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<document id="1027E24BB206BF75185C6BBA9DED42A5" ID-CLB-Dataset="9764" ID-DOI="10.5852/ejt.2020.720.1109" ID-GBIF-Dataset="545aa4ea-1874-4f74-a4c4-a2aeb97ed657" ID-ISSN="2118-9773" ID-Zenodo-Dep="4068355" ID-ZooBank="0CF83E89-2532-407B-921F-C05F9A29C68D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1601993099685" checkinUser="valdenar" docAuthor="Bergsten, Johannes, Ranarilalatiana, Tolotra &amp; Biström, Olof" docDate="2020" docId="03828798D370646FFD94FC16AFC4FE23" docLanguage="en" docName="ejt-720_begsten_ranarilalatinana_bistroem.pdf.imf" docOrigin="European Journal of Taxonomy 720" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bidessus longistriga Regimbart 1895" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="FFBBFFE0D3756468FFAEFFBDAD21FFBB" masterDocTitle="A new species of Bidessus from Anjozorobe-Angavo and a review of Malagasy Bidessus (Coleoptera, Dytiscidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="6" updateTime="1698792528866" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CCA09697DDA78E7F6D7212A73EBA81AB">A new species of Bidessus from Anjozorobe-Angavo and a review of Malagasy Bidessus (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="AF58DE2AF13A7317F536D8C551DCB49B">Bergsten, Johannes</mods:namePart>
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<mods:affiliation id="0DB1AF944E1BF4771C693904F2E93292">Department of Zoology, Swedish Museum of Natural History, Box 50007, SE- 10405 Stockholm, Sweden. Department of Entomology, Faculty of Sciences, Box 906, University of Antananarivo, 101 Antananarivo, Madagascar. Finnish Museum of Natural History, Zoology Unit, P. O. Box 17, FI- 00014 University of Helsinki, Finland.</mods:affiliation>
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<mods:affiliation id="867334EE96EB7FF121AB2FBBB8A7D0D6">Department of Entomology, Faculty of Sciences, Box 906, University of Antananarivo, 101 Antananarivo, Madagascar.</mods:affiliation>
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<emphasis id="B95FEA9CD370646DFD94FC16AE0CFC7D" bold="true" box="[570,813,939,966]" italics="true" pageId="5" pageNumber="6">Bidessus longistriga</emphasis>
<bibRefCitation id="EFBA4B7FD370646DFC9BFC11AEDBFC7C" author="Regimbart M." box="[821,1018,940,967]" pageId="5" pageNumber="6" pagination="1 - 244" refId="ref7828" refString="Regimbart M. 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. Memoires de la Societe entomologique de Belgique 4: 1 - 244." type="journal article" year="1895">Régimbart, 1895</bibRefCitation>
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<figureCitation id="13102A0BD370646DFD7CFC72AE10FC52" box="[722,817,975,1001]" captionStart="Fig" captionStartId="6.[189,232,1922,1948]" captionTargetBox="[223,1365,265,1885]" captionTargetId="figure-14@6.[223,1365,265,1885]" captionTargetPageId="6" captionText="Fig. 2. Habitus photographs of Malagasy species of Bidessus Sharp, 1882. A. B. nesioticus Guignot, 1956 (NHRS-JLKB000025031). B. B. perexiguus Kolbe, 1883 (NHRS-JLKB000025029). C. B. longistriga Régimbart, 1895 (NHRS-JLKB000025028)." figureDoi="http://doi.org/10.5281/zenodo.4068359" httpUri="https://zenodo.org/record/4068359/files/figure.png" pageId="5" pageNumber="6">Figs 2C</figureCitation>
,
<figureCitation id="13102A0BD370646DFC93FC72AE40FC52" box="[829,865,975,1001]" captionStart="Fig" captionStartId="9.[189,232,1888,1914]" captionTargetBox="[366,1209,282,1833]" captionTargetId="figure-11@9.[308,1280,265,1849]" captionTargetPageId="9" captionText="Fig. 3. Male genitalia of the Malagasy species of Bidessus Sharp, 1882. For each subpanel from left to right penis in dorsal view, penis in lateral view and paramere in lateral view. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986. B. B. cf. ceratus Guignot, 1941. C. B. anjozorobe sp. nov. D. B. cf. nero Gschwendtner, 1933. E. B. nesioticus Guignot, 1956. F. B. perexiguus Kolbe, 1883. G. B. longistriga Régimbart, 1895." figureDoi="http://doi.org/10.5281/zenodo.4068361" httpUri="https://zenodo.org/record/4068361/files/figure.png" pageId="5" pageNumber="6">3G</figureCitation>
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<emphasis id="B95FEA9CD370646DFF13FBADAC8DFB91" box="[189,428,1040,1066]" italics="true" pageId="5" pageNumber="6">Bidessus longistriga</emphasis>
<bibRefCitation id="EFBA4B7FD370646DFE1DFBADAF88FB90" author="Regimbart M." box="[435,681,1040,1067]" pageId="5" pageNumber="6" pagination="1 - 244" refId="ref7828" refString="Regimbart M. 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. Memoires de la Societe entomologique de Belgique 4: 1 - 244." type="journal article" year="1895">Régimbart, 1895: 79</bibRefCitation>
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<emphasis id="B95FEA9CD370646DFF13FBEFAC8DFBD7" box="[189,428,1106,1132]" italics="true" pageId="5" pageNumber="6">Bidessus longistriga</emphasis>
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<treatmentCitation id="0A8A109FD370646DFE69FBEFAFB6FBD6" author="Alluaud C." box="[455,663,1106,1133]" page="59" pageId="5" pageNumber="6" year="1900">
<bibRefCitation id="EFBA4B7FD370646DFE69FBEFAFB6FBD6" author="Alluaud C." box="[455,663,1106,1133]" pageId="5" pageNumber="6" pagination="56 - 76" refId="ref7210" refString="Alluaud C. 1900. Liste des insectes coleopteres de la region malgache. In: Grandidier A. (ed.) Histoire physique, naturelle et politique de Madagascar volume 21, Histoire naturelle des Coleopteres, tome I, texte, 1 er partie: 56 - 76. Imprimerie nationale, Paris. https: // doi. org / 10.5962 / bhl. title. 1599" type="book chapter" year="1900">Alluaud 1900: 59</bibRefCitation>
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. —
<treatmentCitation id="0A8A109FD370646DFD69FBEFAEF9FBD6" author="Zimmermann A." box="[711,984,1106,1133]" page="54" pageId="5" pageNumber="6" year="1920">
<bibRefCitation id="EFBA4B7FD370646DFD69FBEFAEF9FBD6" author="Zimmermann A." box="[711,984,1106,1133]" pageId="5" pageNumber="6" pagination="71" refId="ref7976" refString="Zimmermann A. 1920. Pars 71. Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S. (ed.) Coleopterorum Catalogus, Volumen IV. Junk, Berlin." type="book chapter" year="1920">Zimmermann 1920: 54</bibRefCitation>
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<bibRefCitation id="EFBA4B7FD370646DFBA6FBEFA9C8FBD6" author="Guignot F." box="[1032,1257,1106,1133]" pageId="5" pageNumber="6" pagination="1 - 313" refId="ref7502" refString="Guignot F. 1959. Revision des Hydrocanthares d'Afrique (Coleoptera: Dytiscoidea). Premiere Partie. Annales de Musee royal du Congo Belge, Tervuren: Sciences zoologiques 70: 1 - 313." type="journal article" year="1959">Guignot 1959: 254</bibRefCitation>
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. — Bistr̂m 1985: 7. —
<treatmentCitation id="0A8A109FD370646DFED8FBC8AF1AFB2B" author="Rocchi S." box="[374,571,1141,1168]" page="85" pageId="5" pageNumber="6" year="1991">
<bibRefCitation id="EFBA4B7FD370646DFED8FBC8AF1AFB2B" author="Rocchi S." box="[374,571,1141,1168]" pageId="5" pageNumber="6" pagination="71 - 89" refId="ref7858" refString="Rocchi S. 1991. Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera). Frustula Entomologica (Nuova Serie) 14: 71 - 89." type="journal article" year="1991">Rocchi 1991: 85</bibRefCitation>
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<treatmentCitation id="0A8A109FD370646DFDC4FBC8AE63FB2B" author="Nilsson A. N." box="[618,834,1141,1168]" page="115" pageId="5" pageNumber="6" year="2001">
<bibRefCitation id="EFBA4B7FD370646DFDC4FBC8AE63FB2B" author="Nilsson A. N." box="[618,834,1141,1168]" pageId="5" pageNumber="6" pagination="1 - 395" refId="ref7602" refString="Nilsson A. N. 2001. Dytiscidae (Coleoptera). World Catalogue of Insects 3: 1 - 395." type="journal article" year="2001">Nilsson 2001: 115</bibRefCitation>
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<treatmentCitation id="0A8A109FD370646DFCDCFBC8A9F1FB34" author="Pederzani F. &amp; Rocchi S." box="[882,1232,1141,1167]" page="93" pageId="5" pageNumber="6" year="2009">
<bibRefCitation id="EFBA4B7FD370646DFCDCFBC8A9F1FB34" author="Pederzani F. &amp; Rocchi S." box="[882,1232,1141,1167]" pageId="5" pageNumber="6" pagination="81 - 100" refId="ref7701" refString="Pederzani F. &amp; Rocchi S. 2009. On some Madagascan Hydradephaga of The Natural History Museum, London, with the descriptions of five new species of Dytiscidae (Coleoptera Haliplidae, Dytiscidae, Noteridae). Atti dell'Accademia Roveretana degli Agiati, Contributi della Classe di Scienze Matematiche, Fisiche e Naturali (8) 8 B (258) (2008): 81 - 100." type="journal article" year="2009">Pederzani &amp; Rocchi 2009: 93</bibRefCitation>
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<treatmentCitation id="0A8A109FD370646DFB51FBC8AC90FB09" author="Nilsson A. N. &amp; Hajek J." page="104" pageId="5" pageNumber="6" year="2020">
<bibRefCitation id="EFBA4B7FD370646DFB51FBC8AC90FB09" author="Nilsson A. N. &amp; Hajek J." pageId="5" pageNumber="6" pagination="1 - 395" refId="ref7624" refString="Nilsson A. N. &amp; Hajek J. 2020. A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga) Version 1. I. 2020. Electronic and updated version of Nilsson A. N. (2001). Dytiscidae. In: World Catalogue of Insects: 1 - 395. Available from www. waterbeetles. eu [accessed 5 Sep. 2020]." type="book chapter" year="2020">Nilsson &amp; Hajek 2020: 104</bibRefCitation>
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<emphasis id="B95FEA9CD370646DFF13FB64ACEDFB4F" bold="true" box="[189,460,1241,1268]" pageId="5" pageNumber="6">Diagnostic characters</emphasis>
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Head frontally not margined but provided with a shallow, transverse depression slightly posterior to anterior edge of head (between eyes). Pronotum on disc with a broad impunctate area or with a few fine punctures, distinctly sparser than punctures at frontal and posterior edge. Pronotal striae at base deeply impressed. Pronotal striae with a distinct angle (not smoothly curved). Basal striae of elytra long. Sutural lines distinct; anteriorly and posteriorly lines fade away; transformed to row of punctures. Punctures on both side of basal striae of unequal size; distinctly coarser on inner side. Apical sternite with fine, slightly irregular punctures. Female with outline of elytra posteriorly smooth (minor extension absent). Penis in lateral view slender, curved but medially somewhat straightened. Penis apex not distinctly extended (
<figureCitation id="13102A0BD370646DFE99F99FACB4F987" box="[311,405,1570,1596]" captionStart="Fig" captionStartId="9.[189,232,1888,1914]" captionTargetBox="[366,1209,282,1833]" captionTargetId="figure-11@9.[308,1280,265,1849]" captionTargetPageId="9" captionText="Fig. 3. Male genitalia of the Malagasy species of Bidessus Sharp, 1882. For each subpanel from left to right penis in dorsal view, penis in lateral view and paramere in lateral view. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986. B. B. cf. ceratus Guignot, 1941. C. B. anjozorobe sp. nov. D. B. cf. nero Gschwendtner, 1933. E. B. nesioticus Guignot, 1956. F. B. perexiguus Kolbe, 1883. G. B. longistriga Régimbart, 1895." figureDoi="http://doi.org/10.5281/zenodo.4068361" httpUri="https://zenodo.org/record/4068361/files/figure.png" pageId="5" pageNumber="6">Fig. 3G</figureCitation>
).
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Length of body
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<emphasis id="B95FEA9CD370646DFF13F918AC8BF904" bold="true" box="[189,426,1701,1727]" pageId="5" pageNumber="6">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED370646DFF13F96EAE97F8AA" blockId="5.[189,1398,1747,1809]" pageId="5" pageNumber="6">
<specimenCount id="9D2DFD07D370646DFF13F96EAC4BF956" box="[189,362,1747,1773]" pageId="5" pageNumber="6" type="generic">283 specimens</specimenCount>
(
<collectionCode id="ED3AAE4BD370646DFED4F96EACF6F956" box="[378,471,1747,1773]" pageId="5" pageNumber="6">BMNH</collectionCode>
,
<collectionCode id="ED3AAE4BD370646DFE4CF96EAF12F956" box="[482,563,1747,1773]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="5" pageNumber="6">NHRS</collectionCode>
,
<collectionCode id="ED3AAE4BD370646DFD91F969AFABF955" box="[575,650,1748,1774]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="6">NMW</collectionCode>
,
<collectionCode id="ED3AAE4BD370646DFD38F96EAFFEF956" box="[662,735,1747,1773]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="5" pageNumber="6">PBZT</collectionCode>
/
<collectionCode id="ED3AAE4BD370646DFD47F96EAE0AF956" box="[745,811,1747,1773]" country="USA" name="Montgomery Botanical Center" pageId="5" pageNumber="6">MBC</collectionCode>
,
<collectionCode id="ED3AAE4BD370646DFC98F969AEA8F955" box="[822,905,1748,1774]" pageId="5" pageNumber="6">DEUA</collectionCode>
) from provinces
<collectingRegion id="49EFF86CD370646DFBFEF96EA9C0F956" box="[1104,1249,1747,1773]" country="Madagascar" name="Antsiranana" pageId="5" pageNumber="6">Antsiranana</collectingRegion>
,
<collectingRegion id="49EFF86CD370646DFB43F96EA853F956" box="[1261,1394,1747,1773]" country="Madagascar" name="Mahajanga" pageId="5" pageNumber="6">Mahajanga</collectingRegion>
,
<collectingRegion id="49EFF86CD370646DFF13F94BAC32F8AB" box="[189,275,1782,1808]" country="Madagascar" name="Toliara" pageId="5" pageNumber="6">Toliara</collectingRegion>
,
<collectingRegion id="49EFF86CD370646DFEB0F94BAC95F8AB" box="[286,436,1782,1808]" country="Madagascar" name="Fianarantsoa" pageId="5" pageNumber="6">Fianarantsoa</collectingRegion>
and
<collectingRegion id="49EFF86CD370646DFE43F94BAF4DF8AB" box="[493,620,1782,1808]" country="Madagascar" name="Toamasina" pageId="5" pageNumber="6">Toamasina</collectingRegion>
(see Supplementary File 1).
</paragraph>
</subSubSection>
<subSubSection id="C3316505D370646DFF13F885AC13F81F" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8B94368ED370646DFF13F885AC77F8E9" blockId="5.[189,342,1848,1874]" box="[189,342,1848,1874]" pageId="5" pageNumber="6">
<heading id="D0DC81E2D370646DFF13F885AC77F8E9" bold="true" box="[189,342,1848,1874]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B95FEA9CD370646DFF13F885AC77F8E9" bold="true" box="[189,342,1848,1874]" pageId="5" pageNumber="6">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED370646DFF13F8DBAC13F81F" blockId="5.[189,1399,1894,1956]" pageId="5" pageNumber="6">
Endemic species to
<collectingCountry id="F33C761ED370646DFE0BF8DBAF12F83B" box="[421,563,1894,1920]" name="Madagascar" pageId="5" pageNumber="6">Madagascar</collectingCountry>
but widely distributed on the island in lowland areas below
<quantity id="4CD39B6BD370646DFB5EF8DAA818F83A" box="[1264,1337,1895,1921]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="6" unit="m" value="800.0">800 m</quantity>
a.s.l. (
<figureCitation id="13102A0BD370646DFF68F837AC04F81F" box="[198,293,1930,1956]" captionStart="Fig" captionStartId="11.[189,232,1888,1914]" captionTargetBox="[270,1323,299,1826]" captionTargetId="figure-12@11.[232,1355,265,1852]" captionTargetPageId="11" captionText="Fig. 4. Distribution maps of the Malagasy species of Bidessus Sharp, 1882. A. B. longistriga Régimbart, 1895. B. B. perexiguus Kolbe, 1883. C. B. cf. nero Gschwendtner, 1933 (circle), B. nesioticus Guignot, 1956 (square), B. anjozorobe sp. nov. (triangle). D. B. apicidens Biström &amp; Sanfilippo, 1986 (square), B. cf. ceratus Guignot, 1941 (circle)." figureDoi="http://doi.org/10.5281/zenodo.4068363" httpUri="https://zenodo.org/record/4068363/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF546606D373646EFF13F83FAFE7F858" ID-DOI="http://doi.org/10.5281/zenodo.4068359" ID-Zenodo-Dep="4068359" httpUri="https://zenodo.org/record/4068359/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1922,1948]" targetBox="[223,1365,265,1885]" targetPageId="6">
<paragraph id="8B94368ED373646EFF13F83FAFE7F858" blockId="6.[189,1399,1922,2019]" pageId="6" pageNumber="7">
<emphasis id="B95FEA9CD373646EFF13F83FAC29F827" bold="true" box="[189,264,1922,1948]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
Habitus photographs of Malagasy species of
<taxonomicName id="4C2B4D0DD373646EFCBEF83FA92AF827" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[784,1035,1922,1948]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEA9CD373646EFCBEF83FAE56F827" box="[784,887,1922,1948]" italics="true" pageId="6" pageNumber="7">Bidessus</emphasis>
Sharp, 1882
</taxonomicName>
.
<emphasis id="B95FEA9CD373646EFBBAF83FA908F827" bold="true" box="[1044,1065,1922,1948]" pageId="6" pageNumber="7">A</emphasis>
.
<taxonomicName id="4C2B4D0DD373646EFB98F83EA856F826" authority="Guignot, 1956" authorityName="Guignot" authorityYear="1956" box="[1078,1399,1922,1949]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nesioticus">
<emphasis id="B95FEA9CD373646EFB98F83EA9EBF827" box="[1078,1226,1923,1948]" italics="true" pageId="6" pageNumber="7">B. nesioticus</emphasis>
<bibRefCitation id="EFBA4B7FD373646EFB61F83FA856F826" author="Guignot F." box="[1231,1399,1922,1949]" pageId="6" pageNumber="7" pagination="75 - 80" refId="ref7478" refString="Guignot F. 1956. Contribution a la connaissance de la faune dystiscidienne Malgache. Le Naturaliste malgache 8: 75 - 80." type="journal article" year="1956">Guignot, 1956</bibRefCitation>
</taxonomicName>
(NHRS-JLKB000025031).
<emphasis id="B95FEA9CD373646EFDACF81BAF37F87B" bold="true" box="[514,534,1958,1984]" pageId="6" pageNumber="7">B</emphasis>
.
<taxonomicName id="4C2B4D0DD373646EFD88F81BAE43F87B" authority="Kolbe, 1883" authorityName="Kolbe" authorityYear="1883" box="[550,866,1957,1984]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="perexiguus">
<emphasis id="B95FEA9CD373646EFD88F81BAFE6F804" box="[550,711,1958,1983]" italics="true" pageId="6" pageNumber="7">B. perexiguus</emphasis>
<bibRefCitation id="EFBA4B7FD373646EFD61F818AE43F87B" author="Kolbe H. J." box="[719,866,1957,1984]" pageId="6" pageNumber="7" pagination="383 - 427" refId="ref7538" refString="Kolbe H. J. 1883. Uber die madagascarischen Dytisciden des K ˆ nigl. entomologischen Museums zu Berlin. Archiv fur Naturgeschichte 49 (1): 383 - 427." type="journal article" year="1883">Kolbe, 1883</bibRefCitation>
</taxonomicName>
(NHRS-JLKB000025029).
<emphasis id="B95FEA9CD373646EFB01F818A9E5F804" bold="true" box="[1199,1220,1957,1983]" pageId="6" pageNumber="7">C</emphasis>
.
<taxonomicName id="4C2B4D0DD373646EFB7AF81BACA3F858" authority="Regimbart, 1895" authorityName="Regimbart" authorityYear="1895" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longistriga">
<emphasis id="B95FEA9CD373646EFB7AF81BA856F804" box="[1236,1399,1957,1983]" italics="true" pageId="6" pageNumber="7">B. longistriga</emphasis>
<bibRefCitation id="EFBA4B7FD373646EFF13F874ACA3F858" author="Regimbart M." box="[189,386,1992,2019]" pageId="6" pageNumber="7" pagination="1 - 244" refId="ref7828" refString="Regimbart M. 1895. Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. Memoires de la Societe entomologique de Belgique 4: 1 - 244." type="journal article" year="1895">Régimbart, 1895</bibRefCitation>
</taxonomicName>
(NHRS-JLKB000025028).
</paragraph>
</caption>
<subSubSection id="C3316505D372646FFF13FEB5ACD5FE99" box="[189,500,264,290]" pageId="7" pageNumber="8" type="description">
<paragraph id="8B94368ED372646FFF13FEB5ACD5FE99" blockId="7.[189,500,264,290]" box="[189,500,264,290]" pageId="7" pageNumber="8">
<heading id="D0DC81E2D372646FFF13FEB5ACD5FE99" bold="true" box="[189,500,264,290]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B95FEA9CD372646FFF13FEB5ACD5FE99" bold="true" box="[189,500,264,290]" pageId="7" pageNumber="8">Collecting circumstances</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3316505D372646FFF13FE8AAFC4FE23" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B94368ED372646FFF13FE8AAFC4FE23" blockId="7.[189,1399,311,408]" pageId="7" pageNumber="8">
An apt flier that can be collected at light. At low altitudes on both east and west coast, found in a variety of stagnant waters (rice fields, ponds, pools in riverbeds), and in slow-flowing parts of rivers and canals. Often occurs sympatrically with
<taxonomicName id="4C2B4D0DD372646FFD93FEC3AFFFFE2C" ID-CoL="LSRL" authorityName="Kolbe" authorityYear="1883" box="[573,734,382,407]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="perexiguus">
<emphasis id="B95FEA9CD372646FFD93FEC3AFFFFE2C" box="[573,734,382,407]" italics="true" pageId="7" pageNumber="8">B. perexiguus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>