222 lines
26 KiB
XML
222 lines
26 KiB
XML
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<mods:title id="4A829628333429F8F95B9A80BA0CB617">A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)</mods:title>
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<mods:namePart id="3C77CBE94214915ADFFDD5DC55ED66D9">Torres-Carvajal, Omar</mods:namePart>
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<mods:namePart id="2C213D6DC2C7FBC51A26AC3077409084">Etheridge, Richard</mods:namePart>
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<mods:namePart id="9BEABFEA8D40812D39E24A9130AF9C74">Queiroz, Kevin De</mods:namePart>
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<treatment id="038287FDFFA5605969838AFAD0E0F84E" ID-DOI="http://doi.org/10.5281/zenodo.6184975" ID-GBIF-Taxon="119387049" ID-Zenodo-Dep="6184975" LSID="urn:lsid:plazi:treatment:038287FDFFA5605969838AFAD0E0F84E" httpUri="http://treatment.plazi.org/id/038287FDFFA5605969838AFAD0E0F84E" lastPageNumber="31" pageId="30" pageNumber="31">
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<subSubSection id="C3316560FFA5605969838AFAD3B3FE63" pageId="30" pageNumber="31" type="nomenclature">
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<paragraph id="8B9436EBFFA5605969838AFAD3A4FE3E" blockId="30.[151,720,403,430]" box="[151,720,403,430]" pageId="30" pageNumber="31">
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<heading id="D0DC8187FFA5605969838AFAD3A4FE3E" bold="true" box="[151,720,403,430]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
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<taxonomicName id="4C2B4D68FFA5605969838AFAD3A4FE3E" ID-CoL="6FGTM" authority="Torres-Carvajal et al. 2008" authorityName="Torres-Carvajal et al." authorityYear="2008" box="[151,720,403,430]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
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<emphasis id="B95FEAF9FFA5605969838AFAD3A4FE3E" bold="true" box="[151,720,403,430]" pageId="30" pageNumber="31">
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<emphasis id="B95FEAF9FFA5605969838AFAD008FE3D" bold="true" box="[151,380,404,430]" italics="true" pageId="30" pageNumber="31">Enyalioides touzeti</emphasis>
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<bibRefCitation id="EFBA4B1AFFA5605968978AFDD3A4FE3E" author="Torres-Carvajal" box="[387,720,403,430]" pageId="30" pageNumber="41" refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235." type="journal article" year="2008">
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Torres-Carvajal
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<emphasis id="B95FEAF9FFA560596B408AFAD3F3FE3D" bold="true" box="[596,647,404,430]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
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. 2008
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</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9436EBFFA5605969838AB4D3B3FE63" blockId="30.[151,777,474,527]" box="[151,711,474,496]" pageId="30" pageNumber="31">Proposed standard English name: Touzet’s woodlizards</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FFA5605969838A97D27DFD9C" box="[151,777,505,527]" pageId="30" pageNumber="31" type="description">
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<paragraph id="8B9436EBFFA5605969838A97D27DFD9C" blockId="30.[151,777,474,527]" box="[151,777,505,527]" pageId="30" pageNumber="31">Proposed standard Spanish name: lagartijas de palo de Touzet</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FFA5605969838941D292FDF7" pageId="30" pageNumber="31" type="reference_group">
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<paragraph id="8B9436EBFFA5605969838941D292FDF7" blockId="30.[151,1436,559,613]" pageId="30" pageNumber="31">
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<treatmentCitationGroup id="AB3B11C5FFA5605969838941D292FDF7" infered="true" pageId="30" pageNumber="31">
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<taxonomicName id="4C2B4D68FFA5605969838941D3C9FDD5" ID-CoL="6FGTM" authority="Torres-Carvajal et al. (2008:228)" authorityName="Torres-Carvajal et al." authorityPageNumber="228" authorityYear="2008" box="[151,701,559,582]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
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<emphasis id="B95FEAF9FFA5605969838941D02FFDD5" box="[151,347,559,582]" italics="true" pageId="30" pageNumber="31">Enyalioides touzeti</emphasis>
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<treatmentCitation id="0A8A10FAFFA5605968708941D3C2FDD5" author="Torres-Carvajal" box="[356,694,559,582]" httpUri="http://treatment.plazi.org/id/F82287FDFFB6FB5DFCD5F9F221CAB631" page="228" pageId="30" pageNumber="31" year="2008">
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<bibRefCitation id="EFBA4B1AFFA5605968708941D3FDFDD5" author="Torres-Carvajal" box="[356,649,559,582]" pageId="30" pageNumber="41" refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235." type="journal article" year="2008">
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Torres-Carvajal
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<emphasis id="B95FEAF9FFA560596B1F895FD335FDD6" box="[523,577,559,582]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
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(2008
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</bibRefCitation>
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:228
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</treatmentCitation>
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)
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</taxonomicName>
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.
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<typeStatus id="54908849FFA560596BDE8941D258FDD6" box="[714,812,559,581]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
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: EPN 10306, from “Finca La Envidia (3º3'0'S,
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<geoCoordinate id="EE1F502CFFA560596C0A8941D4ECFDD6" box="[1310,1432,559,581]" direction="west" orientation="longitude" pageId="30" pageNumber="31" precision="15" value="-79.69028">79º41'25''W</geoCoordinate>
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,
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<quantity id="4CD39B0EFFA5605969D38921D076FDF6" box="[199,258,591,613]" metricMagnitude="2" metricUnit="m" metricValue="4.33" pageId="30" pageNumber="31" unit="m" value="433.0">433m</quantity>
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) cerro Santa
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<collectingCountry id="F33C767BFFA56059689C8921D0B3FDF7" box="[392,455,591,612]" name="Malta" pageId="30" pageNumber="31">Marta</collectingCountry>
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, Cantón Ponce Enríquez, Provincia Azuay,
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<collectingCountry id="F33C767BFFA560596A958920D2ADFDF7" box="[897,985,590,612]" name="Ecuador" pageId="30" pageNumber="31">Ecuador</collectingCountry>
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.”
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</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FFA56059698389FED564FC86" pageId="30" pageNumber="31" type="diagnosis">
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<paragraph id="8B9436EBFFA56059698389FED564FC86" blockId="30.[151,1437,656,2013]" pageId="30" pageNumber="31">
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<emphasis id="B95FEAF9FFA56059698389FED065FD3A" bold="true" box="[151,273,656,681]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
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<taxonomicName id="4C2B4D68FFA56059680F89FFD081FD3A" box="[283,501,657,681]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
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<emphasis id="B95FEAF9FFA56059680F89FFD081FD3A" box="[283,501,657,681]" italics="true" pageId="30" pageNumber="31">Enyalioides touzeti</emphasis>
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</taxonomicName>
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can be distinguished from other species of
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<taxonomicName id="4C2B4D68FFA560596AE389FFD50FFD3A" box="[1015,1147,657,681]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
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<emphasis id="B95FEAF9FFA560596AE389FFD50FFD3A" box="[1015,1147,657,681]" italics="true" pageId="30" pageNumber="31">Enyalioides</emphasis>
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</taxonomicName>
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by having the following combination of characters: ventrals keeled; dorsals keeled; flank scales homogeneous in size; vertebral scales in neck region in adult males more than twice as high as vertebrals in pelvic region; gular region in males extensively covered by a dark gular patch; and dorsum and limbs without projecting scales.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3316560FFA5605969D1884ED0AAF9C2" pageId="30" pageNumber="31" type="description">
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<paragraph id="8B9436EBFFA5605969D1884ED085FA9E" blockId="30.[151,1437,656,2013]" pageId="30" pageNumber="31">
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<emphasis id="B95FEAF9FFA5605969D1884ED023FCAA" bold="true" box="[197,343,800,825]" pageId="30" pageNumber="31">Description.</emphasis>
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(1) dorsal head scales conical or multicarinate, strongly projecting dorsally; (2) posterior superciliaries not enlarged relative to adjacent scales; (3) scales on lateral edge of skull roof just posterior to superciliaries more projecting than adjacent scales; the projection is more pronounced in adults; (4) one or two enlarged pretympanic scales present; (5) gular scales conical or multicarinate, strongly projecting ventrally; (6) dorsal neck scales homogeneous in size, mostly conical or cycloid and slightly imbricate; lateral neck scales similar in size to dorsal neck scales, sometimes with a few enlarged conical scales; (7) vertebrals larger than adjacent dorsals, forming distinct raised middorsal crest that extends onto tail as a pair of crests; (8) nuchal region with continuous and single middorsal crest; (9) dorsals small, prominently keeled, imbricate, and homogeneous in size; (10) longitudinal row of raised, enlarged scales between dorsals and flank scales present and continuous; (11) scales on flanks homogeneous in size, smaller than dorsals; (12) ventrals keeled; (13) fore limb scales keeled dorsally and ventrally; (14) hind limb scales keeled dorsally and ventrally; scattered enlarged scales absent; dorsal scales of pes homogeneous in size; (15) caudals heterogeneous, increasing in size posteriorly on each segment (6–8 scales in lateral view), not modified as conspicuous spines (
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<figureCitation id="13102A6EFFA560596B6B8FBFD3B6FB7A" box="[639,706,1233,1257]" captionStart="FIGURE 3" captionStartId="12.[151,250,1517,1540]" captionTargetBox="[280,1305,321,1491]" captionTargetId="figure@12.[274,1314,306,1505]" captionTargetPageId="12" captionText="FIGURE 3. Caudal segments from the anterior third of the tail of ten species of Hoplocercinae in lateral view. (A) Enyalioides cofanorum, (B) E. heterolepis, (C) E. laticeps, (D) E. microlepis, (E) E. oshaughnessyi, (F) E. palpebralis, (G) E. praestabilis, (H) E. rubrigularis, (I) E. touzeti, and (J) Morunasaurus annularis. Scale bars = 5 mm. Illustration by D. Paucar." httpUri="https://zenodo.org/record/207076/files/figure.png" pageId="30" pageNumber="31">Fig. 3</figureCitation>
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); (16) tail compressed laterally. Meristic and morphometric characters are presented in
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<tableCitation id="C6A90350FFA56059688C8F9BD09FFA9E" box="[408,491,1269,1293]" captionStart="TABLE 1" captionStartId="6.[151,235,1304,1325]" captionTargetBox="[151,1436,1377,1892]" captionTargetPageId="6" captionText="TABLE 1. Summary of counts and measurements (mm) for species of Hoplocercinae. Range (first line) and mean ± standard" httpUri="http://table.plazi.org/id/DF546663FFBD604169838E76D4E8FABD" pageId="30" pageNumber="31" tableUuid="DF546663FFBD604169838E76D4E8FABD">Table 1</tableCitation>
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.
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</paragraph>
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<paragraph id="8B9436EBFFA5605969D18E76D359FA76" blockId="30.[151,1437,656,2013]" pageId="30" pageNumber="31">
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<emphasis id="B95FEAF9FFA5605969D18E76D0E1FAA2" bold="true" box="[197,405,1304,1329]" pageId="30" pageNumber="31">Coloration in life</emphasis>
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(
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<figureCitation id="13102A6EFFA5605968BC8E77D09BFAA2" box="[424,495,1305,1329]" captionStart="FIGURE 4" captionStartId="14.[151,250,1897,1920]" captionTargetBox="[204,1383,190,1874]" captionTargetId="figure@14.[204,1384,184,1876]" captionTargetPageId="14" captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae." httpUri="https://zenodo.org/record/207077/files/figure.png" pageId="30" pageNumber="31">Fig. 4</figureCitation>
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). Dorsal background olive green or light brown in adults (reddish brown in juveniles), normally with yellow spots in males and large dark blotches (some with background color in the center) in females; supralabials and infralabials normally yellow; light stripe extending from mouth commisure to tympanum in some specimens; light stripe between tympanum and scapular region in some specimens; chin and gular regions in males dark; venter light brown or whitish cream, with dark irregular marks in some specimens; iris reddish brown (
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<bibRefCitation id="EFBA4B1AFFA5605969F98EA3D355FA76" author="Torres-Carvajal" box="[237,545,1485,1509]" pageId="30" pageNumber="41" refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235." type="journal article" year="2008">
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||
Torres-Carvajal
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<emphasis id="B95FEAF9FFA5605968B08EA0D0A9FA76" box="[420,477,1485,1509]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
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2008
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</bibRefCitation>
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).
|
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</paragraph>
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<paragraph id="8B9436EBFFA5605969D18E9FD0AAF9C2" blockId="30.[151,1437,656,2013]" pageId="30" pageNumber="31">
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<emphasis id="B95FEAF9FFA5605969D18E9FD0F7F99A" bold="true" box="[197,387,1520,1545]" pageId="30" pageNumber="31">Natural history.</emphasis>
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Individuals of
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<taxonomicName id="4C2B4D68FFA560596B3A8E9FD270F99A" box="[558,772,1521,1545]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
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||
<emphasis id="B95FEAF9FFA560596B3A8E9FD270F99A" box="[558,772,1521,1545]" italics="true" pageId="30" pageNumber="31">Enyalioides touzeti</emphasis>
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</taxonomicName>
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have been found active on the ground or perching on horizontal branches up to
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<quantity id="4CD39B0EFFA5605968808D7BD0AFF9BE" box="[404,475,1557,1581]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="cm" value="30.0">30 cm</quantity>
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high during the day, or sleeping in a vertical position on tree fern branches at night (
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<bibRefCitation id="EFBA4B1AFFA56059698A8D57D0A6F9C2" author="Torres-Carvajal" box="[158,466,1593,1617]" pageId="30" pageNumber="41" refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235." type="journal article" year="2008">
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||
Torres-Carvajal
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<emphasis id="B95FEAF9FFA5605968428D54D0FBF9C2" box="[342,399,1593,1617]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
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2008
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</bibRefCitation>
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||
).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3316560FFA5605969D18D32D524F92E" pageId="30" pageNumber="31" type="distribution">
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<paragraph id="8B9436EBFFA5605969D18D32D524F92E" blockId="30.[151,1437,656,2013]" pageId="30" pageNumber="31">
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||
<emphasis id="B95FEAF9FFA5605969D18D32D02AF9E6" bold="true" box="[197,350,1628,1653]" pageId="30" pageNumber="31">Distribution.</emphasis>
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<taxonomicName id="4C2B4D68FFA5605968718D33D34FF9E6" box="[357,571,1629,1653]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
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<emphasis id="B95FEAF9FFA5605968718D33D34FF9E6" box="[357,571,1629,1653]" italics="true" pageId="30" pageNumber="31">Enyalioides touzeti</emphasis>
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</taxonomicName>
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occurs on the western slopes of the Andes in southern
|
||
<collectingCountry id="F33C767BFFA560596DBE8D33D473F9E6" box="[1194,1287,1629,1653]" name="Ecuador" pageId="30" pageNumber="31">Ecuador</collectingCountry>
|
||
at elevations of
|
||
<quantity id="4CD39B0EFFA5605969A18DEFD040F90A" box="[181,308,1665,1689]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="30" pageNumber="31" unit="m" value="500.0" valueMax="700.0" valueMin="300.0">300–700 m</quantity>
|
||
(
|
||
<figureCitation id="13102A6EFFA5605968508DEFD0E3F90A" box="[324,407,1665,1689]" captionStart="FIGURE 10" captionStartId="28.[151,250,1666,1689]" captionTargetBox="[161,1426,207,1630]" captionTargetId="figure@28.[151,1436,193,1645]" captionTargetPageId="28" captionText="FIGURE 10. Distribution of Enyalioides praestabilis (dots) and E. touzeti (triangles)." httpUri="https://zenodo.org/record/207087/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
|
||
). This species is not known to occur in sympatry with other hoplocercines, although
|
||
<taxonomicName id="4C2B4D68FFA560596C458DECD184F92E" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="heterolepis">
|
||
<emphasis id="B95FEAF9FFA560596C458DECD184F92E" italics="true" pageId="30" pageNumber="31">E. heterolepis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D68FFA5605968328DC8D09CF92E" box="[294,488,1701,1725]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="oshaughnessyi">
|
||
<emphasis id="B95FEAF9FFA5605968328DC8D09CF92E" box="[294,488,1701,1725]" italics="true" pageId="30" pageNumber="31">E. oshaughnessyi</emphasis>
|
||
</taxonomicName>
|
||
occur at nearby localities (
|
||
<figureCitation id="13102A6EFFA560596A0C8DCBD211F92E" box="[792,869,1701,1725]" captionStart="FIGURE 6" captionStartId="20.[151,250,1959,1982]" captionTargetBox="[159,1429,530,1933]" captionTargetId="figure@20.[151,1436,517,1938]" captionTargetPageId="20" captionText="FIGURE 6. Distribution of Enyalioides heterolepis (triangles) and E. laticeps (dots)." httpUri="https://zenodo.org/record/207083/files/figure.png" pageId="30" pageNumber="31">Figs. 6</figureCitation>
|
||
and
|
||
<figureCitation id="13102A6EFFA560596A888DC8D2D8F92E" box="[924,940,1702,1725]" captionStart="FIGURE 7" captionStartId="23.[151,250,1564,1587]" captionTargetBox="[197,1383,204,1538]" captionTargetId="figure@23.[189,1403,169,1564]" captionTargetPageId="23" captionText="FIGURE 7. Distribution of Enyalioides microlepis (dots) and E. oshaughnessyi (triangles). A, Alto Tambo; B, Bilsa Ecological Reserve (see Remarks under E. oshaughnessyi)." httpUri="https://zenodo.org/record/207084/files/figure.png" pageId="30" pageNumber="31">7</figureCitation>
|
||
, respectively).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316560FFA5605969D18DA7D0E0F84E" pageId="30" pageNumber="31" type="discussion">
|
||
<paragraph id="8B9436EBFFA5605969D18DA7D0E0F84E" blockId="30.[151,1437,656,2013]" pageId="30" pageNumber="31">
|
||
<emphasis id="B95FEAF9FFA5605969D18DA7D04DF972" bold="true" box="[197,313,1737,1761]" pageId="30" pageNumber="31">Remarks.</emphasis>
|
||
Torres-Carvajal & de Queiroz (2009) found that the mtDNA maximum-likelihood corrected genetic distance beween this species and
|
||
<taxonomicName id="4C2B4D68FFA560596B0D8D83D23EF896" box="[537,842,1773,1797]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="oshaughnessyi">
|
||
<emphasis id="B95FEAF9FFA560596B0D8D83D23EF896" box="[537,842,1773,1797]" italics="true" pageId="30" pageNumber="31">Enyalioides oshaughnessyi</emphasis>
|
||
</taxonomicName>
|
||
was relatively low (0.03–0.047) and that
|
||
<taxonomicName id="4C2B4D68FFA560596C3B8D80D4E8F896" box="[1327,1436,1774,1797]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
|
||
<emphasis id="B95FEAF9FFA560596C3B8D80D4E8F896" box="[1327,1436,1774,1797]" italics="true" pageId="30" pageNumber="31">E. touzeti</emphasis>
|
||
</taxonomicName>
|
||
was nested within
|
||
<taxonomicName id="4C2B4D68FFA5605968788C7CD344F8BA" box="[364,560,1809,1833]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="oshaughnessyi">
|
||
<emphasis id="B95FEAF9FFA5605968788C7CD344F8BA" box="[364,560,1809,1833]" italics="true" pageId="30" pageNumber="31">E. oshaughnessyi</emphasis>
|
||
</taxonomicName>
|
||
in a phylogenetic tree including one specimen of
|
||
<taxonomicName id="4C2B4D68FFA560596D668C7CD5AAF8BA" box="[1138,1246,1810,1833]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
|
||
<emphasis id="B95FEAF9FFA560596D668C7CD5AAF8BA" box="[1138,1246,1810,1833]" italics="true" pageId="30" pageNumber="31">E. touzeti</emphasis>
|
||
</taxonomicName>
|
||
and three specimens of
|
||
<taxonomicName id="4C2B4D68FFA5605969EC8C58D0CFF8DE" box="[248,443,1845,1869]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="oshaughnessyi">
|
||
<emphasis id="B95FEAF9FFA5605969EC8C58D0CFF8DE" box="[248,443,1845,1869]" italics="true" pageId="30" pageNumber="31">E. oshaughnessyi</emphasis>
|
||
</taxonomicName>
|
||
. However, morphological evidence supports
|
||
<taxonomicName id="4C2B4D68FFA560596AAD8C58D550F8DE" box="[953,1060,1846,1869]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
|
||
<emphasis id="B95FEAF9FFA560596AAD8C58D550F8DE" box="[953,1060,1846,1869]" italics="true" pageId="30" pageNumber="31">E. touzeti</emphasis>
|
||
</taxonomicName>
|
||
as a separate species, which suggests that this could be a case of fixed morphological differences evolving prior to the evolution of reciprocal monophyly of mtDNA alleles. Alternatively,
|
||
<taxonomicName id="4C2B4D68FFA560596BB48C10D211F806" box="[672,869,1917,1941]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="oshaughnessyi">
|
||
<emphasis id="B95FEAF9FFA560596BB48C10D211F806" box="[672,869,1917,1941]" italics="true" pageId="30" pageNumber="31">E. oshaughnessyi</emphasis>
|
||
</taxonomicName>
|
||
as currently circumscribed might represent more than one species, in which case
|
||
<taxonomicName id="4C2B4D68FFA560596B188CCCD30DF82A" box="[524,633,1954,1977]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="touzeti">
|
||
<emphasis id="B95FEAF9FFA560596B188CCCD30DF82A" box="[524,633,1954,1977]" italics="true" pageId="30" pageNumber="31">E. touzeti</emphasis>
|
||
</taxonomicName>
|
||
would not be nested within
|
||
<taxonomicName id="4C2B4D68FFA560596AD38CCCD5F8F82A" box="[967,1164,1953,1977]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="oshaughnessyi">
|
||
<emphasis id="B95FEAF9FFA560596AD38CCCD5F8F82A" box="[967,1164,1953,1977]" italics="true" pageId="30" pageNumber="31">E. oshaughnessyi</emphasis>
|
||
</taxonomicName>
|
||
(see Remarks under
|
||
<taxonomicName id="4C2B4D68FFA560596C908CCCD04EF84E" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="oshaughnessyi">
|
||
<emphasis id="B95FEAF9FFA560596C908CCCD04EF84E" italics="true" pageId="30" pageNumber="31">E. oshaughnessyi</emphasis>
|
||
</taxonomicName>
|
||
above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |