1750]" name="Angola" pageId="12" pageNumber="451">Angola</collectingCountry>
(
<figureCitation id="13107F82D13DFFAFB3D324E83DA2F951" box="[934,1003,1724,1750]" captionStart="FIGURE 6" captionStartId="13.[152,255,1596,1620]" captionTargetBox="[164,1428,194,1559]" captionTargetId="figure-20@13.[151,1436,181,1572]" captionTargetPageId="13" captionText="FIGURE 6. Distribution of Rhoptropus nivimontanus sp. nov. and the boultoni “group” inAngola. Stars represent the respective type localities." figureDoi="http://doi.org/10.5281/zenodo.14734260" httpUri="https://zenodo.org/record/14734260/files/figure.png" pageId="12" pageNumber="451">Fig. 6</figureCitation>
). Considering the substrate specificity of the genus and the ecological isolation of the inselberg, it is assumed to be endemic to higher elevation areas of Serra da Neve.
</paragraph>
</subSubSection>
<subSubSection id="C331308CD13DFFAFB0B2257C3BD0F871" pageId="12" pageNumber="451" type="biology_ecology">
<paragraph id="8B946307D13DFFAFB0B2257C3BD0F871" blockId="12.[151,1437,1220,2039]" pageId="12" pageNumber="451">
<emphasis id="B95FBF15D13DFFAFB0B2257C3C1CF8C4" bold="true" box="[199,597,1832,1859]" pageId="12" pageNumber="451">Habitat and natural history notes.</emphasis>
Very little is known about the ecology and natural history of
<taxonomicName id="4C2B1884D13DFFAFB49C257D3BD5F8C5" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[1257,1436,1833,1858]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="12" pageNumber="451" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13DFFAFB49C257D3BD5F8C5" box="[1257,1436,1833,1858]" italics="true" pageId="12" pageNumber="451">R. nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13DFFAFB0E225193EA7F8E0" bold="true" box="[151,238,1869,1895]" pageId="12" pageNumber="451">
<taxonomicNameLabel id="A26C026ED13DFFAFB0E225193EA7F8E0" box="[151,238,1869,1895]" pageId="12" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The
<typeStatus id="5490DDA5D13DFFAFB15F25193F13F8E0" box="[298,346,1869,1895]" pageId="12" pageNumber="451">type</typeStatus>
series was collected at roughly
<quantity id="4CD3CEE2D13DFFAFB2C725183D4CF8E0" box="[690,773,1868,1895]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="451" unit="m" value="1600.0">1600 m</quantity>
above sea level in an area dominated by Miombo woodlands on the highlands of the inselberg (
<bibRefCitation id="EFBA1EF6D13DFFAFB27C25243CEFF80C" author="Huntley, B. J." box="[521,678,1904,1931]" pageId="12" pageNumber="451" refId="ref10881" refString="Huntley, B. J. (2023) Ecology of Angola: Terrestrial Biomes and Ecoregions. Springer, Cham, 460 pp. https: // doi. org / 10.1007 / 978 - 3 - 031 - 18923 - 4" type="book" year="2023">Huntley 2023</bibRefCitation>
;
<bibRefCitation id="EFBA1EF6D13DFFAFB2C525253DD5F80C" author="Marques, M. P. &amp; Parrinha, D. &amp; Lopes-Lima, M. &amp; Tiutenko, A. &amp; Bauer, A. M. &amp; Ceriaco, L. M. P." box="[688,924,1904,1931]" pageId="12" pageNumber="451" pagination="167 - 217" refId="ref11774" refString="Marques, M. P., Parrinha, D., Lopes-Lima, M., Tiutenko, A., Bauer, A. M. &amp; Ceriaco, L. M. P. (2024 a) An Island in a sea of sand: a first checklist of the herpetofauna of the Serra da Neve inselberg, southwestern Angola. ZooKeys, 1201, 167 - 217. https: // doi. org / 10.3897 / zookeys. 1201.120750" type="journal article" year="2024">
Marques
<emphasis id="B95FBF15D13DFFAFB36225253D04F80D" box="[791,845,1904,1930]" italics="true" pageId="12" pageNumber="451">et al.</emphasis>
2024a
</bibRefCitation>
). Specimens were found in a riparian area with denser vegetation and barely any granite outcrops (
<figureCitation id="13107F82D13DFFAFB2AA25C03D6CF828" box="[735,805,1940,1967]" captionStart="FIGURE 7" captionStartId="14.[152,255,1061,1085]" captionTargetBox="[151,1435,181,1037]" captionTargetId="figure-25@14.[151,1435,181,1037]" captionTargetPageId="14" captionText="FIGURE 7. Habitat of Rhoptropus nivimontanus sp. nov. at the type locality near Catchi village, on the highlands of Serra da Neve, Namibe Province. Photo by LMPC." figureDoi="http://doi.org/10.5281/zenodo.14734262" httpUri="https://zenodo.org/record/14734262/files/figure.png" pageId="12" pageNumber="451">Fig. 7</figureCitation>
), in contrast to surrounding woodlands in the inselberg occupied by congener
<taxonomicName id="4C2B1884D13DFFAFB1E825ED3C1BF855" box="[413,594,1976,2002]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" isUncertain="true" kingdom="Animalia" pageId="12" pageNumber="451" phylum="Chordata" rank="species" species="barnardi">
<emphasis id="B95FBF15D13DFFAFB1E825ED3FFFF855" box="[413,438,1977,2002]" italics="true" pageId="12" pageNumber="451">R.</emphasis>
aff.
<emphasis id="B95FBF15D13DFFAFB19A25EC3C1BF855" box="[495,594,1976,2002]" italics="true" pageId="12" pageNumber="451">barnardi</emphasis>
</taxonomicName>
. It was observed together with
<taxonomicName id="4C2B1884D13DFFAFB3B025EC3C2AF870" authority="Marques, Ceriaco, Buehler, Bandeira, Janota &amp; Bauer, 2020" authorityName="Marques, Ceriaco, Buehler, Bandeira, Janota &amp; Bauer" authorityYear="2020" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="12" pageNumber="451" phylum="Chordata" rank="species" species="nyaneka">
<emphasis id="B95FBF15D13DFFAFB3B025EC3A88F855" box="[965,1217,1976,2002]" italics="true" pageId="12" pageNumber="451">Lygodactylus nyaneka</emphasis>
<bibRefCitation id="EFBA1EF6D13DFFAFB4BE25ED3C2AF870" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Buehler, M. D. &amp; Bandeira, S. A. &amp; Janota, J. M. &amp; Bauer AM" pageId="12" pageNumber="451" pagination="301 - 352" refId="ref11589" refString="Marques, M. P., Ceriaco, L. M. P., Buehler, M. D., Bandeira, S. A., Janota, J. M. &amp; Bauer AM (2020) A revision of the Dwarf Geckos, genus Lygodactylus (Squamata: Gekkonidae), from Angola, with the description of three new species. Zootaxa, 4853 (3), 301 - 352. https: // doi. org / 10.11646 / zootaxa. 4853.3.1" type="journal article" year="2020">Marques, Ceríaco, Buehler, Bandeira, Janota &amp; Bauer, 2020</bibRefCitation>
</taxonomicName>
on tree branches, suggesting more arboreal rather than rupicolous habits.
</paragraph>
</subSubSection>
<caption id="DF54338FD13CFFAEB0ED24683F65F9FF" ID-DOI="http://doi.org/10.5281/zenodo.14734260" ID-Zenodo-Dep="14734260" httpUri="https://zenodo.org/record/14734260/files/figure.png" pageId="13" pageNumber="452" startId="13.[152,255,1596,1620]" targetBox="[164,1428,194,1559]" targetPageId="13" targetType="figure">
<paragraph id="8B946307D13CFFAEB0ED24683F65F9FF" blockId="13.[151,1437,1596,1656]" pageId="13" pageNumber="452">
<emphasis id="B95FBF15D13CFFAEB0ED24683F5EF9D3" bold="true" box="[152,279,1596,1620]" pageId="13" pageNumber="452">FIGURE 6.</emphasis>
Distribution of
<taxonomicName id="4C2B1884D13CFFAEB1C124683CFEF9D3" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[436,695,1596,1620]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="13" pageNumber="452" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13CFFAEB1C124683CFEF9D3" box="[436,695,1596,1620]" italics="true" pageId="13" pageNumber="452">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13CFFAEB2C924683D42F9D3" bold="true" box="[700,779,1596,1620]" pageId="13" pageNumber="452">
<taxonomicNameLabel id="A26C026ED13CFFAEB2C924683D42F9D3" box="[700,779,1596,1620]" pageId="13" pageNumber="452" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and the
<taxonomicName id="4C2B1884D13CFFAEB32824683DF8F9D3" box="[861,945,1596,1620]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="13" pageNumber="452" phylum="Chordata" rank="species" species="boultoni">
<emphasis id="B95FBF15D13CFFAEB32824683DF8F9D3" box="[861,945,1596,1620]" italics="true" pageId="13" pageNumber="452">boultoni</emphasis>
</taxonomicName>
“group” in Angola. Stars represent the respective type localities.
</paragraph>
</caption>
<subSubSection id="C331308CD13CFFAEB0B2249E3A2AF8F3" pageId="13" pageNumber="452" type="etymology">
<paragraph id="8B946307D13CFFAEB0B2249E3A2AF8F3" blockId="13.[151,1437,1738,1909]" pageId="13" pageNumber="452">
<emphasis id="B95FBF15D13CFFAEB0B2249E3F07F963" bold="true" box="[199,334,1738,1764]" pageId="13" pageNumber="452">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2B1884D13CFFAEB23C249E3CA5F963" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[585,748,1738,1764]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="13" pageNumber="452" phylum="Chordata" rank="species" species="nivimontanus">
“
<emphasis id="B95FBF15D13CFFAEB220249F3CA5F963" box="[597,748,1739,1764]" italics="true" pageId="13" pageNumber="452">nivimontanus</emphasis>
</taxonomicName>
” is formed by the combination of the Latin words “
<emphasis id="B95FBF15D13CFFAEB52E249F3BD9F963" box="[1371,1424,1739,1764]" italics="true" pageId="13" pageNumber="452">nivis</emphasis>
” (genitive singular of
<emphasis id="B95FBF15D13CFFAEB1F724BB3FEDF88F" box="[386,420,1775,1800]" italics="true" pageId="13" pageNumber="452">nix</emphasis>
, i.e., snow) and
<taxonomicName id="4C2B1884D13CFFAEB22A24BA3C90F88F" box="[607,729,1774,1800]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="13" pageNumber="452" phylum="Chordata" rank="species" species="montanus">
“
<emphasis id="B95FBF15D13CFFAEB21924BB3C90F88F" box="[620,729,1775,1800]" italics="true" pageId="13" pageNumber="452">montanus</emphasis>
</taxonomicName>
” (nominative adjectival of
<emphasis id="B95FBF15D13CFFAEB47B24BB3A00F88F" box="[1038,1097,1775,1800]" italics="true" pageId="13" pageNumber="452">mons</emphasis>
, i.e., pertaining to mountains) (
<bibRefCitation id="EFBA1EF6D13CFFAEB0EA25463F78F8AA" author="Brown, R. W." box="[159,305,1810,1837]" pageId="13" pageNumber="452" refId="ref9813" refString="Brown, R. W. (1956) Composition of scientific words. A manual of methods and a lexicon of materials for the practice of logotechnics. Smithsonian Books, Washington D. C., 882 pp." type="book" year="1956">Brown 1956</bibRefCitation>
). It is given in reference to the species’ endemic distribution in the Serra da Neve inselberg, whose Portuguese name directly translates to “Mountain of the Snow”. We suggest the English and Portuguese common names “Serra da Neve Day Gecko” and “Osga Diurna da Serra da Neve”, respectively.
</paragraph>
</subSubSection>
<caption id="DF54338FD13FFFADB0ED26713C0DFBE6" ID-DOI="http://doi.org/10.5281/zenodo.14734262" ID-Zenodo-Dep="14734262" httpUri="https://zenodo.org/record/14734262/files/figure.png" pageId="14" pageNumber="453" startId="14.[152,255,1061,1085]" targetBox="[151,1435,181,1037]" targetPageId="14" targetType="figure">
<paragraph id="8B946307D13FFFADB0ED26713C0DFBE6" blockId="14.[151,1436,1061,1121]" pageId="14" pageNumber="453">
<emphasis id="B95FBF15D13FFFADB0ED26713F5AFBBA" bold="true" box="[152,275,1061,1085]" pageId="14" pageNumber="453">FIGURE 7</emphasis>
. Habitat of
<taxonomicName id="4C2B1884D13FFFADB1F926713CD8FBBA" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[396,657,1061,1085]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="14" pageNumber="453" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13FFFADB1F926713CD8FBBA" box="[396,657,1061,1085]" italics="true" pageId="14" pageNumber="453">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13FFFADB2E226713CA1FBBA" bold="true" box="[663,744,1061,1085]" pageId="14" pageNumber="453">
<taxonomicNameLabel id="A26C026ED13FFFADB2E226713CA1FBBA" box="[663,744,1061,1085]" pageId="14" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at the type locality near Catchi village, on the highlands of Serra da Neve, Namibe Province. Photo by LMPC.
</paragraph>
</caption>
<subSubSection id="C331308CD13FFFADB0B226E73FC9F9DD" pageId="14" pageNumber="453" type="conservation">
<paragraph id="8B946307D13FFFADB0B226E73FC9F9DD" blockId="14.[151,1437,1203,1626]" pageId="14" pageNumber="453">
<emphasis id="B95FBF15D13FFFADB0B226E73F26FB4A" bold="true" box="[199,367,1203,1229]" pageId="14" pageNumber="453">Conservation.</emphasis>
<taxonomicName id="4C2B1884D13FFFADB10626E73CD9FB4A" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[371,656,1203,1229]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="14" pageNumber="453" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13FFFADB10626E73CD9FB4A" box="[371,656,1203,1229]" italics="true" pageId="14" pageNumber="453">Rhoptropus nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13FFFADB2E126E03CA2FB49" bold="true" box="[660,747,1204,1230]" pageId="14" pageNumber="453">
<taxonomicNameLabel id="A26C026ED13FFFADB2E126E03CA2FB49" box="[660,747,1204,1230]" pageId="14" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from
<specimenCount id="9D2DA88ED13FFFADB3BC26E73A33FB4A" box="[969,1146,1203,1229]" count="3" pageId="14" pageNumber="453" type="generic">three specimens</specimenCount>
and is apparently endemic to the Serra da Neve inselberg. Assuming that the species is homogeneously distributed throughout the inselberg, we estimated an extent of occurrence of approximately
<quantity id="4CD3CEE2D13FFFADB35B26AF3DC1FA92" box="[814,904,1275,1301]" metricMagnitude="5" metricUnit="m" metricValue="6.3" pageId="14" pageNumber="453" unit="km" value="630.0">630 km</quantity>
<superScript id="7C5ECE4FD13FFFADB3FD26AF3DD9FA8E" attach="left" box="[904,912,1275,1289]" fontSize="6" pageId="14" pageNumber="453">2</superScript>
using the GeoCAT Geospatial Conservation Assessment Tool (
<bibRefCitation id="EFBA1EF6D13FFFADB112274B3C13FABD" author="Bachman, S. &amp; Moat, J. &amp; Hill, A. W. &amp; Torre, J. &amp; Scott, B." box="[359,602,1311,1338]" pageId="14" pageNumber="453" pagination="117 - 126" refId="ref9123" refString="Bachman, S., Moat, J., Hill, A. W., Torre, J. &amp; Scott, B. (2011) Supporting Red List threat assessment withGeoCAT: Geospatial conservation assessment tool. ZooKeys, 150, 117 - 126. https: // doi. org / 10.3897 / zookeys. 150.2109" type="journal article" year="2011">
Bachman
<emphasis id="B95FBF15D13FFFADB1AE27743C47FABE" box="[475,526,1311,1337]" italics="true" pageId="14" pageNumber="453">et al</emphasis>
. 2011
</bibRefCitation>
). Despite its remoteness, the landscape near human settlements is visibly transformed, with large deforested areas dedicated to agriculture or cattle pastures (
<figureCitation id="13107F82D13FFFADB43C27173AC7FAD9" box="[1097,1166,1347,1374]" captionStart="FIGURE 8" captionStartId="15.[152,255,1123,1147]" captionTargetBox="[151,1436,181,1099]" captionTargetId="figure-20@15.[151,1436,181,1099]" captionTargetPageId="15" captionText="FIGURE 8. General view of Catchi village and plateau, with riparian gallery and type locality of Rhoptropus nivimontanus sp. nov. in foreground (red arrow). Photo by LMPC." figureDoi="http://doi.org/10.5281/zenodo.14734266" httpUri="https://zenodo.org/record/14734266/files/figure.png" pageId="14" pageNumber="453">Fig. 8</figureCitation>
;
<bibRefCitation id="EFBA1EF6D13FFFADB4E927103BD8FAD9" author="Marques, M. P. &amp; Parrinha, D. &amp; Lopes-Lima, M. &amp; Tiutenko, A. &amp; Bauer, A. M. &amp; Ceriaco, L. M. P." box="[1180,1425,1347,1374]" pageId="14" pageNumber="453" pagination="167 - 217" refId="ref11774" refString="Marques, M. P., Parrinha, D., Lopes-Lima, M., Tiutenko, A., Bauer, A. M. &amp; Ceriaco, L. M. P. (2024 a) An Island in a sea of sand: a first checklist of the herpetofauna of the Serra da Neve inselberg, southwestern Angola. ZooKeys, 1201, 167 - 217. https: // doi. org / 10.3897 / zookeys. 1201.120750" type="journal article" year="2024">
Marques
<emphasis id="B95FBF15D13FFFADB57327103B09FADA" box="[1286,1344,1347,1373]" italics="true" pageId="14" pageNumber="453">et al.</emphasis>
2024a
</bibRefCitation>
). Considering the apparent association of
<taxonomicName id="4C2B1884D13FFFADB22C273C3D46FA06" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[601,783,1384,1409]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="14" pageNumber="453" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13FFFADB22C273C3D46FA06" box="[601,783,1384,1409]" italics="true" pageId="14" pageNumber="453">R. nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13FFFADB363273C3D26FA05" bold="true" box="[790,879,1384,1410]" pageId="14" pageNumber="453">
<taxonomicNameLabel id="A26C026ED13FFFADB363273C3D26FA05" box="[790,879,1384,1410]" pageId="14" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with arboreal habitats and riparian areas, it might become particularly vulnerable to deforestation, an alarming cause of habitat loss in the highlands of the Angolan Central Plateau (
<bibRefCitation id="EFBA1EF6D13FFFADB12427FB3C6BFA4D" author="Mendelsohn, J. M." box="[337,546,1455,1482]" pageId="14" pageNumber="453" pagination="123 - 140" refId="ref11955" refString="Mendelsohn, J. M. (2019) Landscape changes in Angola. In: Huntley, B. J., Russo, V., Lages, F. &amp; Ferrand, N. (Eds.), Biodiversity of Angola: Science &amp; Conservation, a modern synthesis. Springer, Cham, pp. 123 - 140. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 8" type="book chapter" year="2019">Mendelsohn 2019</bibRefCitation>
). Currently available data does not allow for an unambiguous assessment of the conservation status of
<taxonomicName id="4C2B1884D13FFFADB1E627803C03FA6A" authorityName="Parrinha &amp; Marques &amp; Tiutenko &amp; Heinicke &amp; Bauer &amp; Ceríaco" authorityYear="2025" box="[403,586,1492,1517]" class="Squamata" family="Gekkonidae" genus="Rhoptropus" kingdom="Animalia" pageId="14" pageNumber="453" phylum="Chordata" rank="species" species="nivimontanus" status="sp. nov.">
<emphasis id="B95FBF15D13FFFADB1E627803C03FA6A" box="[403,586,1492,1517]" italics="true" pageId="14" pageNumber="453">R. nivimontanus</emphasis>
</taxonomicName>
<emphasis id="B95FBF15D13FFFADB22427803CE2FA69" bold="true" box="[593,683,1492,1518]" pageId="14" pageNumber="453">
<taxonomicNameLabel id="A26C026ED13FFFADB22427803CE2FA69" box="[593,683,1492,1518]" pageId="14" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
While we recommend the status of Data Deficient for this species, further surveys targeted at Serra da Neve and other inselbergs in the region, and a formal conservation assessment are warranted, especially considering the susceptibility of montane species to climate change (
<bibRefCitation id="EFBA1EF6D13FFFADB4C9244F3BD1F9B1" author="Sinervo, B. &amp; Mendez-de-la-Cruz, F. &amp; Miles, D. B. &amp; Heulin, B. &amp; Bastiaans, E. &amp; Villagran-Santa Cruz, M. &amp; Lara-Resendiz, R. &amp; Martinez-Mendez, N. &amp; Calderon-Espinosa, M. L. &amp; Meza-Lazaro, R. N. &amp; Gadsden, H. &amp; Avila, L. J. &amp; Morando, M. &amp; De la Riva, I. J. &amp; Sepulveda, P. V. &amp; Rocha, C. F. D. &amp; Ibarguengoytia, N. &amp; Puntriano, C. A. &amp; Massot, M. &amp; Lepetz, V. &amp; Oksanen, T. A. &amp; Chapple, D. G. &amp; Bauer, A. M. &amp; Branch, W. R. &amp; Clobert, J. &amp; Sites Jr., J. W." box="[1212,1432,1563,1590]" pageId="14" pageNumber="453" pagination="894 - 899" refId="ref12460" refString="Sinervo, B., Mendez-de-la-Cruz, F., Miles, D. B., Heulin, B., Bastiaans, E., Villagran-Santa Cruz, M., Lara-Resendiz, R., Martinez-Mendez, N., Calderon-Espinosa, M. L., Meza-Lazaro, R. N., Gadsden, H., Avila, L. J., Morando, M., De la Riva, I. J., Sepulveda, P. V., Rocha, C. F. D., Ibarguengoytia, N., Puntriano, C. A., Massot, M., Lepetz, V., Oksanen, T. A., Chapple, D. G., Bauer, A. M., Branch, W. R., Clobert, J. &amp; Sites Jr., J. W. (2010) Erosion of lizard diversity by climate change and altered thermal niches. Science, 328 (5980), 894 - 899. https: // doi. org / 10.1126 / science. 1184695" type="journal article" year="2010">
Sinervo
<emphasis id="B95FBF15D13FFFADB56924483B1CF9B2" box="[1308,1365,1563,1589]" italics="true" pageId="14" pageNumber="453">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EFBA1EF6D13FFFADB0E2246B3F3DF9DD" author="Moreira, M. O. &amp; Fonseca, C. &amp; Rojas, D." box="[151,372,1599,1626]" pageId="14" pageNumber="453" pagination="676 - 689" refId="ref12199" refString="Moreira, M. O., Fonseca, C. &amp; Rojas, D. (2023) Potential persistence of high-mountain lizards. Ecological Research, 38 (5), 676 - 689. https: // doi. org / 10.1111 / 1440 - 1703.12410" type="journal article" year="2023">
Moreira
<emphasis id="B95FBF15D13FFFADB08D24143F78F9DE" box="[248,305,1599,1625]" italics="true" pageId="14" pageNumber="453">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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