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<document id="3B9A41CBB122142E3AC31CE5846FB71A" ID-CLB-Dataset="25546" ID-DOI="10.11646/zootaxa.4647.1.28" ID-GBIF-Dataset="ce397603-2e1b-45d7-a660-ab4771887774" ID-ISSN="1175-5326" ID-Zenodo-Dep="3353292" ID-ZooBank="76D3D826-7173-444A-BDB4-F2537F325155" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564250778104" checkinUser="plazi" docAuthor="Klimov, Pavel B., Oconnor, Barry &amp; Khaustov, Alexander A." docDate="2019" docId="03DB8787FFD8FF897DD449A6FD4EACBE" docLanguage="en" docName="zootaxa.4647.1.28.pdf" docOrigin="Zootaxa 4647 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naiadacarus sidorchukae Klimov &amp; Oconnor &amp; Khaustov 2019, sp. n." docType="treatment" docVersion="10" lastPageNumber="450" masterDocId="FFE2FFFFFFDFFF807D43497CFFC4AB28" masterDocTitle="Naiadacarus sidorchukae sp. n. (Acariformes: Acaridae) from Western Siberia closely related to a North American species, with a new generic synonymy" masterLastPageNumber="456" masterPageNumber="441" pageNumber="448" updateTime="1698747673551" updateUser="ExternalLinkService">
<mods:mods id="84B0E8DC13830A7AB0C17B721F280C1E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B40EF067AF684C980CB9848A5F5E9183">Naiadacarus sidorchukae sp. n. (Acariformes: Acaridae) from Western Siberia closely related to a North American species, with a new generic synonymy</mods:title>
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<mods:namePart id="3544E7F1831C22C518BB7BD0B56CECDA">Klimov, Pavel B.</mods:namePart>
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<mods:namePart id="28A12B9440D2708AABB95B209FCB914A">Oconnor, Barry</mods:namePart>
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<mods:namePart id="CD9715336D0207282E37326D8EB16829">Khaustov, Alexander A.</mods:namePart>
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<mods:date id="D1376924DDC6FBE151BC903278886F29">2019</mods:date>
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<mods:number id="850A63E6052AD13549D8214C9826CD9D">2019-07-26</mods:number>
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<mods:number id="452E32765A78869EE4A34D040DE6EF14">4647</mods:number>
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<treatment id="03DB8787FFD8FF897DD449A6FD4EACBE" ID-GBIF-Taxon="159398376" LSID="urn:lsid:plazi:treatment:03DB8787FFD8FF897DD449A6FD4EACBE" httpUri="http://treatment.plazi.org/id/03DB8787FFD8FF897DD449A6FD4EACBE" lastPageId="9" lastPageNumber="450" pageId="7" pageNumber="448">
<subSubSection id="C368651AFFD8FF877DD449A6FD0EAA18" pageId="7" pageNumber="448" type="nomenclature">
<paragraph id="8BCD3691FFD8FF877DD449A6FDDBABDD" blockId="7.[151,543,218,245]" box="[151,543,218,245]" pageId="7" pageNumber="448">
<heading id="D08581FDFFD8FF877DD449A6FDDBABDD" bold="true" box="[151,543,218,245]" fontSize="11" level="1" pageId="7" pageNumber="448" reason="1">
<emphasis id="B906EA83FFD8FF877DD449A6FDDBABDD" bold="true" box="[151,543,218,245]" pageId="7" pageNumber="448">
<taxonomicName id="4C724D12FFD8FF877DD449A6FE09ABDD" authority="Klimov &amp; Oconnor &amp; Khaustov, 2019" authorityName="Klimov &amp; Oconnor &amp; Khaustov" authorityYear="2019" box="[151,461,218,245]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="448" phylum="Arthropoda" rank="species" species="sidorchukae" status="sp. nov.">
<emphasis id="B906EA83FFD8FF877DD449A6FE09ABDD" bold="true" box="[151,461,218,245]" italics="true" pageId="7" pageNumber="448">Naiadacarus sidorchukae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23557F8FFD8FF877C9F49A7FDDBABDD" box="[476,543,219,245]" pageId="7" pageNumber="448" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCD3691FFD8FF877DD44864FD0EAA18" blockId="7.[151,714,280,305]" box="[151,714,280,305]" pageId="7" pageNumber="448">
<taxonomicName id="4C724D12FFD8FF877DD44864FEDEAA18" authorityName="Fashing" authorityYear="1974" box="[151,282,280,304]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA83FFD8FF877DD44864FEDEAA18" box="[151,282,280,304]" italics="true" pageId="7" pageNumber="448">Naiadacarus</emphasis>
</taxonomicName>
sp. Khaustov
<emphasis id="B906EA83FFD8FF877CE84865FE1DAA18" box="[427,473,280,304]" italics="true" pageId="7" pageNumber="448">at al</emphasis>
., 2018: 33, Fig. 20C,D.
</paragraph>
</subSubSection>
<subSubSection id="C368651AFFD8FF897DD44824FD4EACBE" lastPageId="9" lastPageNumber="450" pageId="7" pageNumber="448" type="materials_examined">
<paragraph id="8BCD3691FFD8FF877DD44824FA5FA92A" blockId="7.[151,1437,343,1127]" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877DD44824FE83AA5A" bold="true" box="[151,327,343,370]" pageId="7" pageNumber="448">Type material.</emphasis>
<typeStatus id="54C98833FFD8FF877C0C4824FE78AA5A" box="[335,444,344,370]" pageId="7" pageNumber="448" type="holotype">Holotype</typeStatus>
: phoretic deutonymph (slide 28.1)—
<collectingCountry id="F3657601FFD8FF877E1E4824FC00AA5A" box="[861,964,344,370]" name="Russia" pageId="7" pageNumber="448">RUSSIA</collectingCountry>
,
<collectingRegion id="49B6F873FFD8FF877E934825FBBAAA5A" box="[976,1150,344,371]" country="Russia" name="Tyumen'" pageId="7" pageNumber="448">Tyumen region</collectingRegion>
, near Uspenka,
<geoCoordinate id="EE465056FFD8FF877876482BFA5CAA5B" box="[1333,1432,343,371]" degrees="57" direction="north" minutes="04" orientation="latitude" pageId="7" pageNumber="448" precision="925" value="57.066666">57°04 N</geoCoordinate>
,
<geoCoordinate id="EE465056FFD8FF877DD44807FF31AABF" box="[151,245,379,407]" degrees="65" direction="east" minutes="04" orientation="longitude" pageId="7" pageNumber="448" precision="925" value="65.066666">65°04 E</geoCoordinate>
, pine-spruce coniferous forest, ethanol wash of pheromone trap with numerous specimens of
<taxonomicName id="4C724D12FFD8FF8778514801FF08AA92" class="Insecta" family="Curculionidae" genus="Ips" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="species" species="typographus">
<emphasis id="B906EA83FFD8FF8778514801FF08AA92" italics="true" pageId="7" pageNumber="448">Ips typographus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C724D12FFD8FF877D9948DCFE9FAA92" authorityName="Linnaeus" authorityYear="1758" box="[218,347,416,442]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C724D12FFD8FF877C2548DCFDCDAA92" authorityName="Latreille" authorityYear="1802" box="[358,521,416,442]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="4C724D12FFD8FF877F5048DCFD4FAA93" authorityName="Latreille" authorityYear="1804" box="[531,651,416,443]" class="Insecta" family="Scolytinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="448" phylum="Arthropoda" rank="family">Scolytinae</taxonomicName>
) collected by this trap,
<date id="FFCC1051FFD8FF877ED348DCFBECAA92" box="[912,1064,416,443]" pageId="7" pageNumber="448" value="2017-05-22">22 May 2017</date>
, A. Khaustov.
<typeStatus id="54C98833FFD8FF87798848DDFAF8AA93" box="[1227,1340,417,443]" pageId="7" pageNumber="448" type="paratype">Paratypes</typeStatus>
: 30 phoretic deutonymphs (slides 10, 11, 13, 14, 15, 16, 17, 18, 24, 27, 28, 64, 67, some slides with multiple specimens)— same data; 1 phoretic deutonymph—same data,
<date id="FFCC1051FFD8FF877FF14894FC8EA92A" box="[690,842,488,515]" pageId="7" pageNumber="448" value="2017-05-28">28 May 2017</date>
; 11 phoretic deutonymphs—same data,
<date id="FFCC1051FFD8FF877848489BFA5CA92A" box="[1291,1432,487,515]" pageId="7" pageNumber="448" value="2017-06-05">05 Jun 2017</date>
.
</paragraph>
<paragraph id="8BCD3691FFD8FF877D844B70FD45A962" blockId="7.[151,1437,343,1127]" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877D844B70FE4EA90E" bold="true" box="[199,394,523,550]" pageId="7" pageNumber="448">Type deposition.</emphasis>
<materialsCitation id="3B1A3CCCFFD8FF877CD34B70FD45A962" ID-GBIF-Occurrence="2413160329" collectionCode="ZISP, UMMZ, TSUMZ" pageId="7" pageNumber="448" specimenCount="42" typeStatus="holotype">
<typeStatus id="54C98833FFD8FF877CD34B70FE3CA90E" box="[400,504,524,550]" pageId="7" pageNumber="448" type="holotype">Holotype</typeStatus>
and
<specimenCount id="9D74FD18FFD8FF877F6F4B70FD64A90F" box="[556,672,524,551]" pageId="7" pageNumber="448" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(slide 28) deposited in
<collectionCode id="ED63AE54FFD8FF877EE04B70FC24A90E" box="[931,992,524,550]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="448">ZISP</collectionCode>
;
<specimenCount id="9D74FD18FFD8FF877EA84B70FBBCA90F" box="[1003,1144,524,551]" pageId="7" pageNumber="448" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
(slides 13, 18) in
<collectionCode id="ED63AE54FFD8FF87787E4B71FA5CA90F" box="[1341,1432,525,551]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="7" pageNumber="448" type="University or college">UMMZ</collectionCode>
;
<specimenCount id="9D74FD18FFD8FF877DD44B4CFEE1A963" box="[151,293,560,587]" pageId="7" pageNumber="448" type="generic" typeStatus="paratypes">31 paratypes</specimenCount>
(remaining slides) in
<collectionCode id="ED63AE54FFD8FF877F584B4CFDBAA962" box="[539,638,560,586]" pageId="7" pageNumber="448">TSUMZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCD3691FFD8FF877D844B2FFCBFA80A" blockId="7.[151,1437,343,1127]" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877D844B2FFE09A946" bold="true" box="[199,461,595,622]" pageId="7" pageNumber="448">Phoretic deutonymph</emphasis>
(
<tableCitation id="C6F0032AFFD8FF877C9D4B28FDFCA946" box="[478,568,596,622]" captionStart="TABLE 1" captionStartId="2.[151,244,728,755]" captionTargetBox="[163,1396,770,1874]" captionTargetPageId="2" captionText="TABLE 1. Measurements of phoretic deutonymphs of N. sidorchukae, sp. n." httpUri="http://table.plazi.org/id/DF0D6619FFDDFF827DD44BA4FC35A9DB" pageId="7" pageNumber="448" tableUuid="DF0D6619FFDDFF827DD44BA4FC35A9DB">Table 1</tableCitation>
: measurements). Body broadly ovoid, widest in sejugal region (
<figureCitation id="13492A14FFD8FF87785C4B28FA4AA946" box="[1311,1422,596,622]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1797,1822]" captionStartId-1="4.[151,250,1983,2008]" captionStartId-2="5.[151,250,1945,1970]" captionStartId-3="6.[151,250,1983,2008]" captionTargetBox-0="[168,1383,184,1771]" captionTargetBox-1="[172,1414,181,1957]" captionTargetBox-2="[168,1419,189,1920]" captionTargetBox-3="[173,1415,181,1957]" captionTargetId-0="figure@3.[151,1435,181,1771]" captionTargetId-1="figure@4.[172,1414,181,1957]" captionTargetId-2="figure@5.[168,1419,181,1920]" captionTargetId-3="figure@6.[173,1415,181,1957]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Dorsal view, photo (DIC)." captionText-1="FIGURE 2. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Dorsal view, line drawing." captionText-2="FIGURE 3. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Ventral view, photo (DIC)." captionText-3="FIGURE 4. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Ventral view, line drawing." httpUri-0="https://zenodo.org/record/3353294/files/figure.png" httpUri-1="https://zenodo.org/record/3353296/files/figure.png" httpUri-2="https://zenodo.org/record/3353298/files/figure.png" httpUri-3="https://zenodo.org/record/3353300/files/figure.png" pageId="7" pageNumber="448">Figs. 1–4</figureCitation>
); idiosomal length 147–167 (158±8, n=10), width 107–131 (117±7, n=10). Gnathosoma consisting of relatively elongate subcapitular remnant and very short, rounded palps bearing palpal solenidia (ω) apically and filiform dorsal palpal setae
<emphasis id="B906EA83FFD8FF877C614BBCFE92A9F2" box="[290,342,704,730]" italics="true" pageId="7" pageNumber="448">(dm)</emphasis>
dorsolaterally; supracoxal setae of palps (
<emphasis id="B906EA83FFD8FF877E734BBCFCA2A9F2" box="[816,870,704,730]" italics="true" pageId="7" pageNumber="448">elc p</emphasis>
) absent from subcapitular remnant, their positions marked by somewhat refractile spots (
<figureCitation id="13492A14FFD8FF877F014B98FD55A9D6" box="[578,657,740,766]" captionStart="FIGURE 2" captionStartId="4.[151,250,1983,2008]" captionTargetBox="[172,1414,181,1957]" captionTargetId="figure@4.[172,1414,181,1957]" captionTargetPageId="4" captionText="FIGURE 2. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Dorsal view, line drawing." httpUri="https://zenodo.org/record/3353296/files/figure.png" pageId="7" pageNumber="448">Figs. 2</figureCitation>
,
<figureCitation id="13492A14FFD8FF877FDE4B98FD6AA9D6" box="[669,686,740,766]" captionStart="FIGURE 4" captionStartId="6.[151,250,1983,2008]" captionTargetBox="[173,1415,181,1957]" captionTargetId="figure@6.[173,1415,181,1957]" captionTargetPageId="6" captionText="FIGURE 4. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Ventral view, line drawing." httpUri="https://zenodo.org/record/3353300/files/figure.png" pageId="7" pageNumber="448">4</figureCitation>
). Positions of ventral subcapitulars and ventral palpal tibials on the gnathosomal remnants also marked by refractile spots (
<figureCitation id="13492A14FFD8FF877E414A74FC95A80A" box="[770,849,776,802]" captionStart="FIGURE 2" captionStartId="4.[151,250,1983,2008]" captionTargetBox="[172,1414,181,1957]" captionTargetId="figure@4.[172,1414,181,1957]" captionTargetPageId="4" captionText="FIGURE 2. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Dorsal view, line drawing." httpUri="https://zenodo.org/record/3353296/files/figure.png" pageId="7" pageNumber="448">Figs. 2</figureCitation>
,
<figureCitation id="13492A14FFD8FF877E1E4A74FCAAA80A" box="[861,878,776,802]" captionStart="FIGURE 4" captionStartId="6.[151,250,1983,2008]" captionTargetBox="[173,1415,181,1957]" captionTargetId="figure@6.[173,1415,181,1957]" captionTargetPageId="6" captionText="FIGURE 4. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Ventral view, line drawing." httpUri="https://zenodo.org/record/3353300/files/figure.png" pageId="7" pageNumber="448">4</figureCitation>
).
</paragraph>
<paragraph id="8BCD3691FFD8FF877D844A51FD99AF4F" blockId="7.[151,1437,343,1127]" pageId="7" pageNumber="448">
Dorsum (
<figureCitation id="13492A14FFD8FF877C724A50FEBAA86E" box="[305,382,812,838]" captionStart="FIGURE 1" captionStartId="3.[151,250,1797,1822]" captionTargetBox="[168,1383,184,1771]" captionTargetId="figure@3.[151,1435,181,1771]" captionTargetPageId="3" captionText="FIGURE 1. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Dorsal view, photo (DIC)." httpUri="https://zenodo.org/record/3353294/files/figure.png" pageId="7" pageNumber="448">Figs. 1</figureCitation>
,
<figureCitation id="13492A14FFD8FF877CCF4A50FE58A86E" box="[396,412,812,838]" captionStart="FIGURE 2" captionStartId="4.[151,250,1983,2008]" captionTargetBox="[172,1414,181,1957]" captionTargetId="figure@4.[172,1414,181,1957]" captionTargetPageId="4" captionText="FIGURE 2. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Dorsal view, line drawing." httpUri="https://zenodo.org/record/3353296/files/figure.png" pageId="7" pageNumber="448">2</figureCitation>
,
<figureCitation id="13492A14FFD8FF877CEB4A57FE09A86E" box="[424,461,811,838]" captionStart="FIGURE 5" captionStartId="7.[151,250,1975,2000]" captionTargetBox="[158,1430,1148,1950]" captionTargetId="figure@7.[158,1430,1148,1950]" captionTargetPageId="7" captionText="FIGURE 5. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Propodosoma, dorsal (A) and ventral (B) views, photo (DIC)." httpUri="https://zenodo.org/record/3353302/files/figure.png" pageId="7" pageNumber="448">5A</figureCitation>
,
<figureCitation id="13492A14FFD8FF877C9A4A50FE3AA86F" box="[473,510,812,839]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1051]" captionTargetBox="[151,1435,181,1001]" captionTargetId="figure@8.[151,1435,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Hysterosoma, dorsal (A) and ventral (B) views, photo (DIC)." httpUri="https://zenodo.org/record/3353304/files/figure.png" pageId="7" pageNumber="448">6A</figureCitation>
). Propodosoma and hysterosoma each covered by heavily punctate shield. Internal vertical setae (
<emphasis id="B906EA83FFD8FF877C7D4A2DFE96A842" box="[318,338,849,874]" italics="true" pageId="7" pageNumber="448">vi</emphasis>
) short, positioned slightly anterior to apex of propodosomal sclerite, with adjacent bases; external vertical setae (
<emphasis id="B906EA83FFD8FF877C244A09FE44A8A6" box="[359,384,885,910]" italics="true" pageId="7" pageNumber="448">ve</emphasis>
) absent; internal scapular setae (
<emphasis id="B906EA83FFD8FF877FB64A09FCCCA8A6" box="[757,776,885,910]" italics="true" pageId="7" pageNumber="448">si</emphasis>
) distinctly posterior and medial to external scapulars (
<emphasis id="B906EA83FFD8FF8778374A09FA4FA8A6" box="[1396,1419,885,910]" italics="true" pageId="7" pageNumber="448">se</emphasis>
); supracoxal setae of legs I (
<emphasis id="B906EA83FFD8FF877C894AE5FE2AA89A" box="[458,494,921,946]" italics="true" pageId="7" pageNumber="448">scx</emphasis>
) filiform, positioned on thin sclerites above legs I. Sejugal furrow well developed. Hysterosoma with 11 pairs of simple, filiform setae on hysterosomal shield (
<emphasis id="B906EA83FFD8FF8779454AC1FBD6A8FE" box="[1030,1042,957,982]" italics="true" pageId="7" pageNumber="448">c</emphasis>
<subScript id="17F634D4FFD8FF8779504AB2FBDFA8F4" attach="right" box="[1043,1051,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779504AB2FBDFA8F4" box="[1043,1051,974,988]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8779684AC1FBF3A8FE" box="[1067,1079,957,982]" italics="true" pageId="7" pageNumber="448">c</emphasis>
<subScript id="17F634D4FFD8FF8779744AB2FBFBA8F4" attach="left" box="[1079,1087,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779744AB2FBFBA8F4" box="[1079,1087,974,988]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8779134AC1FB98A8FE" box="[1104,1116,957,982]" italics="true" pageId="7" pageNumber="448">c</emphasis>
<subScript id="17F634D4FFD8FF87791F4AB2FBA0A8F4" attach="left" box="[1116,1124,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF87791F4AB2FBA0A8F4" box="[1116,1124,974,988]" italics="true" pageId="7" pageNumber="448">p</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8779374AC0FB46A8FE" box="[1140,1154,956,982]" italics="true" pageId="7" pageNumber="448">d</emphasis>
<subScript id="17F634D4FFD8FF8779C14AB2FB4EA8F4" attach="left" box="[1154,1162,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779C14AB2FB4EA8F4" box="[1154,1162,974,988]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8779D94AC0FB6CA8FE" box="[1178,1192,956,982]" italics="true" pageId="7" pageNumber="448">d</emphasis>
<subScript id="17F634D4FFD8FF8779EB4AB2FB74A8F4" attach="left" box="[1192,1200,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779EB4AB2FB74A8F4" box="[1192,1200,974,988]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8779834AC1FB08A8FE" box="[1216,1228,957,982]" italics="true" pageId="7" pageNumber="448">e</emphasis>
<subScript id="17F634D4FFD8FF87798E4AB2FB11A8F4" attach="right" box="[1229,1237,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF87798E4AB2FB11A8F4" box="[1229,1237,974,988]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8779A64AC1FB35A8FE" box="[1253,1265,957,982]" italics="true" pageId="7" pageNumber="448">e</emphasis>
<subScript id="17F634D4FFD8FF8779B24AB2FB3DA8F4" attach="left" box="[1265,1273,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779B24AB2FB3DA8F4" box="[1265,1273,974,988]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8778494AC0FAD6A8FE" box="[1290,1298,956,982]" italics="true" pageId="7" pageNumber="448">f</emphasis>
<subScript id="17F634D4FFD8FF8778524AB2FADDA8F4" attach="left" box="[1297,1305,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8778524AB2FADDA8F4" box="[1297,1305,974,988]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF87786A4AC0FAF3A8FE" box="[1321,1335,956,982]" italics="true" pageId="7" pageNumber="448">h</emphasis>
<subScript id="17F634D4FFD8FF8778744AB2FAFBA8F4" attach="left" box="[1335,1343,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8778744AB2FAFBA8F4" box="[1335,1343,974,988]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8778134AC0FA9AA8FE" box="[1360,1374,956,982]" italics="true" pageId="7" pageNumber="448">h</emphasis>
<subScript id="17F634D4FFD8FF87781D4AB2FAA2A8F4" attach="left" box="[1374,1382,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF87781D4AB2FAA2A8F4" box="[1374,1382,974,988]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF8778354AC0FA40A8FE" box="[1398,1412,956,982]" italics="true" pageId="7" pageNumber="448">h</emphasis>
<subScript id="17F634D4FFD8FF8778C74AB2FA48A8F4" attach="left" box="[1412,1420,974,988]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8778C74AB2FA48A8F4" box="[1412,1420,974,988]" italics="true" pageId="7" pageNumber="448">3</emphasis>
</subScript>
), setae
<emphasis id="B906EA83FFD8FF877D954A9CFF20A8D2" box="[214,228,992,1018]" italics="true" pageId="7" pageNumber="448">h</emphasis>
<subScript id="17F634D4FFD8FF877DA74A8EFF28AF28" attach="left" box="[228,236,1010,1024]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877DA74A8EFF28AF28" box="[228,236,1010,1024]" italics="true" pageId="7" pageNumber="448">3</emphasis>
</subScript>
distinctly longer than others. Distance
<emphasis id="B906EA83FFD8FF877FE74A9CFD76A8D2" box="[676,690,992,1018]" italics="true" pageId="7" pageNumber="448">d</emphasis>
<emphasis id="B906EA83FFD8FF877FF14A8EFD0BA8D2" box="[690,719,993,1024]" italics="true" pageId="7" pageNumber="448">
<subScript id="17F634D4FFD8FF877FF14A8EFD7EAF28" attach="left" box="[690,698,1010,1024]" fontSize="6" pageId="7" pageNumber="448">1</subScript>
-c
</emphasis>
<date id="FFCC1051FFD8FF877F934A8EFCE4A8D2" box="[720,800,992,1024]" pageId="7" pageNumber="448" value="1919-10-01">
<subScript id="17F634D4FFD8FF877F934A8EFD1CAF28" attach="left" box="[720,728,1010,1024]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877F934A8EFD1CAF28" box="[720,728,1010,1024]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
10-19
</date>
(14±3, n=10),
<emphasis id="B906EA83FFD8FF877E8A4A9CFC13A8D2" box="[969,983,992,1018]" italics="true" pageId="7" pageNumber="448">d</emphasis>
<emphasis id="B906EA83FFD8FF877E944A8EFC3DA8D2" box="[983,1017,993,1024]" italics="true" pageId="7" pageNumber="448">
<subScript id="17F634D4FFD8FF877E944A8EFC1BAF28" attach="left" box="[983,991,1010,1024]" fontSize="6" pageId="7" pageNumber="448">1</subScript>
–e
</emphasis>
<date id="FFCC1051FFD8FF877EB94A8EFB8BA8D2" box="[1018,1103,992,1024]" pageId="7" pageNumber="448" value="1938-01-26">
<subScript id="17F634D4FFD8FF877EB94A8EFBC6AF28" attach="left" box="[1018,1026,1010,1024]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877EB94A8EFBC6AF28" box="[1018,1026,1010,1024]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
26–38
</date>
(31±3, n=10); ratio
<emphasis id="B906EA83FFD8FF8778704A9CFA85A8D2" box="[1331,1345,992,1018]" italics="true" pageId="7" pageNumber="448">d</emphasis>
<emphasis id="B906EA83FFD8FF8778024A8EFA9AA8D2" box="[1345,1374,993,1024]" italics="true" pageId="7" pageNumber="448">
<subScript id="17F634D4FFD8FF8778024A8EFA8DAF28" attach="left" box="[1345,1353,1010,1024]" fontSize="6" pageId="7" pageNumber="448">1</subScript>
-c
</emphasis>
<subScript id="17F634D4FFD8FF87781C4A8EFAA3AF28" attach="left" box="[1375,1383,1010,1024]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF87781C4A8EFAA3AF28" box="[1375,1383,1010,1024]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
/
<emphasis id="B906EA83FFD8FF87783E4A9CFA4FA8D2" box="[1405,1419,992,1018]" italics="true" pageId="7" pageNumber="448">d</emphasis>
<emphasis id="B906EA83FFD8FF8778C84A8EFF67AF36" italics="true" pageId="7" pageNumber="448">
<subScript id="17F634D4FFD8FF8778C84A8EFA57AF28" attach="left" box="[1419,1427,1010,1024]" fontSize="6" pageId="7" pageNumber="448">1</subScript>
- e
</emphasis>
<subScript id="17F634D4FFD8FF877DE74D6AFF68AF0C" attach="left" box="[164,172,1046,1060]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877DE74D6AFF68AF0C" box="[164,172,1046,1060]" italics="true" pageId="7" pageNumber="448">1</emphasis>
</subScript>
1.57–3.57 (2.34±0.58, n=10). Opisthonotal gland openings (gla) between setae
<emphasis id="B906EA83FFD8FF87796F4D79FBFCAF36" box="[1068,1080,1029,1054]" italics="true" pageId="7" pageNumber="448">c</emphasis>
<subScript id="17F634D4FFD8FF87797B4D6AFB84AF0C" attach="left" box="[1080,1088,1046,1060]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF87797B4D6AFB84AF0C" box="[1080,1088,1046,1060]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
and
<emphasis id="B906EA83FFD8FF8779344D79FB47AF36" box="[1143,1155,1029,1054]" italics="true" pageId="7" pageNumber="448">e</emphasis>
<subScript id="17F634D4FFD8FF8779C04D6AFB4FAF0C" attach="left" box="[1155,1163,1046,1060]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779C04D6AFB4FAF0C" box="[1155,1163,1046,1060]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
, situated much closer to latter. Of four fundamental pairs of cupules, only two pairs observed:
<emphasis id="B906EA83FFD8FF877EE84D55FC05AF6A" box="[939,961,1065,1090]" italics="true" pageId="7" pageNumber="448">ia</emphasis>
between setae
<emphasis id="B906EA83FFD8FF87792E4D55FBBDAF6A" box="[1133,1145,1065,1090]" italics="true" pageId="7" pageNumber="448">c</emphasis>
<subScript id="17F634D4FFD8FF8779394D46FB46AF60" attach="left" box="[1146,1154,1082,1096]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779394D46FB46AF60" box="[1146,1154,1082,1096]" italics="true" pageId="7" pageNumber="448">2</emphasis>
</subScript>
and
<emphasis id="B906EA83FFD8FF8779FA4D55FB01AF6A" box="[1209,1221,1065,1090]" italics="true" pageId="7" pageNumber="448">c</emphasis>
<subScript id="17F634D4FFD8FF8779854D46FB0AAF60" attach="right" box="[1222,1230,1082,1096]" fontSize="6" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF8779854D46FB0AAF60" box="[1222,1230,1082,1096]" italics="true" pageId="7" pageNumber="448">p</emphasis>
</subScript>
,
<emphasis id="B906EA83FFD8FF87799E4D54FB37AF6A" box="[1245,1267,1064,1090]" italics="true" pageId="7" pageNumber="448">ih</emphasis>
ventral, lateral to posterior portion of attachment organ.
</paragraph>
<caption id="DF0D6619FFD8FF877DD44ECBFEDFACDC" httpUri="https://zenodo.org/record/3353302/files/figure.png" pageId="7" pageNumber="448" startId="7.[151,250,1975,2000]" targetBox="[158,1430,1148,1950]" targetPageId="7">
<paragraph id="8BCD3691FFD8FF877DD44ECBFEDFACDC" blockId="7.[151,1436,1975,2036]" pageId="7" pageNumber="448">
<emphasis id="B906EA83FFD8FF877DD44ECBFED7ACF8" bold="true" box="[151,275,1975,2000]" pageId="7" pageNumber="448">FIGURE 5.</emphasis>
<taxonomicName id="4C724D12FFD8FF877C594EC4FDDBACF8" authority="Klimov &amp; Oconnor &amp; Khaustov, 2019" authorityName="Klimov &amp; Oconnor &amp; Khaustov" authorityYear="2019" box="[282,543,1976,2000]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="448" phylum="Arthropoda" rank="species" species="sidorchukae" status="sp. nov.">
<emphasis id="B906EA83FFD8FF877C594EC4FDDBACF8" box="[282,543,1976,2000]" italics="true" pageId="7" pageNumber="448">Naiadacarus sidorchukae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F8FFD8FF877F664EC4FD99ACE7" box="[549,605,1976,1999]" pageId="7" pageNumber="448" rank="species">sp. n.</taxonomicNameLabel>
, phoretic deutonymph, holotype. Propodosoma, dorsal (A) and ventral (B) views, photo (DIC).
</paragraph>
</caption>
<caption id="DF0D6619FFD7FF887DD44D7EFEDFAF17" httpUri="https://zenodo.org/record/3353304/files/figure.png" pageId="8" pageNumber="449" startId="8.[151,250,1026,1051]" targetBox="[151,1435,181,1001]" targetPageId="8">
<paragraph id="8BCD3691FFD7FF887DD44D7EFEDFAF17" blockId="8.[151,1436,1026,1087]" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF887DD44D7EFED0AF33" bold="true" box="[151,276,1026,1051]" pageId="8" pageNumber="449">FIGURE 6.</emphasis>
<taxonomicName id="4C724D12FFD7FF887C584D7FFDE4AF33" authority="Klimov &amp; Oconnor &amp; Khaustov, 2019" authorityName="Klimov &amp; Oconnor &amp; Khaustov" authorityYear="2019" box="[283,544,1027,1051]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="8" pageNumber="449" phylum="Arthropoda" rank="species" species="sidorchukae" status="sp. nov.">
<emphasis id="B906EA83FFD7FF887C584D7FFDE4AF33" box="[283,544,1027,1051]" italics="true" pageId="8" pageNumber="449">Naiadacarus sidorchukae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F8FFD7FF887F644D7FFD9BAF32" box="[551,607,1027,1050]" pageId="8" pageNumber="449" rank="species">sp. n.</taxonomicNameLabel>
, phoretic deutonymph, holotype. Hysterosoma, dorsal (A) and ventral (B) views, photo (DIC).
</paragraph>
</caption>
<paragraph id="8BCD3691FFD7FF887D844DEEFB43AC38" blockId="8.[151,1437,1168,2024]" pageId="8" pageNumber="449">
Venter (
<figureCitation id="13492A14FFD7FF887C634DEDFEABAF83" box="[288,367,1169,1195]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[168,1419,189,1920]" captionTargetId="figure@5.[168,1419,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Ventral view, photo (DIC)." httpUri="https://zenodo.org/record/3353298/files/figure.png" pageId="8" pageNumber="449">Figs. 3</figureCitation>
,
<figureCitation id="13492A14FFD7FF887C3E4DEDFE49AF83" box="[381,397,1169,1195]" captionStart="FIGURE 4" captionStartId="6.[151,250,1983,2008]" captionTargetBox="[173,1415,181,1957]" captionTargetId="figure@6.[173,1415,181,1957]" captionTargetPageId="6" captionText="FIGURE 4. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Ventral view, line drawing." httpUri="https://zenodo.org/record/3353300/files/figure.png" pageId="8" pageNumber="449">4</figureCitation>
,
<figureCitation id="13492A14FFD7FF887CD94DECFE78AF83" box="[410,444,1168,1195]" captionStart="FIGURE 5" captionStartId="7.[151,250,1975,2000]" captionTargetBox="[158,1430,1148,1950]" captionTargetId="figure@7.[158,1430,1148,1950]" captionTargetPageId="7" captionText="FIGURE 5. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Propodosoma, dorsal (A) and ventral (B) views, photo (DIC)." httpUri="https://zenodo.org/record/3353302/files/figure.png" pageId="8" pageNumber="449">5B</figureCitation>
,
<figureCitation id="13492A14FFD7FF887C8A4DEDFE2AAF84" box="[457,494,1169,1196]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1051]" captionTargetBox="[151,1435,181,1001]" captionTargetId="figure@8.[151,1435,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Hysterosoma, dorsal (A) and ventral (B) views, photo (DIC)." httpUri="https://zenodo.org/record/3353304/files/figure.png" pageId="8" pageNumber="449">6B</figureCitation>
). Coxal fields sclerotized, covered with longitudinal wrinkles; coxal fields IV more heavily sclerotized in anterior half. Anterior apodemes of coxal fields I fused forming sternum. Anterior apodemes of coxal fields II curved medially. Posterior apodemes of coxal fields II broad, curved medially; there is substantial distance between their posterior ends and apices of anterior apodemes III; very finely granular sclerotization present in this area. Anterior apodemes of coxal fields III fused with each other and anterior apodemes of coxal fields IV. Posterior medial apodeme well developed, extending from anterior apodemes IV almost to genital opening. Posterior apodemes IV short, transverse, positioned directly anterior to attachment organ. Dorsal hysterosomal shield separated from ventral surface by a distinct suture on each side. Subhumeral setae (
<emphasis id="B906EA83FFD7FF8879754CF2FB86AE8F" box="[1078,1090,1422,1447]" italics="true" pageId="8" pageNumber="449">c</emphasis>
<subScript id="17F634D4FFD7FF8879004CE3FB8FAE85" attach="right" box="[1091,1099,1439,1453]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF8879004CE3FB8FAE85" box="[1091,1099,1439,1453]" italics="true" pageId="8" pageNumber="449">3</emphasis>
</subScript>
) short, filiform, positioned on ventral surface between legs II–III, adjacent to sutures separating dorsal shield from venter. Coxal setae
<emphasis id="B906EA83FFD7FF8878664CCEFA86AEE4" box="[1317,1346,1458,1484]" italics="true" pageId="8" pageNumber="449">1a</emphasis>
,
<emphasis id="B906EA83FFD7FF88780C4CCEFAAFAEE4" box="[1359,1387,1458,1484]" italics="true" pageId="8" pageNumber="449">3a</emphasis>
,
<emphasis id="B906EA83FFD7FF88783A4CCDFA52AEE3" box="[1401,1430,1457,1483]" italics="true" pageId="8" pageNumber="449">4b</emphasis>
,
<emphasis id="B906EA83FFD7FF887DD44CAAFF61AEC7" box="[151,165,1494,1519]" italics="true" pageId="8" pageNumber="449">g</emphasis>
filiform;
<emphasis id="B906EA83FFD7FF887C534CAAFEE8AED8" box="[272,300,1494,1520]" italics="true" pageId="8" pageNumber="449">4a</emphasis>
in form of small, rounded conoids. Genital region in posterior portion of coxal fields IV; opening elongate, with two pairs of genital papillae within genital atrium; papillae relatively large, two segmented, with rounded apices (
<figureCitation id="13492A14FFD7FF887DB34F61FE88AD10" box="[240,332,1565,1592]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1051]" captionTargetBox="[151,1435,181,1001]" captionTargetId="figure@8.[151,1435,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Hysterosoma, dorsal (A) and ventral (B) views, photo (DIC)." httpUri="https://zenodo.org/record/3353304/files/figure.png" pageId="8" pageNumber="449">Fig. 6B</figureCitation>
). Coxal setae (
<emphasis id="B906EA83FFD7FF887CBE4F61FDDDAD1F" box="[509,537,1565,1591]" italics="true" pageId="8" pageNumber="449">4b</emphasis>
) positioned at junctions of anterior coxal apodemes IV and posterior median apodeme; genital setae (
<emphasis id="B906EA83FFD7FF887CE74F3EFE76AD73" box="[420,434,1602,1627]" italics="true" pageId="8" pageNumber="449">g</emphasis>
) laterad of genital opening. Attachment organ posterior to coxal fields (
<figureCitation id="13492A14FFD7FF88798E4F3DFAD8AD73" box="[1229,1308,1601,1627]" captionStart="FIGURE 4" captionStartId="6.[151,250,1983,2008]" captionTargetBox="[173,1415,181,1957]" captionTargetId="figure@6.[173,1415,181,1957]" captionTargetPageId="6" captionText="FIGURE 4. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Ventral view, line drawing." httpUri="https://zenodo.org/record/3353300/files/figure.png" pageId="8" pageNumber="449">Figs. 4</figureCitation>
,
<figureCitation id="13492A14FFD7FF8878644F3DFA8FAD74" box="[1319,1355,1601,1628]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1051]" captionTargetBox="[151,1435,181,1001]" captionTargetId="figure@8.[151,1435,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Hysterosoma, dorsal (A) and ventral (B) views, photo (DIC)." httpUri="https://zenodo.org/record/3353304/files/figure.png" pageId="8" pageNumber="449">6B</figureCitation>
). Anterior suckers (
<emphasis id="B906EA83FFD7FF887C684F19FE83AD57" box="[299,327,1637,1663]" italics="true" pageId="8" pageNumber="449">ad</emphasis>
<subScript id="17F634D4FFD7FF887C044F0BFE8BADAD" attach="left" box="[327,335,1655,1669]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF887C044F0BFE8BADAD" box="[327,335,1655,1669]" italics="true" pageId="8" pageNumber="449">3</emphasis>
</subScript>
) round, median suckers (
<emphasis id="B906EA83FFD7FF887F2E4F19FD4DAD57" box="[621,649,1637,1663]" italics="true" pageId="8" pageNumber="449">ad</emphasis>
<subScript id="17F634D4FFD7FF887FCA4F0BFD60ADAD" attach="left" box="[649,676,1655,1669]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF887FCA4F0BFD60ADAD" box="[649,676,1655,1669]" italics="true" pageId="8" pageNumber="449">1+2</emphasis>
</subScript>
) slightly larger, consisting of large sclerotized margins surrounding paired vestigial alveoli. Pair of small refractile spots anterolateral to median suckers (
<emphasis id="B906EA83FFD7FF88792A4FF6FB46AD8B" box="[1129,1154,1674,1699]" italics="true" pageId="8" pageNumber="449">ps</emphasis>
<subScript id="17F634D4FFD7FF8879C14FE7FB4EAD81" attach="left" box="[1154,1162,1691,1705]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF8879C14FE7FB4EAD81" box="[1154,1162,1691,1705]" italics="true" pageId="8" pageNumber="449">3</emphasis>
</subScript>
). Lateral conoidal setae of attachment organ (
<emphasis id="B906EA83FFD7FF887CCA4FD2FE66ADEF" box="[393,418,1710,1735]" italics="true" pageId="8" pageNumber="449">ps</emphasis>
<subScript id="17F634D4FFD7FF887CE14FC3FE6EADE5" attach="left" box="[418,426,1727,1741]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF887CE14FC3FE6EADE5" box="[418,426,1727,1741]" italics="true" pageId="8" pageNumber="449">2</emphasis>
</subScript>
) situated slightly posterior to a line joining centers of median suckers; posterior conoidal setae (
<emphasis id="B906EA83FFD7FF887D9C4FAEFF3CADC3" box="[223,248,1746,1771]" italics="true" pageId="8" pageNumber="449">ps</emphasis>
<subScript id="17F634D4FFD7FF887DBB4F9FFEC4ADD9" attach="left" box="[248,256,1763,1777]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF887DBB4F9FFEC4ADD9" box="[248,256,1763,1777]" italics="true" pageId="8" pageNumber="449">1</emphasis>
</subScript>
) posterior to median suckers (
<emphasis id="B906EA83FFD7FF887F174FADFDB4ADC3" box="[596,624,1745,1771]" italics="true" pageId="8" pageNumber="449">ad</emphasis>
<subScript id="17F634D4FFD7FF887F334F9FFD4FADD9" attach="left" box="[624,651,1763,1777]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF887F334F9FFD4FADD9" box="[624,651,1763,1777]" italics="true" pageId="8" pageNumber="449">1+2</emphasis>
</subScript>
) and slightly more medial. Anterior and posterior lateral and posterior median cuticular conoids well developed. Anus positioned between anterior suckers (
<emphasis id="B906EA83FFD7FF8879104F89FBABAC27" box="[1107,1135,1781,1807]" italics="true" pageId="8" pageNumber="449">ad</emphasis>
<subScript id="17F634D4FFD7FF88792C4E7BFBB3AC3D" attach="left" box="[1135,1143,1799,1813]" fontSize="6" pageId="8" pageNumber="449">
<emphasis id="B906EA83FFD7FF88792C4E7BFBB3AC3D" box="[1135,1143,1799,1813]" italics="true" pageId="8" pageNumber="449">3</emphasis>
</subScript>
).
</paragraph>
<paragraph id="8BCD3691FFD7FF897D844E66FD05AADF" blockId="8.[151,1437,1168,2024]" lastBlockId="9.[151,1437,153,503]" lastPageId="9" lastPageNumber="450" pageId="8" pageNumber="449">
Legs (
<figureCitation id="13492A14FFD7FF887C4D4E65FE4EAC1B" box="[270,394,1817,1843]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="9.[151,250,1556,1581]" captionStartId-1="10.[151,250,1218,1243]" captionStartId-2="12.[151,250,1489,1514]" captionStartId-3="13.[151,250,1290,1315]" captionTargetBox-0="[151,1435,525,1531]" captionTargetBox-1="[151,1436,181,1193]" captionTargetBox-2="[154,1435,535,1461]" captionTargetBox-3="[151,1436,345,1265]" captionTargetId-0="figure@9.[151,1435,525,1531]" captionTargetId-1="figure@10.[151,1436,181,1193]" captionTargetId-2="figure@12.[151,1435,525,1464]" captionTargetId-3="figure@13.[151,1436,345,1265]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 7. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg I, dorsal (A), ventral view (B); leg II, dorsal (C), ventral view (D). Photo (DIC)." captionText-1="FIGURE 8. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg I, dorsal (A), ventral view (B); leg II, dorsal (C), ventral view (D). Line drawing." captionText-2="FIGURE 9. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg III, dorsal (A), ventral view (B); leg IV, dorsal (C), ventral view (D). Photo (DIC)." captionText-3="FIGURE 10. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg III, dorsal (A), ventral view (B); leg IV, dorsal (C), ventral view (D). Line drawing." httpUri-0="https://zenodo.org/record/3353306/files/figure.png" httpUri-1="https://zenodo.org/record/3353308/files/figure.png" httpUri-2="https://zenodo.org/record/3353310/files/figure.png" httpUri-3="https://zenodo.org/record/3353312/files/figure.png" pageId="8" pageNumber="449">Figs. 7–10</figureCitation>
). Legs elongate, all segments free. Trochanters I–III each with filiform seta (
<emphasis id="B906EA83FFD7FF8879AE4E66FAC8AC1B" box="[1261,1292,1818,1843]" italics="true" pageId="8" pageNumber="449">pR</emphasis>
I–II,
<emphasis id="B906EA83FFD7FF88780F4E66FAACAC1B" box="[1356,1384,1818,1843]" italics="true" pageId="8" pageNumber="449">sR</emphasis>
III). Femoral setation 1-1-0-1; setae
<emphasis id="B906EA83FFD7FF887F444E42FDE1AC7F" box="[519,549,1854,1879]" italics="true" pageId="8" pageNumber="449">vF</emphasis>
I–II and
<emphasis id="B906EA83FFD7FF887FD74E42FD7CAC7F" box="[660,696,1854,1879]" italics="true" pageId="8" pageNumber="449">wF</emphasis>
IV filiform, smooth. Genual setation 2-2-0-0; setae
<emphasis id="B906EA83FFD7FF8878544E42FAFBAC70" box="[1303,1343,1854,1880]" italics="true" pageId="8" pageNumber="449">mG</emphasis>
and
<emphasis id="B906EA83FFD7FF88783F4E42FA59AC70" box="[1404,1437,1854,1880]" italics="true" pageId="8" pageNumber="449">cG</emphasis>
I–II somewhat spine-like, seta
<emphasis id="B906EA83FFD7FF887CB34E1EFDD6AC54" box="[496,530,1890,1916]" italics="true" pageId="8" pageNumber="449">nG</emphasis>
III absent. Tibial setation 2-2-1-1; setae
<emphasis id="B906EA83FFD7FF887E9F4E1EFC3EAC53" box="[988,1018,1890,1915]" italics="true" pageId="8" pageNumber="449">gT</emphasis>
and
<emphasis id="B906EA83FFD7FF8879734E1DFB8AAC53" box="[1072,1102,1889,1915]" italics="true" pageId="8" pageNumber="449">hT</emphasis>
I–II and
<emphasis id="B906EA83FFD7FF8879F64E1DFB15AC53" box="[1205,1233,1889,1915]" italics="true" pageId="8" pageNumber="449">kT</emphasis>
III–IV spine-like. Tarsal setation 8-9-8-8; tarsus I with 5 thinly foliate setae (
<emphasis id="B906EA83FFD7FF887E7B4EFAFC95ACB7" box="[824,849,1926,1951]" italics="true" pageId="8" pageNumber="449">ra</emphasis>
,
<emphasis id="B906EA83FFD7FF887E234EF9FCB2ACB7" box="[864,886,1925,1951]" italics="true" pageId="8" pageNumber="449">la</emphasis>
,
<emphasis id="B906EA83FFD7FF887EC54EFAFC50ACB7" box="[902,916,1926,1951]" italics="true" pageId="8" pageNumber="449">p</emphasis>
,
<emphasis id="B906EA83FFD7FF887EE04EFAFC75ACB7" box="[931,945,1926,1951]" italics="true" pageId="8" pageNumber="449">q</emphasis>
, and
<emphasis id="B906EA83FFD7FF887EB14EF9FC3EACB7" box="[1010,1018,1925,1951]" italics="true" pageId="8" pageNumber="449">f</emphasis>
), seta
<emphasis id="B906EA83FFD7FF8879054EF9FB90ACB7" box="[1094,1108,1925,1951]" italics="true" pageId="8" pageNumber="449">d</emphasis>
elongate and filiform, seta
<emphasis id="B906EA83FFD7FF8878D34EFAFA58ACB7" box="[1424,1436,1926,1951]" italics="true" pageId="8" pageNumber="449">e</emphasis>
spoon-shaped, seta
<emphasis id="B906EA83FFD7FF887C324ED6FE56ACEB" box="[369,402,1962,1987]" italics="true" pageId="8" pageNumber="449">wa</emphasis>
filiform to somewhat spine-like, setae
<emphasis id="B906EA83FFD7FF887E074ED6FCA4ACEB" box="[836,864,1962,1987]" italics="true" pageId="8" pageNumber="449">aa</emphasis>
and
<emphasis id="B906EA83FFD7FF887ED64ED5FC75ACEB" box="[917,945,1961,1987]" italics="true" pageId="8" pageNumber="449">ba</emphasis>
absent (
<figureCitation id="13492A14FFD7FF88794D4ED5FB9EACEC" box="[1038,1114,1961,1988]" captionStart="FIGURE 7" captionStartId="9.[151,250,1556,1581]" captionTargetBox="[151,1435,525,1531]" captionTargetId="figure@9.[151,1435,525,1531]" captionTargetPageId="9" captionText="FIGURE 7. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg I, dorsal (A), ventral view (B); leg II, dorsal (C), ventral view (D). Photo (DIC)." httpUri="https://zenodo.org/record/3353306/files/figure.png" pageId="8" pageNumber="449">Figs. 7</figureCitation>
A-B, 8A-B); tarsus II similar to tarsus I except seta
<emphasis id="B906EA83FFD7FF887CCF4EB1FE6CACCF" box="[396,424,1997,2023]" italics="true" pageId="8" pageNumber="449">ba</emphasis>
present and filiform and spoon-shaped apex of seta
<emphasis id="B906EA83FFD7FF887EAF4EB2FC3CACCF" box="[1004,1016,1998,2023]" italics="true" pageId="8" pageNumber="449">e</emphasis>
virtually absent (
<figureCitation id="13492A14FFD7FF8879F94EB1FAC2ACCF" box="[1210,1286,1997,2023]" captionStart="FIGURE 7" captionStartId="9.[151,250,1556,1581]" captionTargetBox="[151,1435,525,1531]" captionTargetId="figure@9.[151,1435,525,1531]" captionTargetPageId="9" captionText="FIGURE 7. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg I, dorsal (A), ventral view (B); leg II, dorsal (C), ventral view (D). Photo (DIC)." httpUri="https://zenodo.org/record/3353306/files/figure.png" pageId="8" pageNumber="449">Figs. 7</figureCitation>
C–D, 8C–D). Tarsus III with 8 setae (
<emphasis id="B906EA83FFD6FF897CDF49E6FE6BAB9B" box="[412,431,154,179]" italics="true" pageId="9" pageNumber="450">w</emphasis>
,
<emphasis id="B906EA83FFD6FF897CFF49E6FE03AB9B" box="[444,455,154,179]" italics="true" pageId="9" pageNumber="450">r</emphasis>
,
<emphasis id="B906EA83FFD6FF897C9749E6FE1BAB9B" box="[468,479,154,179]" italics="true" pageId="9" pageNumber="450">s</emphasis>
,
<emphasis id="B906EA83FFD6FF897CAF49E6FE3EAB9B" box="[492,506,154,179]" italics="true" pageId="9" pageNumber="450">p</emphasis>
,
<emphasis id="B906EA83FFD6FF897F4449E6FDD1AB9B" box="[519,533,154,179]" italics="true" pageId="9" pageNumber="450">q</emphasis>
,
<emphasis id="B906EA83FFD6FF897F6049E6FDEBAB9B" box="[547,559,154,179]" italics="true" pageId="9" pageNumber="450">e</emphasis>
,
<emphasis id="B906EA83FFD6FF897F7F49E5FD80AB9B" box="[572,580,153,179]" italics="true" pageId="9" pageNumber="450">f</emphasis>
,
<emphasis id="B906EA83FFD6FF897F1249E5FD9BAB9B" box="[593,607,153,179]" italics="true" pageId="9" pageNumber="450">d</emphasis>
) smooth, setae
<emphasis id="B906EA83FFD6FF897E4E49E6FCDDAB9B" box="[781,793,154,179]" italics="true" pageId="9" pageNumber="450">e</emphasis>
,
<emphasis id="B906EA83FFD6FF897E6549E5FCEAAB9B" box="[806,814,153,179]" italics="true" pageId="9" pageNumber="450">f</emphasis>
, and
<emphasis id="B906EA83FFD6FF897E2949E6FCB9AB9B" box="[874,893,154,179]" italics="true" pageId="9" pageNumber="450">w</emphasis>
slightly flattened apically (
<figureCitation id="13492A14FFD6FF8979EF49E5FB3DAB9B" box="[1196,1273,153,179]" captionStart="FIGURE 9" captionStartId="12.[151,250,1489,1514]" captionTargetBox="[154,1435,535,1461]" captionTargetId="figure@12.[151,1435,525,1464]" captionTargetPageId="12" captionText="FIGURE 9. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg III, dorsal (A), ventral view (B); leg IV, dorsal (C), ventral view (D). Photo (DIC)." httpUri="https://zenodo.org/record/3353310/files/figure.png" pageId="9" pageNumber="450">Figs. 9</figureCitation>
A-B, 10 A–B). Tarsus IV similar to tarsus III except seta
<emphasis id="B906EA83FFD6FF897F3249C2FD40ABFF" box="[625,644,190,215]" italics="true" pageId="9" pageNumber="450">w</emphasis>
longer, thicker and distinctly barbed, and setae
<emphasis id="B906EA83FFD6FF8979E249C1FB6BABFF" box="[1185,1199,189,215]" italics="true" pageId="9" pageNumber="450">d</emphasis>
and
<emphasis id="B906EA83FFD6FF8979A449C1FB2BABFF" box="[1255,1263,189,215]" italics="true" pageId="9" pageNumber="450">f</emphasis>
longer (
<figureCitation id="13492A14FFD6FF89780D49C1FA58ABFF" box="[1358,1436,189,215]" captionStart="FIGURE 9" captionStartId="12.[151,250,1489,1514]" captionTargetBox="[154,1435,535,1461]" captionTargetId="figure@12.[151,1435,525,1464]" captionTargetPageId="12" captionText="FIGURE 9. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg III, dorsal (A), ventral view (B); leg IV, dorsal (C), ventral view (D). Photo (DIC)." httpUri="https://zenodo.org/record/3353310/files/figure.png" pageId="9" pageNumber="450">Figs. 9</figureCitation>
C–D, 10 C–D). Solenidia ω
<subScript id="17F634D4FFD6FF897C8D498EFE12AA28" attach="left" box="[462,470,242,256]" fontSize="6" pageId="9" pageNumber="450">1</subScript>
on tarsi I–II cylindrical, with rounded apices; ω
<subScript id="17F634D4FFD6FF897EB7498EFC38AA28" attach="left" box="[1012,1020,242,256]" fontSize="6" pageId="9" pageNumber="450">3</subScript>
on tarsus I shorter than ω
<subScript id="17F634D4FFD6FF89785C498EFAE3AA28" attach="left" box="[1311,1319,242,256]" fontSize="6" pageId="9" pageNumber="450">1</subScript>
, with bul- bous apex, positioned directly adjacent to ω
<subScript id="17F634D4FFD6FF897FC6486AFD49AA0C" attach="left" box="[645,653,278,292]" fontSize="6" pageId="9" pageNumber="450">1</subScript>
; ω
<subScript id="17F634D4FFD6FF897FEC486AFD73AA0C" attach="left" box="[687,695,278,292]" fontSize="6" pageId="9" pageNumber="450">2</subScript>
of tarsus I thin, expanding slightly apically, positioned somewhat more basal and posterior to ω
<subScript id="17F634D4FFD6FF897CA54846FE2AAA60" attach="left" box="[486,494,314,328]" fontSize="6" pageId="9" pageNumber="450">1</subScript>
+ ω
<subScript id="17F634D4FFD6FF897F594846FDE6AA60" attach="left" box="[538,546,314,328]" fontSize="6" pageId="9" pageNumber="450">3</subScript>
group (in the
<typeStatus id="54C98833FFD6FF897F824855FCE2AA6B" box="[705,806,297,323]" pageId="9" pageNumber="450" type="holotype">holotype</typeStatus>
, ω
<subScript id="17F634D4FFD6FF897E074846FC88AA60" attach="left" box="[836,844,314,328]" fontSize="6" pageId="9" pageNumber="450">2</subScript>
on the same level as ω
<subScript id="17F634D4FFD6FF8979124846FB9DAA60" attach="left" box="[1105,1113,314,328]" fontSize="6" pageId="9" pageNumber="450">1</subScript>
+ ω
<subScript id="17F634D4FFD6FF8979C74846FB48AA60" attach="left" box="[1156,1164,314,328]" fontSize="6" pageId="9" pageNumber="450">3</subScript>
group); φ of tibiae I–III elongate, tapering; φ
<date id="FFCC1051FFD6FF897CC94831FE2EAA4F" box="[394,490,333,359]" pageId="9" pageNumber="450" value="1932-02-20">II 20–32</date>
(27±3, n=23), shorter than tarsus II, ratio tibial solenidion φ II/tarsus II: 0.62–0.93 (0.79±0.08, n=23); φ IV short and stout; σ of genu I elongate, tapering slightly, σ of genu II shorter, cylindrical, σ of genu III absent (when absent, position marked by refractile spot) or present (short). Famulus ε of tarsus I short, pointed, adjacent to solenidion ω
<subScript id="17F634D4FFD6FF897F4648B6FDC9AAF0" attach="left" box="[517,525,458,472]" fontSize="6" pageId="9" pageNumber="450">1</subScript>
. All pretarsi consisting of hooked empodial claws arising from tarsal apices, short paired condylophores visible within tarsal apices.
</paragraph>
<caption id="DF0D6619FFD6FF897DD44F68FE3BAD79" httpUri="https://zenodo.org/record/3353306/files/figure.png" pageId="9" pageNumber="450" startId="9.[151,250,1556,1581]" targetBox="[151,1435,525,1531]" targetPageId="9">
<paragraph id="8BCD3691FFD6FF897DD44F68FE3BAD79" blockId="9.[151,1436,1556,1617]" pageId="9" pageNumber="450">
<emphasis id="B906EA83FFD6FF897DD44F68FED7AD05" bold="true" box="[151,275,1556,1581]" pageId="9" pageNumber="450">FIGURE 7.</emphasis>
<taxonomicName id="4C724D12FFD6FF897C5A4F69FDDAAD05" authority="Klimov &amp; Oconnor &amp; Khaustov, 2019" authorityName="Klimov &amp; Oconnor &amp; Khaustov" authorityYear="2019" box="[281,542,1557,1581]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="sidorchukae" status="sp. nov.">
<emphasis id="B906EA83FFD6FF897C5A4F69FDDAAD05" box="[281,542,1557,1581]" italics="true" pageId="9" pageNumber="450">Naiadacarus sidorchukae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F8FFD6FF897F674F69FD98AD04" box="[548,604,1557,1580]" pageId="9" pageNumber="450" rank="species">sp. n.</taxonomicNameLabel>
, phoretic deutonymph, holotype. Leg I, dorsal (A), ventral view (B); leg II, dorsal (C), ventral view (D). Photo (DIC).
</paragraph>
</caption>
<paragraph id="8BCD3691FFD6FF897D844FDEFBA0AC01" blockId="9.[151,1437,1698,1942]" pageId="9" pageNumber="450">
<emphasis id="B906EA83FFD6FF897D844FDEFE86AD95" bold="true" box="[199,322,1698,1725]" pageId="9" pageNumber="450">Diagnosis.</emphasis>
Phoretic deutonymphs of
<taxonomicName id="4C724D12FFD6FF897F304FDFFC51AD95" authorityName="Klimov &amp; Oconnor &amp; Khaustov" authorityYear="2019" box="[627,917,1699,1725]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="sidorchukae">
<emphasis id="B906EA83FFD6FF897F304FDFFC51AD95" box="[627,917,1699,1725]" italics="true" pageId="9" pageNumber="450">Naiadacarus sidorchukae</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4C724D12FFD6FF8979CF4FD8FACDAD95" box="[1164,1289,1699,1725]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="fashingi">
<emphasis id="B906EA83FFD6FF8979CF4FD8FACDAD95" box="[1164,1289,1699,1725]" italics="true" pageId="9" pageNumber="450">N. fashingi</emphasis>
</taxonomicName>
by a shorter tibial solenidion φ II, 20–32 (27±3, n=23); this solenidion is shorter than tarsus II, ratio tibial solenidion φ II/tarsus II is 0.62–0.93 (0.79±0.08, n=23) (
<figureCitation id="13492A14FFD6FF897F5B4F97FDBDAC2D" box="[536,633,1771,1797]" captionStart="FIGURE 7" captionStartId="9.[151,250,1556,1581]" captionTargetBox="[151,1435,525,1531]" captionTargetId="figure@9.[151,1435,525,1531]" captionTargetPageId="9" captionText="FIGURE 7. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg I, dorsal (A), ventral view (B); leg II, dorsal (C), ventral view (D). Photo (DIC)." httpUri="https://zenodo.org/record/3353306/files/figure.png" pageId="9" pageNumber="450">Figs. 7C</figureCitation>
,
<figureCitation id="13492A14FFD6FF897FC74F97FD6CAC2D" box="[644,680,1771,1797]" captionStart="FIGURE 8" captionStartId="10.[151,250,1218,1243]" captionTargetBox="[151,1436,181,1193]" captionTargetId="figure@10.[151,1436,181,1193]" captionTargetPageId="10" captionText="FIGURE 8. Naiadacarus sidorchukae sp. n., phoretic deutonymph, holotype. Leg I, dorsal (A), ventral view (B); leg II, dorsal (C), ventral view (D). Line drawing." httpUri="https://zenodo.org/record/3353308/files/figure.png" pageId="9" pageNumber="450">8C</figureCitation>
). In
<taxonomicName id="4C724D12FFD6FF897F9A4F90FC97AC2D" box="[729,851,1771,1797]" class="Arachnida" family="Acaridae" genus="Naiadacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="fashingi">
<emphasis id="B906EA83FFD6FF897F9A4F90FC97AC2D" box="[729,851,1771,1797]" italics="true" pageId="9" pageNumber="450">N. fashingi</emphasis>
</taxonomicName>
, tibial solenidion φ II is longer, 35–49 (42±4, n=23), as long as tarsus II, ratio tibial solenidion φ II/tarsus II is 0.94–1.17 (1.02±0.05, n=23).
</paragraph>
<paragraph id="8BCD3691FFD6FF897D844E4FFD4EACBE" blockId="9.[151,1437,1698,1942]" pageId="9" pageNumber="450">
<emphasis id="B906EA83FFD6FF897D844E4FFE8AAC65" bold="true" box="[199,334,1843,1869]" pageId="9" pageNumber="450">Etymology.</emphasis>
The new species is named after the late Ekaterina (Katya) Sidorchuk (
<date id="FFCC1051FFD6FF89793E4E4FFB3FAC65" box="[1149,1275,1843,1869]" pageId="9" pageNumber="450" value="1981" valueMax="2019">1981-2019</date>
), who made a major contribution in mite paleontology by introducing breakthrough techniques in studying fossil specimens and describing a large number of important taxa.
</paragraph>
</subSubSection>
</treatment>
</document>
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