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<document id="7442442783FBEE0FE440196FB9D00CD8" ID-CLB-Dataset="23985" ID-DOI="10.5281/zenodo.3673247" ID-GBIF-Dataset="775d8166-02ea-49c5-a526-c7dd6a387744" ID-ISSN="1942-1354" ID-Zenodo-Dep="3673247" ID-ZooBank="E4E34D66-128E-43D7-833C-456BB172908D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027355183" checkinUser="felipe" docAuthor="Otto, Robert L." docDate="2019" docId="038287A56B479E4536B8FB1DA372FDEF" docLanguage="en" docName="InsectaMundi.0702.1-6.pdf.imf" docOrigin="Insecta Mundi 702 (702)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Cretopityobius pankowskiorum Otto 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFBBFFDD6B429E433678FFA0A017FFEE" masterDocTitle="Descriptions of two new elateroid beetles (Coleoptera: Eucnemidae, Elateridae) from Burmese amber" masterLastPageNumber="6" masterPageNumber="1" pageNumber="4" updateTime="1698784538992" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="F9FA82B0190D6047C7D9EC5B1CD3E6D9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5C1724591F9EE0523372B4495E85C654">
<mods:title id="54F5FA0F648D9D51AAF53F3A81F1AF32">Descriptions of two new elateroid beetles (Coleoptera: Eucnemidae, Elateridae) from Burmese amber</mods:title>
</mods:titleInfo>
<mods:name id="FBBF4EA36570A0B8AFBF64ED5B2EC447" type="personal">
<mods:role id="D413F41A58344AA24D3870FF828F9A37">
<mods:roleTerm id="569B2AC2B7D16A816EE62D5D0565E8B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6FF184AA2DDEFBFB0061693C5E16DEA">Otto, Robert L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1CCB3D844C45404B99D33F8B62D5CE03">text</mods:typeOfResource>
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<mods:titleInfo id="8C44718B4D171F07D8338B52FA2A9854">
<mods:title id="4B7EC41786DE00D1D21E4384C17EC7EF">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="00BA4847F7FB1CD3041AF4EFFC310CFD">
<mods:date id="D8F3F7C3622E3FECDA0CBFB60A271C52">2019</mods:date>
<mods:detail id="8F5A8C3DDE9C34B852BF5CFB3A30C12C" type="pubDate">
<mods:number id="60B1887A72CEB4E0207F4399D02C1706">2019-05-31</mods:number>
</mods:detail>
<mods:detail id="4B5D5D91AE22C0515F45E2F49E7270A6" type="volume">
<mods:number id="D51CBD2B2503748A8CD4FFCAB4794699">702</mods:number>
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<mods:detail id="106BB1D122ADE050C4A00B3A1C01CDB4" type="issue">
<mods:number id="04FC66A2954FBA85FE5F606DBD5FA44D">702</mods:number>
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<mods:classification id="5786C3823C91AF76E31FA462E06FF940">journal article</mods:classification>
<mods:identifier id="7A7DB40ED3C4C3F2EE2B6C08968850D3" type="CLB-Dataset">23985</mods:identifier>
<mods:identifier id="BBE557BCE20FA1FD279DE24EB3006928" type="DOI">10.5281/zenodo.3673247</mods:identifier>
<mods:identifier id="173E3BA86BDEEA91DAF096EDE5351DD7" type="GBIF-Dataset">775d8166-02ea-49c5-a526-c7dd6a387744</mods:identifier>
<mods:identifier id="ADCD865B06835E67BF89104FE559D9B7" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="4FA95EBC3A610E31F3DEFA5C04F67011" type="Zenodo-Dep">3673247</mods:identifier>
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</mods:mods>
<treatment id="038287A56B479E4536B8FB1DA372FDEF" ID-DOI="http://doi.org/10.5281/zenodo.3679943" ID-GBIF-Taxon="162161604" ID-Zenodo-Dep="3679943" LSID="urn:lsid:plazi:treatment:038287A56B479E4536B8FB1DA372FDEF" httpUri="http://treatment.plazi.org/id/038287A56B479E4536B8FB1DA372FDEF" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
<subSubSection id="C33165386B479E4636B8FB1DA331FB3B" box="[192,806,1213,1237]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8B9436B36B479E4636B8FB1DA331FB3B" blockId="5.[192,806,1213,1270]" box="[192,806,1213,1237]" pageId="5" pageNumber="4">
<heading id="D0DC81DF6B479E4636B8FB1DA331FB3B" bold="true" box="[192,806,1213,1237]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
<emphasis id="B95FEAA16B479E4636B8FB1DA331FB3B" bold="true" box="[192,806,1213,1237]" pageId="5" pageNumber="4">
<taxonomicName id="4C2B4D306B479E4636B8FB1DA2A7FB3B" authority="Otto" authorityName="Otto" authorityYear="2019" box="[192,688,1213,1237]" class="Insecta" family="Eucnemidae" genus="Cretopityobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="pankowskiorum" status="sp. nov.">
<emphasis id="B95FEAA16B479E4636B8FB1DA27DFB3B" bold="true" box="[192,618,1213,1237]" italics="true" pageId="5" pageNumber="4">Cretopityobius pankowskiorum</emphasis>
Otto
</taxonomicName>
,
<taxonomicNameLabel id="A26C57DA6B479E4634C6FB1DA331FB3B" box="[702,806,1213,1237]" pageId="5" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33165386B479E4636B8FB7DA134FB18" box="[192,291,1245,1270]" pageId="5" pageNumber="4" type="description">
<paragraph id="8B9436B36B479E4636B8FB7DA134FB18" blockId="5.[192,806,1213,1270]" box="[192,291,1245,1270]" pageId="5" pageNumber="4">
<figureCitation id="13102A366B479E4636B8FB7DA134FB18" box="[192,291,1245,1270]" captionStart="Figures 1–4" captionStartId="7.[192,286,1848,1870]" captionTargetBox="[272,1360,272,1785]" captionTargetId="figure@7.[272,1360,272,1785]" captionTargetPageId="7" captionText="Figures 1–4. Fossil elateroids. 1) Cenomana clavata holotype, dorsal habitus. 2) Cenomana clavata holotype, ventral habitus. 3) Cretopityobius pankowskiorum holotype, dorsal habitus. 4) Cretopityobius pankowskiorum holotype, ventral habitus. (Scale: 1–4 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.3673249" httpUri="https://zenodo.org/record/3673249/files/figure.png" pageId="5" pageNumber="4">Fig. 3–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33165386B479E4636B8FAAEA26DF9E9" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8B9436B36B479E4636B8FAAEA26DF9E9" blockId="5.[192,1441,1294,1543]" pageId="5" pageNumber="4">
<emphasis id="B95FEAA16B479E4636B8FAAEA152FAC8" bold="true" box="[192,325,1294,1318]" pageId="5" pageNumber="4">
<typeStatus id="549088116B479E4636B8FAAEA128FAC8" box="[192,319,1294,1318]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EF460686B479E463735FAAEA180FAC8" LSID="urn:lsid:plazi:treatment:038287A56B479E4536B8FB1DA372FDEF:8EF460686B479E463735FAAEA180FAC8" box="[333,407,1294,1318]" country="Myanmar" county="Cenomanian" municipality="Piece" name="Small" pageId="5" pageNumber="4">Small</location>
, round, clear, polished yellow-colored amber piece with air bubbles, smaller insects, plant and soil debris intermixed with the specimen.
<collectingMunicipality id="6BF0ACC96B479E463558FA8EA348FAA8" box="[800,863,1326,1350]" pageId="5" pageNumber="4">Piece</collectingMunicipality>
was excavated from northern
<collectingCountry id="F33C76236B479E4632BAFA8EA52FFAA8" box="[1218,1336,1326,1350]" name="Myanmar" pageId="5" pageNumber="4">Myanmar</collectingCountry>
believed to be
<collectingCounty id="62F54E3F6B479E46377AFAEEA189FA88" box="[258,414,1358,1382]" pageId="5" pageNumber="4">Cenomanian</collectingCounty>
in age, approximately 96 million years old.
<emphasis id="B95FEAA16B479E4635CDFAEEA42CFA88" bold="true" box="[949,1083,1358,1382]" pageId="5" pageNumber="4">
<typeStatus id="549088116B479E4635CDFAEEA422FA88" box="[949,1077,1358,1382]" pageId="5" pageNumber="4" type="paratype">Paratype</typeStatus>
.
</emphasis>
Small, round, clear, polished yellow-colored amber piece with debris, including disarticulated antennae intermixed with the specimen. Piece was excavated from northern
<collectingCountry id="F33C76236B479E4634D6FA2FA331FA49" box="[686,806,1423,1447]" name="Myanmar" pageId="5" pageNumber="4">Myanmar</collectingCountry>
believed to be Cenomanian in age, approximately 96 million years old.
<materialsCitation id="3B433CEE6B479E4637E3FA0EA129FA09" ID-GBIF-Occurrence="2565950090" collectionCode="USNM" country="Myanmar" location="Washington" municipality="District of Columbia" pageId="5" pageNumber="4" specimenCount="1" typeStatus="holotype">
<typeStatus id="549088116B479E4637E3FA0EA21EFA28" box="[411,521,1454,1478]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
is deposited in the Smithsonian Institute (
<collectionCode id="ED3AAE766B479E46326DFA0FA479FA29" box="[1045,1134,1455,1479]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="4" type="Museum">USNM</collectionCode>
),
<location id="8EF460686B479E4632FAFA0EA50EFA28" LSID="urn:lsid:plazi:treatment:038287A56B479E4536B8FB1DA372FDEF:8EF460686B479E4632FAFA0EA50EFA28" box="[1154,1305,1454,1478]" country="Myanmar" municipality="District of Columbia" name="Washington" pageId="5" pageNumber="4">Washington</location>
,
<collectingMunicipality id="6BF0ACC96B479E46335DFA0EA12DFA09" pageId="5" pageNumber="4">District of Columbia</collectingMunicipality>
.
</materialsCitation>
<materialsCitation id="3B433CEE6B479E46373DFA6FA26DF9E9" ID-GBIF-Occurrence="2565950088" collectionCode="USNM" country="Myanmar" location="Washington" municipality="District of Columbia" pageId="5" pageNumber="4" specimenCount="1" typeStatus="paratype">
<typeStatus id="549088116B479E46373DFA6FA1A2FA09" box="[325,437,1487,1511]" pageId="5" pageNumber="4" type="paratype">Paratype</typeStatus>
is deposited in the Insect Research Collection (
<collectionCode id="ED3AAE766B479E46358EFA6FA451FA09" box="[1014,1094,1487,1511]" pageId="5" pageNumber="4">WIRC</collectionCode>
), UW-Madison, Department of Entomology, Madison, Wisconsin.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33165386B479E4636B8F9BEA2B2F866" pageId="5" pageNumber="4" type="description">
<paragraph id="8B9436B36B479E4636B8F9BEA338F8F6" blockId="5.[192,1441,1566,1816]" pageId="5" pageNumber="4">
<emphasis id="B95FEAA16B479E4636B8F9BEA1E0F9D9" bold="true" box="[192,503,1566,1591]" pageId="5" pageNumber="4">
Description.
<typeStatus id="549088116B479E46370AF9BFA1E6F9D9" box="[370,497,1567,1591]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
:
</emphasis>
Length,
<quantity id="4CD39B566B479E46341EF9BFA2D1F9D8" box="[614,710,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="4" unit="mm" value="5.5">5.5 mm</quantity>
. Width,
<quantity id="4CD39B566B479E463555F9BFA39AF9D8" box="[813,909,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="4" unit="mm" value="1.5">1.5 mm</quantity>
. Elongate and tapering towards the elytral apex; head, pronotum, elytra and venter dark brown; antennae and legs dark brown; head, pronotum and elytra clothed with long, erect setae (
<figureCitation id="13102A366B479E46340EF9FFA2ACF999" box="[630,699,1631,1655]" captionStart="Figures 1–4" captionStartId="7.[192,286,1848,1870]" captionTargetBox="[272,1360,272,1785]" captionTargetId="figure@7.[272,1360,272,1785]" captionTargetPageId="7" captionText="Figures 1–4. Fossil elateroids. 1) Cenomana clavata holotype, dorsal habitus. 2) Cenomana clavata holotype, ventral habitus. 3) Cretopityobius pankowskiorum holotype, dorsal habitus. 4) Cretopityobius pankowskiorum holotype, ventral habitus. (Scale: 1–4 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.3673249" httpUri="https://zenodo.org/record/3673249/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
).
<emphasis id="B95FEAA16B479E4634B7F9FFA308F999" bold="true" box="[719,799,1631,1655]" pageId="5" pageNumber="4">Head:</emphasis>
Dorso-ventrally flattened; surfaces shiny with closely spaced, shallow punctures; remaining areas of head obscured by air bubble; dark brown.
<emphasis id="B95FEAA16B479E46336CF9DFA5B7F979" bold="true" box="[1300,1440,1663,1687]" pageId="5" pageNumber="4">Antennae:</emphasis>
Long, extending almost 2/3 the length of its body; dark brown.
<emphasis id="B95FEAA16B479E4635CCF93FA45FF959" bold="true" box="[948,1096,1695,1719]" pageId="5" pageNumber="4">Pronotum:</emphasis>
Convex; surfaces glossy and transversely, shallowly rugose; base sinuous; about 1/2 of surface obscured by air bubble; dark brown.
<emphasis id="B95FEAA16B479E4636B8F940A109F916" bold="true" box="[192,286,1760,1784]" pageId="5" pageNumber="4">Elytra:</emphasis>
Surfaces somewhat shiny and transversely rugose; dark brown.
<emphasis id="B95FEAA16B479E46325AF940A497F916" bold="true" box="[1058,1152,1760,1784]" pageId="5" pageNumber="4">Venter</emphasis>
(
<figureCitation id="13102A366B479E4632F6F940A4C3F916" box="[1166,1236,1760,1784]" captionStart="Figures 1–4" captionStartId="7.[192,286,1848,1870]" captionTargetBox="[272,1360,272,1785]" captionTargetId="figure@7.[272,1360,272,1785]" captionTargetPageId="7" captionText="Figures 1–4. Fossil elateroids. 1) Cenomana clavata holotype, dorsal habitus. 2) Cenomana clavata holotype, ventral habitus. 3) Cretopityobius pankowskiorum holotype, dorsal habitus. 4) Cretopityobius pankowskiorum holotype, ventral habitus. (Scale: 1–4 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.3673249" httpUri="https://zenodo.org/record/3673249/files/figure.png" pageId="5" pageNumber="4">Fig. 4</figureCitation>
): Surfaces glossy with shallow indications of punctures; dark brown.
</paragraph>
<paragraph id="8B9436B36B479E4636B8F890A2B2F866" blockId="5.[192,1440,1840,1929]" pageId="5" pageNumber="4">
<emphasis id="B95FEAA16B479E4636B8F890A15CF8A6" bold="true" box="[192,331,1840,1864]" pageId="5" pageNumber="4">Variation.</emphasis>
The
<typeStatus id="549088116B479E4637F3F891A1ECF8A7" box="[395,507,1841,1865]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
, although in its disarticulated condition in a piece of amber, is much larger and wider than the
<typeStatus id="549088116B479E4637C3F8F0A234F886" box="[443,547,1872,1896]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
at 7.0 mm long and 2.0 mm wide. There are no exoskeletal differences between the
<typeStatus id="549088116B479E463724F8D1A1DEF867" box="[348,457,1905,1929]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
and the
<typeStatus id="549088116B479E46344EF8D0A2B5F866" box="[566,674,1904,1928]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33165386B449E4536B8FF79A2A8FEFF" pageId="6" pageNumber="5" type="etymology">
<paragraph id="8B9436B36B449E4536B8FF79A2A8FEFF" blockId="6.[192,1440,217,273]" pageId="6" pageNumber="5">
<emphasis id="B95FEAA16B449E4536B8FF79A14CFF1F" bold="true" box="[192,347,217,241]" pageId="6" pageNumber="5">Etymology.</emphasis>
The specific epithet is dedicated to the Pankowski family of Rockville,
<collectingRegion id="49EFF8516B449E4532DDFF79A50AFF1F" box="[1189,1309,217,241]" country="United States of America" name="Maryland" pageId="6" pageNumber="5">Maryland</collectingRegion>
for provid- ing these interesting specimens for study.
</paragraph>
</subSubSection>
<subSubSection id="C33165386B449E4536B8FE89A372FDEF" pageId="6" pageNumber="5" type="discussion">
<paragraph id="8B9436B36B449E4536B8FE89A372FDEF" blockId="6.[192,1440,297,513]" pageId="6" pageNumber="5">
<emphasis id="B95FEAA16B449E4536B8FE89A156FEAF" bold="true" box="[192,321,297,321]" pageId="6" pageNumber="5">Remarks.</emphasis>
These specimens are assigned to the subfamily
<taxonomicName id="4C2B4D306B449E45350FFE89A417FEAF" authorityName="Hyslop" authorityYear="1917" box="[887,1024,297,321]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Pityobiinae">Pityobiinae</taxonomicName>
based on the smaller antennomere III in relation to II, bipectinate antennae, and general appearance similar to Nearctic species of
<taxonomicName id="4C2B4D306B449E453321FEEAA0E3FE6F" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Elateridae" genus="Pityobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA16B449E453321FEEAA0E3FE6F" italics="true" pageId="6" pageNumber="5">Pityobius</emphasis>
</taxonomicName>
. In addition to these traits, both genera also have impressed frons above the frontoclypeal region. The new species differs from
<taxonomicName id="4C2B4D306B449E453466FE29A287FE4F" authorityName="LeConte" authorityYear="1853" box="[542,656,393,417]" class="Insecta" family="Elateridae" genus="Pityobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA16B449E453466FE29A287FE4F" box="[542,656,393,417]" italics="true" pageId="6" pageNumber="5">Pityobius</emphasis>
</taxonomicName>
based on its overall relative size, being smaller in
<taxonomicName id="4C2B4D306B449E453297FE29A5B7FE4F" authority="Otto, 2019" authorityName="Otto" authorityYear="2019" box="[1263,1440,393,417]" class="Insecta" family="Eucnemidae" genus="Cretopityobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FEAA16B449E453297FE29A5B7FE4F" box="[1263,1440,393,417]" italics="true" pageId="6" pageNumber="5">Cretopityobius</emphasis>
</taxonomicName>
(5.5–7.0 mm long), larger in
<taxonomicName id="4C2B4D306B449E453461FE09A29CFE2F" authorityName="LeConte" authorityYear="1853" box="[537,651,425,449]" class="Insecta" family="Elateridae" genus="Pityobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA16B449E453461FE09A29CFE2F" box="[537,651,425,449]" italics="true" pageId="6" pageNumber="5">Pityobius</emphasis>
</taxonomicName>
(15.0–39.0 mm long) as well as the placement of rami in relation to each antennal segment. The rami arise from the base of each segment in
<taxonomicName id="4C2B4D306B449E453224FE69A51AFE0F" authority="Otto, 2019" authorityName="Otto" authorityYear="2019" box="[1116,1293,457,481]" class="Insecta" family="Eucnemidae" genus="Cretopityobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95FEAA16B449E453224FE69A51AFE0F" box="[1116,1293,457,481]" italics="true" pageId="6" pageNumber="5">Cretopityobius</emphasis>
</taxonomicName>
, while rami arise from the apices of each antennomere in
<taxonomicName id="4C2B4D306B449E453494FE49A349FDEF" authorityName="LeConte" authorityYear="1853" box="[748,862,489,513]" class="Insecta" family="Elateridae" genus="Pityobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA16B449E453494FE49A349FDEF" box="[748,862,489,513]" italics="true" pageId="6" pageNumber="5">Pityobius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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