treatments-xml/data/03/AC/87/03AC87EC816FFFC2B4C5FE3DFC50FC90.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.5643881" ID-GBIF-Dataset="2d26eac2-e73d-493c-a17d-5f71cdece746" ID-GBIF-Taxon="189980073" ID-Zenodo-Dep="5643881" approvedBy="valdenar" checkinTime="1635965709029" checkinUser="valdenar" docAuthor="Kaltenbach, Thomas, Surbakti, Suriani, Kluge, Nikita J., Gattolliat, JeanJLuc, Sartori, Michel &amp; Balke, Michael" docDate="2021" docId="03AC87EC816FFFC2B4C5FE3DFC50FC90" docLanguage="en" docName="Treubia.48.1.37-54.pdf" docOrigin="Treubia 48 (1)" docTitle="Labiobaetis academicus Kaltenbach, Surbakti &amp; Kluge 2021, sp. nov." docType="treatment" docVersion="3" lastPageId="7" lastPageNumber="44" masterDocId="FF95FF94816CFFC5B415FFB6FFEDFFA6" masterDocTitle="Discovery Of A New Mayfly Species (Ephemeroptera, Baetidae) Near Cenderawasih University Campus In Papua, Indonesia" masterLastPageNumber="54" masterPageNumber="37" pageId="3" pageNumber="40" updateTime="1635968351674" updateUser="ExternalLinkService">
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<mods:title>Discovery Of A New Mayfly Species (Ephemeroptera, Baetidae) Near Cenderawasih University Campus In Papua, Indonesia</mods:title>
</mods:titleInfo>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaltenbach, Thomas</mods:namePart>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CHJ 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CHJ 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Surbakti, Suriani</mods:namePart>
<mods:affiliation>Department of Biology, Cenderawasih University (UNCEN), Jl. Kumbolker Perumnas III, Yabansai, Heram, Kota Jayapura, Papua 99224, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kluge, Nikita J.</mods:namePart>
<mods:affiliation>Department of Entomology, Biological Faculty, SaintJPetersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gattolliat, JeanJLuc</mods:namePart>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CHJ 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CHJ 1015 Lausanne, Switzerland</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sartori, Michel</mods:namePart>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CHJ 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CHJ 1015 Lausanne, Switzerland</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Balke, Michael</mods:namePart>
<mods:affiliation>SNSBJZoologische Staatssammlung München (ZSM), Münchhausenstrasse 21, DJ 81247 München, Germany</mods:affiliation>
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<mods:title>Treubia</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-06-30</mods:number>
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<mods:number>48</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>37</mods:start>
<mods:end>54</mods:end>
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<mods:identifier type="DOI">10.14203/treubia.v48i1.4020</mods:identifier>
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<subSubSection box="[208,1040,395,424]" pageId="3" pageNumber="40" type="nomenclature">
<paragraph blockId="3.[208,1040,395,424]" box="[208,1040,395,424]" pageId="3" pageNumber="40">
<emphasis bold="true" box="[208,1040,395,424]" pageId="3" pageNumber="40">
<taxonomicName authority="Kaltenbach, Surbakti &amp; Kluge" authorityName="Kaltenbach, Surbakti &amp; Kluge" authorityYear="2021" box="[208,931,395,424]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="academicus" status="sp. nov.">
<emphasis bold="true" box="[208,524,395,424]" italics="true" pageId="3" pageNumber="40">Labiobaetis academicus</emphasis>
Kaltenbach, Surbakti &amp; Kluge
</taxonomicName>
<taxonomicNameLabel box="[939,1040,395,424]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection box="[208,424,445,474]" pageId="3" pageNumber="40" type="description">
<paragraph blockId="3.[208,424,445,474]" box="[208,424,445,474]" pageId="3" pageNumber="40">
<figureCitation box="[208,362,445,474]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="8.[299,375,1785,1809]" captionStartId-1="9.[223,299,1847,1871]" captionStartId-2="10.[299,375,1296,1320]" captionTargetBox-0="[341,1378,240,1734]" captionTargetBox-1="[239,1320,231,1786]" captionTargetBox-2="[299,1435,469,1219]" captionTargetId-0="figure-12@8.[299,1438,204,1815]" captionTargetId-1="figure-10@9.[223,1358,210,1815]" captionTargetId-2="figure-12@10.[299,1435,469,1245]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." captionText-1="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." captionText-2="Figure 3. Labiobaetis academicus sp. nov., habitus, larva: a Dorsal view b Ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5643873" figureDoi-1="http://doi.org/10.5281/zenodo.5643877" figureDoi-2="http://doi.org/10.5281/zenodo.5643879" httpUri-0="https://zenodo.org/record/5643873/files/figure.png" httpUri-1="https://zenodo.org/record/5643877/files/figure.png" httpUri-2="https://zenodo.org/record/5643879/files/figure.png" pageId="3" pageNumber="40">Figures 1–3</figureCitation>
, 
<figureCitation box="[376,424,445,474]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStart-3="Figure 8" captionStartId-0="12.[299,375,1812,1836]" captionStartId-1="13.[223,299,1745,1769]" captionStartId-2="14.[299,375,964,988]" captionStartId-3="14.[299,375,1904,1928]" captionTargetBox-0="[323,1396,244,1698]" captionTargetBox-1="[258,1314,204,1701]" captionTargetBox-2="[299,1331,204,889]" captionTargetBox-3="[299,1329,1063,1836]" captionTargetId-0="figure-12@12.[299,1406,204,1769]" captionTargetId-1="figure-10@13.[226,1326,177,1731]" captionTargetId-2="figure-12@14.[299,1331,204,911]" captionTargetId-3="figure-25@14.[299,1329,1063,1852]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="14" captionText-0="Figure 5. Labiobaetis academicus sp. nov., male imago: a Habitus, dorsal view b Abdomen c Habitus, lateral view d Cercus e Genital structure f Subimaginal gonostyli extracted from mature male larva with fragment of larval protogonostylus." captionText-1="Figure 6. Labiobaetis academicus sp. nov., female imago: a, b Habitus, dorsal view; male imago: c Middle leg; subimago (same in male and female): d Mesonotum e Mesopleuron." captionText-2="Figure 7. Distribution of Labiobaetis claudiae group in New Guinea." captionText-3="Figure 8. Habitat of L. academicus sp. nov. (photo N.J. Kluge)." figureDoi-0="http://doi.org/10.5281/zenodo.5643887" figureDoi-1="http://doi.org/10.5281/zenodo.5643889" figureDoi-2="http://doi.org/10.5281/zenodo.5643891" figureDoi-3="http://doi.org/10.5281/zenodo.5643893" httpUri-0="https://zenodo.org/record/5643887/files/figure.png" httpUri-1="https://zenodo.org/record/5643889/files/figure.png" httpUri-2="https://zenodo.org/record/5643891/files/figure.png" httpUri-3="https://zenodo.org/record/5643893/files/figure.png" pageId="3" pageNumber="40">5–8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="40" type="diagnosis">
<paragraph blockId="3.[151,1361,528,760]" pageId="3" pageNumber="40">
<emphasis bold="true" box="[208,616,528,557]" pageId="3" pageNumber="40">Differential diagnosis. Larva.</emphasis>
Following combination of characters differentiate the new species from 
<taxonomicName authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2018" box="[386,529,579,608]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis box="[386,529,579,608]" italics="true" pageId="3" pageNumber="40">L. claudiae</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2018" box="[593,737,580,608]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="stagnum">
<emphasis box="[593,737,580,608]" italics="true" pageId="3" pageNumber="40">L. stagnum</emphasis>
</taxonomicName>
: A) maxilla medially with 7–9 medium to long, spineJlike setae; B) shape of labial palp segment II as 
<figureCitation box="[844,937,630,659]" captionStart="Figure 1" captionStartId="8.[299,375,1785,1809]" captionTargetBox="[341,1378,240,1734]" captionTargetId="figure-12@8.[299,1438,204,1815]" captionTargetPageId="8" captionText="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." figureDoi="http://doi.org/10.5281/zenodo.5643873" httpUri="https://zenodo.org/record/5643873/files/figure.png" pageId="3" pageNumber="40">Fig. 1h</figureCitation>
; segment III slightly pentagonal; C) shape of tergalius IV as 
<figureCitation box="[530,667,680,709]" captionStart="Figure 2" captionStartId="9.[223,299,1847,1871]" captionTargetBox="[239,1320,231,1786]" captionTargetId="figure-10@9.[223,1358,210,1815]" captionTargetPageId="9" captionText="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." figureDoi="http://doi.org/10.5281/zenodo.5643877" httpUri="https://zenodo.org/record/5643877/files/figure.png" pageId="3" pageNumber="40">Fig. 2d; F</figureCitation>
) paraproct distally not expanded, with 20–25 stout, marginal spines.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="43" pageId="3" pageNumber="40" type="description">
<paragraph blockId="3.[151,1361,806,886]" pageId="3" pageNumber="40">
<emphasis bold="true" box="[208,470,807,836]" pageId="3" pageNumber="40">Description. Larva</emphasis>
(
<figureCitation box="[489,607,806,835]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="8.[299,375,1785,1809]" captionStartId-1="9.[223,299,1847,1871]" captionStartId-2="10.[299,375,1296,1320]" captionTargetBox-0="[341,1378,240,1734]" captionTargetBox-1="[239,1320,231,1786]" captionTargetBox-2="[299,1435,469,1219]" captionTargetId-0="figure-12@8.[299,1438,204,1815]" captionTargetId-1="figure-10@9.[223,1358,210,1815]" captionTargetId-2="figure-12@10.[299,1435,469,1245]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." captionText-1="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." captionText-2="Figure 3. Labiobaetis academicus sp. nov., habitus, larva: a Dorsal view b Ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5643873" figureDoi-1="http://doi.org/10.5281/zenodo.5643877" figureDoi-2="http://doi.org/10.5281/zenodo.5643879" httpUri-0="https://zenodo.org/record/5643873/files/figure.png" httpUri-1="https://zenodo.org/record/5643877/files/figure.png" httpUri-2="https://zenodo.org/record/5643879/files/figure.png" pageId="3" pageNumber="40">Figs 1–3</figureCitation>
). Cerci ca. 2/3 of body length, paracercus ca. 2/3 of cerci length, antenna approx. twice as long as head length.
</paragraph>
<paragraph blockId="3.[151,1360,921,1103]" pageId="3" pageNumber="40">
<emphasis box="[208,356,921,950]" italics="true" pageId="3" pageNumber="40">Coloration.</emphasis>
Head, thorax and abdomen dorsally brown, with pattern as in 
<figureCitation box="[1185,1282,921,950]" captionStart="Figure 3" captionStartId="10.[299,375,1296,1320]" captionTargetBox="[299,1435,469,1219]" captionTargetId="figure-12@10.[299,1435,469,1245]" captionTargetPageId="10" captionText="Figure 3. Labiobaetis academicus sp. nov., habitus, larva: a Dorsal view b Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5643879" httpUri="https://zenodo.org/record/5643879/files/figure.png" pageId="3" pageNumber="40">Fig. 3a</figureCitation>
. Fore protoptera brown with bright striation. Head, thorax and abdomen ventrally light brown. Legs light brown; femur with a large, distomedial brown spot, dorsal margin and apex brown; tibia in basal part with brown spot, bordered by patellotibial suture. Caudalii light brown.
</paragraph>
<paragraph blockId="3.[151,1360,1138,1218]" pageId="3" pageNumber="40">
<emphasis box="[208,315,1139,1167]" italics="true" pageId="3" pageNumber="40">Antenna</emphasis>
(
<figureCitation box="[335,431,1138,1167]" captionStart="Figure 2" captionStartId="9.[223,299,1847,1871]" captionTargetBox="[239,1320,231,1786]" captionTargetId="figure-10@9.[223,1358,210,1815]" captionTargetPageId="9" captionText="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." figureDoi="http://doi.org/10.5281/zenodo.5643877" httpUri="https://zenodo.org/record/5643877/files/figure.png" pageId="3" pageNumber="40">Fig. 2h</figureCitation>
) with scape and pedicel subcylindrical, without distolateral process at scape.
</paragraph>
<paragraph blockId="3.[151,1361,1253,1536]" pageId="3" pageNumber="40">
<emphasis box="[208,309,1253,1282]" italics="true" pageId="3" pageNumber="40">Labrum</emphasis>
(
<figureCitation box="[330,427,1253,1282]" captionStart="Figure 1" captionStartId="8.[299,375,1785,1809]" captionTargetBox="[341,1378,240,1734]" captionTargetId="figure-12@8.[299,1438,204,1815]" captionTargetPageId="8" captionText="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." figureDoi="http://doi.org/10.5281/zenodo.5643873" httpUri="https://zenodo.org/record/5643873/files/figure.png" pageId="3" pageNumber="40">Fig. 1a</figureCitation>
). Rectangular, length 0.7x maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of one plus 5–7 long, simple setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid, pectinate setae; ventral surface with ca. seven short, spineJlike setae near lateral and anterolateral margin.
</paragraph>
<paragraph blockId="3.[151,1360,1571,1753]" pageId="3" pageNumber="40">
<emphasis box="[208,405,1571,1600]" italics="true" pageId="3" pageNumber="40">Right mandible</emphasis>
(
<figureCitation box="[424,552,1571,1600]" captionStart="Figure 1" captionStartId="8.[299,375,1785,1809]" captionTargetBox="[341,1378,240,1734]" captionTargetId="figure-12@8.[299,1438,204,1815]" captionTargetPageId="8" captionText="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." figureDoi="http://doi.org/10.5281/zenodo.5643873" httpUri="https://zenodo.org/record/5643873/files/figure.png" pageId="3" pageNumber="40">Fig. 1b, c</figureCitation>
). Incisor and kinetodontium fused. Incisor with four denticles; kinetodontium with four denticles, inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex. Tuft of setae at apex of mola present.
</paragraph>
<paragraph blockId="3.[151,1364,1788,1919]" pageId="3" pageNumber="40">
<emphasis box="[208,387,1788,1817]" italics="true" pageId="3" pageNumber="40">Left mandible</emphasis>
(
<figureCitation box="[408,539,1788,1817]" captionStart="Figure 1" captionStartId="8.[299,375,1785,1809]" captionTargetBox="[341,1378,240,1734]" captionTargetId="figure-12@8.[299,1438,204,1815]" captionTargetPageId="8" captionText="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." figureDoi="http://doi.org/10.5281/zenodo.5643873" httpUri="https://zenodo.org/record/5643873/files/figure.png" pageId="3" pageNumber="40">Fig. 1d, e</figureCitation>
). Incisor and kinetodontium fused. Incisor with four denticles; kinetodontium with four denticles. Prostheca robust, apically with small denticles and combJ shaped structure. Margin between prostheca and mola slightly convex, with minute denticles
</paragraph>
<paragraph blockId="4.[234,1428,202,332]" pageId="4" pageNumber="41">toward subtriangular process. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola present.</paragraph>
<paragraph blockId="4.[234,1428,367,447]" pageId="4" pageNumber="41">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
<paragraph blockId="4.[234,1428,482,664]" pageId="4" pageNumber="41">
<emphasis box="[291,713,482,511]" italics="true" pageId="4" pageNumber="41">Hypopharynx and superlinguae</emphasis>
(
<figureCitation box="[739,837,482,511]" captionStart="Figure 1" captionStartId="8.[299,375,1785,1809]" captionTargetBox="[341,1378,240,1734]" captionTargetId="figure-12@8.[299,1438,204,1815]" captionTargetPageId="8" captionText="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." figureDoi="http://doi.org/10.5281/zenodo.5643873" httpUri="https://zenodo.org/record/5643873/files/figure.png" pageId="4" pageNumber="41">Fig. 1f</figureCitation>
). Lingua approx. as long as superlinguae. Lingua longer than broad; medial tuft of stout setae well developed, short; distal half laterally expanded. Superlinguae distally rounded; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph blockId="4.[234,1428,699,1033]" pageId="4" pageNumber="41">
<emphasis box="[291,391,699,728]" italics="true" pageId="4" pageNumber="41">Maxilla</emphasis>
(
<figureCitation box="[412,508,699,728]" captionStart="Figure 1" captionStartId="8.[299,375,1785,1809]" captionTargetBox="[341,1378,240,1734]" captionTargetId="figure-12@8.[299,1438,204,1815]" captionTargetPageId="8" captionText="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." figureDoi="http://doi.org/10.5281/zenodo.5643873" httpUri="https://zenodo.org/record/5643873/files/figure.png" pageId="4" pageNumber="41">Fig. 1g</figureCitation>
). GaleaJlacinia ventrally with two simple, apical setae under canines. Inner dorsal row of setae with three dentiJsetae, distal dentiJseta toothJlike, middle and proximal dentiJsetae slender, bifid and pectinate. Medially with one bipectinate, spineJlike seta and 7–9 medium to long, simple setae. Maxillary palp approx. as long as length of galeaJlacinia; 2Jsegmented; palp segment II 1.1x length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments Iand II; apex of last segment constricted, without excavation at inner distolateral margin.
</paragraph>
<paragraph blockId="4.[234,1429,1068,1707]" pageId="4" pageNumber="41">
<emphasis box="[291,389,1068,1097]" italics="true" pageId="4" pageNumber="41">Labium</emphasis>
(
<figureCitation box="[416,556,1068,1097]" captionStart="Figure 1" captionStartId="8.[299,375,1785,1809]" captionTargetBox="[341,1378,240,1734]" captionTargetId="figure-12@8.[299,1438,204,1815]" captionTargetPageId="8" captionText="Figure 1. Labiobaetis academicus sp. nov., larva morphology: a Labrum (left, ventral view; right, dorsal view) b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx and superlinguae g Maxilla h Labium (left, ventral view; right, dorsal view) i Apex of paraglossa." figureDoi="http://doi.org/10.5281/zenodo.5643873" httpUri="https://zenodo.org/record/5643873/files/figure.png" pageId="4" pageNumber="41">Fig. 1h, i</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca. 12 short, stout, spineJlike setae increasing in length distally; apex with two long and one short, robust setae; outer margin with six spineJlike setae; ventral surface with few fine, simple, scattered setae. Paraglossa subJrectangular, curved inward; apex rounded; ventrally with three rows of long, robust, distally pectinate setae in apical area and one medium, simple seta in anteromedial area; dorsally with a row of four long, spineJlike setae near inner margin. Labial palp with segment I approx. as long as segments II and III combined. Segment Iventrally with short, fine, simple setae. Segment II with narrow, thumbJlike distomedial protuberance; distomedial protuberance 0.5x width of base of segment III; ventral surface with short, fine, simple setae; dorsally with a row of 3–6 medium, spineJlike, simple setae near outer margin. Segment III slightly pentagonal; length 0.9x width; ventrally covered with short, spineJlike, simple setae and short, fine, simple setae.
</paragraph>
<paragraph blockId="4.[291,599,1743,1772]" box="[291,599,1743,1772]" pageId="4" pageNumber="41">
<emphasis box="[291,500,1743,1772]" italics="true" pageId="4" pageNumber="41">Hind protoptera</emphasis>
absent.
</paragraph>
<paragraph blockId="4.[234,1428,1807,1938]" pageId="4" pageNumber="41">
<emphasis box="[291,351,1808,1836]" italics="true" pageId="4" pageNumber="41">Legs</emphasis>
(
<figureCitation box="[377,520,1807,1836]" captionStart="Figure 2" captionStartId="9.[223,299,1847,1871]" captionTargetBox="[239,1320,231,1786]" captionTargetId="figure-10@9.[223,1358,210,1815]" captionTargetPageId="9" captionText="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." figureDoi="http://doi.org/10.5281/zenodo.5643877" httpUri="https://zenodo.org/record/5643877/files/figure.png" pageId="4" pageNumber="41">Fig. 2a, b</figureCitation>
). Ratio of foreleg segments 1.3:1.0:0.6:0.2. 
<emphasis box="[1131,1216,1808,1836]" italics="true" pageId="4" pageNumber="41">Femur</emphasis>
. Length ca. 3x maximum width. Dorsal margin with a row of 25–32 medium, curved, spineJlike setae; length of setae 0.12x maximum width of femur. Apex rounded, with medium to short,
</paragraph>
<paragraph blockId="5.[159,1353,193,527]" pageId="5" pageNumber="42">
curved, spineJlike setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch present. 
<emphasis box="[351,426,244,273]" italics="true" pageId="5" pageNumber="42">Tibia.</emphasis>
Dorsal margin with a row of short to medium, stout, spineJlike setae. Ventral margin with a row of short, curved, spineJlike setae, on apex some longer, spineJlike setae and a tuft of fine, simple setae. Anterior and posterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 1/2 area. 
<emphasis box="[973,1067,398,426]" italics="true" pageId="5" pageNumber="42">Tarsus.</emphasis>
Dorsal margin with a row of short, spineJlike setae. Ventral margin with a row of curved, spineJlike setae. Claw with one row of 10–12 denticles; distally pointed; with 4–5 stripes; subapical setae absent.
</paragraph>
<paragraph blockId="5.[159,1353,562,795]" pageId="5" pageNumber="42">
<emphasis box="[215,336,562,591]" italics="true" pageId="5" pageNumber="42">Abdomen</emphasis>
(
<figureCitation box="[364,470,562,591]" captionStart="Figure 2" captionStartId="9.[223,299,1847,1871]" captionTargetBox="[239,1320,231,1786]" captionTargetId="figure-10@9.[223,1358,210,1815]" captionTargetPageId="9" captionText="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." figureDoi="http://doi.org/10.5281/zenodo.5643877" httpUri="https://zenodo.org/record/5643877/files/figure.png" pageId="5" pageNumber="42">Fig. 2c</figureCitation>
). Surface with dense, irregular rows of UJshaped scale bases. Posterior margin of terga II–IX with triangular spines, approx. as long as wide; spines diminished on middle of posterior margin of tergum IX behind pair of submedian setae. Posterior margin of tergum Xwith longer and narrower spines. Posterior margins of sterna VII–IX with shorter triangular spines.
</paragraph>
<paragraph blockId="5.[159,1353,830,1012]" pageId="5" pageNumber="42">
<emphasis box="[215,318,830,859]" italics="true" pageId="5" pageNumber="42">Tergalii</emphasis>
(
<figureCitation box="[343,473,830,859]" captionStart="Figure 2" captionStartId="9.[223,299,1847,1871]" captionTargetBox="[239,1320,231,1786]" captionTargetId="figure-10@9.[223,1358,210,1815]" captionTargetPageId="9" captionText="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." figureDoi="http://doi.org/10.5281/zenodo.5643877" httpUri="https://zenodo.org/record/5643877/files/figure.png" pageId="5" pageNumber="42">Fig. 2d–f</figureCitation>
). Present on segments II–VII. Margin with denticles of different sizes, intercalating both medium and long, fine simple setae. Tracheae extending from main trunk to inner and outer margins. Tergalius IV as long as length of segments Vand VI combined. Tergalius VII as long as length of segments VIII and 1/2 IX combined.
</paragraph>
<paragraph blockId="5.[159,1353,1047,1177]" pageId="5" pageNumber="42">
<emphasis box="[215,347,1048,1076]" italics="true" pageId="5" pageNumber="42">Paraproct</emphasis>
(
<figureCitation box="[367,467,1047,1076]" captionStart="Figure 2" captionStartId="9.[223,299,1847,1871]" captionTargetBox="[239,1320,231,1786]" captionTargetId="figure-10@9.[223,1358,210,1815]" captionTargetPageId="9" captionText="Figure 2. Labiobaetis academicus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Tergalius IV e, f margin of tergalius IV g Paraproct h Base of antenna." figureDoi="http://doi.org/10.5281/zenodo.5643877" httpUri="https://zenodo.org/record/5643877/files/figure.png" pageId="5" pageNumber="42">Fig. 2g</figureCitation>
). Distally not expanded, with 20–25 stout, marginal spines. Surface scattered with UJshaped scale bases and fine, simple setae. Cercotractor with numerous small, marginal spines.
</paragraph>
<paragraph blockId="5.[159,1353,1231,1514]" pageId="5" pageNumber="42">
<emphasis bold="true" box="[215,352,1232,1261]" pageId="5" pageNumber="42">Subimago</emphasis>
(both sexes; 
<figureCitation box="[535,664,1231,1260]" captionStart="Figure 6" captionStartId="13.[223,299,1745,1769]" captionTargetBox="[258,1314,204,1701]" captionTargetId="figure-10@13.[226,1326,177,1731]" captionTargetPageId="13" captionText="Figure 6. Labiobaetis academicus sp. nov., female imago: a, b Habitus, dorsal view; male imago: c Middle leg; subimago (same in male and female): d Mesonotum e Mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.5643889" httpUri="https://zenodo.org/record/5643889/files/figure.png" pageId="5" pageNumber="42">Fig. 6d, e</figureCitation>
). 
<emphasis box="[692,958,1231,1260]" italics="true" pageId="5" pageNumber="42">Cuticular coloration</emphasis>
. Pronotum brown with lighter areas. Mesonotum brown with medioparapsidal suture contrastingly colorless, other sutures darker brown (
<figureCitation box="[350,448,1333,1362]" captionStart="Figure 6" captionStartId="13.[223,299,1745,1769]" captionTargetBox="[258,1314,204,1701]" captionTargetId="figure-10@13.[226,1326,177,1731]" captionTargetPageId="13" captionText="Figure 6. Labiobaetis academicus sp. nov., female imago: a, b Habitus, dorsal view; male imago: c Middle leg; subimago (same in male and female): d Mesonotum e Mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.5643889" httpUri="https://zenodo.org/record/5643889/files/figure.png" pageId="5" pageNumber="42">Fig. 6d</figureCitation>
). Thoracic pleura with brown sclerites and colorless membranes (
<figureCitation captionStart="Figure 6" captionStartId="13.[223,299,1745,1769]" captionTargetBox="[258,1314,204,1701]" captionTargetId="figure-10@13.[226,1326,177,1731]" captionTargetPageId="13" captionText="Figure 6. Labiobaetis academicus sp. nov., female imago: a, b Habitus, dorsal view; male imago: c Middle leg; subimago (same in male and female): d Mesonotum e Mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.5643889" httpUri="https://zenodo.org/record/5643889/files/figure.png" pageId="5" pageNumber="42">Fig. 6e</figureCitation>
). Legs mostly colorless; femur colored with brown at base, outer and inner margins, with indistinct brownish spot in distal part; tibia colored with brown at base. Abdominal terga uniformly light brown, sterna lighter. Cerci uniformly colorless.
</paragraph>
<paragraph blockId="5.[215,848,1550,1579]" box="[215,848,1550,1579]" pageId="5" pageNumber="42">
<emphasis box="[215,516,1550,1579]" italics="true" pageId="5" pageNumber="42">Hypodermal coloration</emphasis>
. As in imago (see below).
</paragraph>
<paragraph blockId="5.[215,1270,1614,1643]" box="[215,1270,1614,1643]" pageId="5" pageNumber="42">
<emphasis box="[215,313,1615,1643]" italics="true" pageId="5" pageNumber="42">Texture</emphasis>
. On all legs of both sexes, all tarsomeres covered with blunt microlepides.
</paragraph>
<paragraph blockId="5.[159,1353,1697,1980]" pageId="5" pageNumber="42">
<heading bold="true" box="[215,310,1697,1726]" fontSize="12" level="2" pageId="5" pageNumber="42" reason="2">
<emphasis bold="true" box="[215,310,1697,1726]" pageId="5" pageNumber="42">Imago,</emphasis>
</heading>
<emphasis bold="true" box="[330,396,1697,1726]" pageId="5" pageNumber="42">male</emphasis>
(
<figureCitation box="[423,559,1697,1726]" captionStart="Figure 5" captionStartId="12.[299,375,1812,1836]" captionTargetBox="[323,1396,244,1698]" captionTargetId="figure-12@12.[299,1406,204,1769]" captionTargetPageId="12" captionText="Figure 5. Labiobaetis academicus sp. nov., male imago: a Habitus, dorsal view b Abdomen c Habitus, lateral view d Cercus e Genital structure f Subimaginal gonostyli extracted from mature male larva with fragment of larval protogonostylus." figureDoi="http://doi.org/10.5281/zenodo.5643887" httpUri="https://zenodo.org/record/5643887/files/figure.png" pageId="5" pageNumber="42">Fig. 5a–d</figureCitation>
, 
<figureCitation box="[582,616,1697,1726]" captionStart="Figure 6" captionStartId="13.[223,299,1745,1769]" captionTargetBox="[258,1314,204,1701]" captionTargetId="figure-10@13.[226,1326,177,1731]" captionTargetPageId="13" captionText="Figure 6. Labiobaetis academicus sp. nov., female imago: a, b Habitus, dorsal view; male imago: c Middle leg; subimago (same in male and female): d Mesonotum e Mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.5643889" httpUri="https://zenodo.org/record/5643889/files/figure.png" pageId="5" pageNumber="42">6c</figureCitation>
). Head ocher. Turbinate eyes high widened apically, orange. Thorax light brown with ocher areas. Wing membrane colorless, veins ocher; costal brace and adjacent areas with contrasting dark brown stripes. Pterostigma with several oblique veins. Hind wing absent. Legs of all pairs with similar coloration (
<figureCitation box="[1152,1249,1849,1878]" captionStart="Figure 6" captionStartId="13.[223,299,1745,1769]" captionTargetBox="[258,1314,204,1701]" captionTargetId="figure-10@13.[226,1326,177,1731]" captionTargetPageId="13" captionText="Figure 6. Labiobaetis academicus sp. nov., female imago: a, b Habitus, dorsal view; male imago: c Middle leg; subimago (same in male and female): d Mesonotum e Mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.5643889" httpUri="https://zenodo.org/record/5643889/files/figure.png" pageId="5" pageNumber="42">Fig. 6c</figureCitation>
): femur light ocher, with inner margin and apex bordered with brown, with reddish macula near apex; tibia with base and apex whitish, other part light brownish on inner side, lighter on
</paragraph>
<paragraph blockId="6.[251,1429,179,564]" pageId="6" pageNumber="43">
outer side; tarsus light brownish. Tarsus of middle and hind legs with two apical spines, on segments 1st+2nd and 3rd. Abdominal segments I–VI whitish, with peculiar maculae: each tergum II–VII with pair of lateral brown maculae on spiracles; each tergum III–VI with pair of sublateral brown maculae; each tergum III–IV with unpaired reddish stripe on posterior margin and with more or less expressed triangular macula arising from this stripe in anterior direction. Abdominal segments VII–VIII uniformly brown. Abdominal terga IX–X lighter. Each segment of cerci white at base and at apex, contrastingly brown at middle; segments with longer and shorter brown areas irregularly alternating (
<figureCitation box="[1144,1240,535,564]" captionStart="Figure 5" captionStartId="12.[299,375,1812,1836]" captionTargetBox="[323,1396,244,1698]" captionTargetId="figure-12@12.[299,1406,204,1769]" captionTargetPageId="12" captionText="Figure 5. Labiobaetis academicus sp. nov., male imago: a Habitus, dorsal view b Abdomen c Habitus, lateral view d Cercus e Genital structure f Subimaginal gonostyli extracted from mature male larva with fragment of larval protogonostylus." figureDoi="http://doi.org/10.5281/zenodo.5643887" httpUri="https://zenodo.org/record/5643887/files/figure.png" pageId="6" pageNumber="43">Fig. 5d</figureCitation>
).
</paragraph>
<paragraph blockId="6.[251,1432,599,831]" pageId="6" pageNumber="43">
<emphasis box="[307,863,599,628]" italics="true" pageId="6" pageNumber="43">Male genital structure and development.</emphasis>
Imaginal genitals as in 
<figureCitation box="[1212,1313,599,628]" captionStart="Figure 5" captionStartId="12.[299,375,1812,1836]" captionTargetBox="[323,1396,244,1698]" captionTargetId="figure-12@12.[299,1406,204,1769]" captionTargetPageId="12" captionText="Figure 5. Labiobaetis academicus sp. nov., male imago: a Habitus, dorsal view b Abdomen c Habitus, lateral view d Cercus e Genital structure f Subimaginal gonostyli extracted from mature male larva with fragment of larval protogonostylus." figureDoi="http://doi.org/10.5281/zenodo.5643887" httpUri="https://zenodo.org/record/5643887/files/figure.png" pageId="6" pageNumber="43">Fig. 5e</figureCitation>
. SternoJ styligeral muscle completely absent. Each gonovectis with small hook at apex. Penial bridge without prominent median projection. Unistyligers cylindrical, with straight inner margins. Gonostylus with 1st segment smoothly narrowed toward apex; 2nd segment slightly widened toward apex; 3rd segment short.
</paragraph>
<paragraph blockId="6.[251,1428,866,1048]" pageId="6" pageNumber="43">
Protogonostyli of male larva represent very shallow convexities of posterior margin of abdominal sternum IX. In mature larva ready to molt to subimago, subimaginal gonostyli packed under larval cuticle in « 
<taxonomicName authority="Jpose" authorityName="Jpose" box="[657,876,968,997]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis box="[657,806,968,997]" italics="true" pageId="6" pageNumber="43">Labiobaetis</emphasis>
Jpose
</taxonomicName>
» (
<bibRefCitation author="Kluge, N. J." box="[910,1076,968,997]" journalOrPublisher="Academic Publishers, Dordrecht" pageId="6" pageNumber="43" pagination="1 - 442" refId="ref5728" refString="Kluge, N. J. 2004. The phylogenetic system of Ephemeroptera. Academic Publishers, Dordrecht, 1 - 442." title="The phylogenetic system of Ephemeroptera" type="book chapter" year="2004">Kluge, 2004</bibRefCitation>
: fig. 29I): second segments directed medially and bent (
<figureCitation box="[608,699,1019,1048]" captionStart="Figure 5" captionStartId="12.[299,375,1812,1836]" captionTargetBox="[323,1396,244,1698]" captionTargetId="figure-12@12.[299,1406,204,1769]" captionTargetPageId="12" captionText="Figure 5. Labiobaetis academicus sp. nov., male imago: a Habitus, dorsal view b Abdomen c Habitus, lateral view d Cercus e Genital structure f Subimaginal gonostyli extracted from mature male larva with fragment of larval protogonostylus." figureDoi="http://doi.org/10.5281/zenodo.5643887" httpUri="https://zenodo.org/record/5643887/files/figure.png" pageId="6" pageNumber="43">Fig. 5f</figureCitation>
).
</paragraph>
<paragraph blockId="6.[251,1432,1097,1431]" pageId="6" pageNumber="43">
<emphasis bold="true" box="[307,510,1097,1126]" pageId="6" pageNumber="43">Imago, female</emphasis>
(
<figureCitation box="[534,675,1097,1126]" captionStart="Figure 6" captionStartId="13.[223,299,1745,1769]" captionTargetBox="[258,1314,204,1701]" captionTargetId="figure-10@13.[226,1326,177,1731]" captionTargetPageId="13" captionText="Figure 6. Labiobaetis academicus sp. nov., female imago: a, b Habitus, dorsal view; male imago: c Middle leg; subimago (same in male and female): d Mesonotum e Mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.5643889" httpUri="https://zenodo.org/record/5643889/files/figure.png" pageId="6" pageNumber="43">Fig. 6a, b</figureCitation>
). Head ocher with brownish. Prothorax and anteroJ lateral areas of mesonotum with contrasting light ocher and dark brown markings; other areas of mesonotum and metanotum light brown; ventral side of thorax light uniformly ocher. Wings as in male. Leg coloration as in male. Tarsus of fore leg with two apical spines, on 2nd and 3rd segments (as on middle and hind legs). Each abdominal tergum III and IV with reddish median macula as in male; other markings of abdominal terga and sterna vary individually. Cerci as in male.
</paragraph>
<paragraph blockId="6.[251,1428,1479,1559]" pageId="6" pageNumber="43">
<emphasis bold="true" box="[307,461,1480,1509]" pageId="6" pageNumber="43">Dimension.</emphasis>
Body length of larvae 4.2–6.5 mm. Fore wing length (and approximate body length) of subimagos, male and female imagos ca. 5 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="43" type="etymology">
<paragraph blockId="6.[251,1427,1607,1687]" pageId="6" pageNumber="43">
<emphasis bold="true" box="[307,463,1608,1637]" pageId="6" pageNumber="43">Etymology.</emphasis>
Refering to the fact that the specimens were collected just behind the campus of Cenderawasih University (UNCEN) in Jayapura.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="44" pageId="6" pageNumber="43" type="distribution">
<paragraph blockId="6.[307,972,1736,1765]" box="[307,972,1736,1765]" pageId="6" pageNumber="43">
<emphasis bold="true" box="[307,482,1736,1765]" pageId="6" pageNumber="43">Distribution.</emphasis>
Indonesia: Papua Province (
<figureCitation box="[876,956,1736,1765]" captionStart="Figure 7" captionStartId="14.[299,375,964,988]" captionTargetBox="[299,1331,204,889]" captionTargetId="figure-12@14.[299,1331,204,911]" captionTargetPageId="14" captionText="Figure 7. Distribution of Labiobaetis claudiae group in New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5643891" httpUri="https://zenodo.org/record/5643891/files/figure.png" pageId="6" pageNumber="43">Fig. 7</figureCitation>
).
</paragraph>
<paragraph blockId="6.[251,1428,1813,1995]" pageId="6" pageNumber="43">
<emphasis bold="true" box="[307,566,1814,1843]" pageId="6" pageNumber="43">Biological aspects.</emphasis>
The specimens were collected at an altitude of 160 m in a medium sized, shallow, moderately flowing river with stones and mud on the bottom, with few vegetation (
<figureCitation box="[471,555,1915,1944]" captionStart="Figure 8" captionStartId="14.[299,375,1904,1928]" captionTargetBox="[299,1329,1063,1836]" captionTargetId="figure-25@14.[299,1329,1063,1852]" captionTargetPageId="14" captionText="Figure 8. Habitat of L. academicus sp. nov. (photo N.J. Kluge)." figureDoi="http://doi.org/10.5281/zenodo.5643893" httpUri="https://zenodo.org/record/5643893/files/figure.png" pageId="6" pageNumber="43">Fig. 8</figureCitation>
). The new species was collected together with 12 other mayfly species: four other, yet undescribed species of 
<taxonomicName authority="Novikova &amp; Kluge, 1987" authorityName="Novikova &amp; Kluge (Novikova &amp; Kluge" authorityYear="1987" box="[905,1423,1966,1995]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis box="[905,1054,1966,1995]" italics="true" pageId="6" pageNumber="43">Labiobaetis</emphasis>
<bibRefCitation author="Novikova, E. A. &amp; Kluge, N. J." box="[1070,1423,1966,1995]" journalOrPublisher="Vestnik Zoologii" pageId="6" pageNumber="43" pagination="8 - 19" part="1987" refId="ref5916" refString="Novikova, E. A. &amp; Kluge, N. J. 1987. Systematics of the genus Baetis (Ephemeroptera, Baetidae), with descriptions of new species from Middle Asia. Vestnik Zoologii, 1987 (4): 8 - 19 (in Russian)." title="Systematics of the genus Baetis (Ephemeroptera, Baetidae), with descriptions of new species from Middle Asia" type="journal article" year="1987">Novikova &amp; Kluge, 1987</bibRefCitation>
</taxonomicName>
;
</paragraph>
<paragraph blockId="7.[159,1353,201,332]" pageId="7" pageNumber="44">
<taxonomicName authority="McCafferty &amp; Sun, 2005" authorityName="McCafferty &amp; Sun" authorityYear="2005" box="[159,779,202,231]" class="Insecta" family="Baetidae" genus="Mystaxiops" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="venatoris">
<emphasis box="[159,435,203,231]" italics="true" pageId="7" pageNumber="44">Mystaxiops venatoris</emphasis>
McCafferty &amp; Sun, 2005
</taxonomicName>
; two species of 
<taxonomicName authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1999" box="[996,1161,201,230]" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[996,1161,201,230]" italics="true" pageId="7" pageNumber="44">Papuanatula</emphasis>
</taxonomicName>
<bibRefCitation author="LugoJOrtiz, C. R. &amp; McCafferty, W. P. &amp; Waltz, R. D." journalOrPublisher="Transactions of the American Entomological Society" pageId="7" pageNumber="44" pagination="1 - 37" part="125" refId="ref5862" refString="LugoJOrtiz, C. R., McCafferty, W. P. &amp; Waltz, R. D. 1999. Definition and reorganization of the genus Pseudocloeon (Ephemeroptera: Baetidae) with new species descriptions and combinations. Transactions of the American Entomological Society, 125: 1 - 37." title="Definition and reorganization of the genus Pseudocloeon (Ephemeroptera: Baetidae) with new species descriptions and combinations" type="journal article" year="1999">LugoJOrtiz &amp; McCafferty, 1999</bibRefCitation>
; two species of 
<taxonomicName authority="Braasch &amp; Soldan, 1980" authorityName="Braasch &amp; Soldan" authorityYear="1980" box="[599,1084,252,281]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[599,759,252,281]" italics="true" pageId="7" pageNumber="44">Centroptella</emphasis>
Braasch &amp; Soldán, 1980
</taxonomicName>
; one 
<taxonomicName box="[1152,1293,252,281]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[1152,1244,252,281]" italics="true" pageId="7" pageNumber="44">Cloeon</emphasis>
sp.
</taxonomicName>
; one 
<taxonomicName box="[159,292,304,332]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[159,248,304,332]" italics="true" pageId="7" pageNumber="44">Caenis</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName authority="Kluge, 2013" authorityName="Kluge" authorityYear="2013" box="[356,814,303,332]" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="reductus">
<emphasis box="[356,645,303,332]" italics="true" pageId="7" pageNumber="44">Nonnullidens reductus</emphasis>
Kluge, 2013
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="44" type="materials_examined">
<paragraph blockId="7.[159,1353,386,822]" pageId="7" pageNumber="44">
<emphasis bold="true" box="[215,568,387,416]" pageId="7" pageNumber="44">
TypeJmaterial. 
<materialsCitation ID-GBIF-Occurrence="3397333301" collectingDate="2019-05-26" collectionCode="MZB" collectorName="Surbakti &amp; Kellis &amp; Sumoked" country="Indonesia" elevation="160" latitude="-2.5686111" location="Waena" longLatPrecision="21" longitude="140.64749" municipality="Jayapura" pageId="7" pageNumber="44" specimenCode="GBIFCH00592382" specimenCount="1" specimenCount-larva="1" stateProvince="Papua Province" typeStatus="holotype">
<typeStatus box="[435,562,387,416]" pageId="7" pageNumber="44">Holotype</typeStatus>
. 
<collectingCountry box="[584,765,386,415]" name="Indonesia" pageId="7" pageNumber="44">INDONESIA</collectingCountry>
• 
<specimenCount box="[805,873,386,415]" pageId="7" pageNumber="44" type="larva">larva</specimenCount>
; 
<collectingRegion box="[892,1101,386,415]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="44">Papua Province</collectingRegion>
, 
<collectingMunicipality box="[1120,1237,387,415]" pageId="7" pageNumber="44">Jayapura</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03AC87EC816FFFC2B4C5FE3DFC50FC90:8EDA6021816BFFC2B0FDFE35FAA9FE39" box="[1256,1348,387,415]" country="Indonesia" latitude="-2.5686111" longLatPrecision="21" longitude="140.64749" municipality="Jayapura" name="Waena" pageId="7" pageNumber="44" stateProvince="Papua Province">Waena</location>
, 
<location LSID="urn:lsid:plazi:treatment:03AC87EC816FFFC2B4C5FE3DFC50FC90:8EDA6021816BFFC2B48AFE00FE8DFE74" box="[159,352,437,466]" country="Indonesia" latitude="-2.5686111" longLatPrecision="21" longitude="140.64749" municipality="Jayapura" name="Kamp Wolker" pageId="7" pageNumber="44" stateProvince="Papua Province">Kamp Wolker</location>
; 
<locationDeviation box="[371,668,437,466]" pageId="7" pageNumber="44">near UNCEN campus</locationDeviation>
; 
<geoCoordinate box="[687,835,437,466]" degrees="02" direction="south" minutes="34" orientation="latitude" pageId="7" pageNumber="44" precision="15" seconds="07" value="-2.5686111">02°34'07&quot;S</geoCoordinate>
, 
<geoCoordinate box="[855,1022,437,466]" degrees="140" direction="east" minutes="38" orientation="longitude" pageId="7" pageNumber="44" precision="15" seconds="51" value="140.64749">140°38'51&quot;E</geoCoordinate>
; 
<quantity box="[1042,1133,437,466]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="44" unit="m" value="160.0">
<elevation box="[1042,1133,437,466]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="44" unit="m" value="160.0">160 m</elevation>
</quantity>
; 
<date box="[1153,1285,437,466]" pageId="7" pageNumber="44" value="2019-05-26">
<collectingDate box="[1153,1285,437,466]" pageId="7" pageNumber="44" value="2019-05-26">26.v.2019</collectingDate>
</date>
; leg. 
<collectorName box="[159,271,488,517]" pageId="7" pageNumber="44">Surbakti</collectorName>
, 
<collectorName box="[291,367,488,517]" pageId="7" pageNumber="44">Kellis</collectorName>
&amp; 
<collectorName box="[425,550,488,517]" pageId="7" pageNumber="44">Sumoked</collectorName>
; (PAP080); on slide; 
<specimenCode box="[856,1106,488,517]" pageId="7" pageNumber="44">GBIFCH00592382</specimenCode>
; 
<collectionCode box="[1127,1200,489,517]" collectionName="MZB" pageId="7" pageNumber="44">MZB</collectionCode>
</materialsCitation>
</emphasis>
. 
<materialsCitation ID-GBIF-Occurrence="3397333303" collectingDate="2019-05-26" collectionCode="MZB, KSP" collectorName="Surbakti &amp; Kellis &amp; Sumoked" country="Indonesia" elevation="160" latitude="-2.5686111" location="INDONESIA" longLatPrecision="21" longitude="140.64749" municipality="Jayapura" pageId="7" pageNumber="44" specimenCode="GBIFCH00673069, GBIFCH00673081, GBIFCH00592381, GBIFCH00515503" specimenCount="17" specimenCount-larva="17" stateProvince="Papua Province" typeStatus="paratype">
<emphasis bold="true" box="[1220,1353,488,517]" pageId="7" pageNumber="44">
<typeStatus box="[1220,1347,488,517]" pageId="7" pageNumber="44">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry box="[159,333,539,568]" name="Indonesia" pageId="7" pageNumber="44">INDONESIA</collectingCountry>
• 
<specimenCount box="[379,510,539,568]" pageId="7" pageNumber="44" type="larva">17 larvae</specimenCount>
; same data as holotype; 3 on slides; GenBank 
<accessionNumber box="[1190,1348,539,568]" pageId="7" pageNumber="44">MW041241</accessionNumber>
, 
<accessionNumber box="[159,318,590,619]" pageId="7" pageNumber="44">MW041242</accessionNumber>
; 
<specimenCode box="[332,580,590,619]" pageId="7" pageNumber="44">GBIFCH00673069</specimenCode>
, 
<specimenCode box="[595,843,590,619]" pageId="7" pageNumber="44">GBIFCH00673081</specimenCode>
, 
<specimenCode box="[858,1108,590,619]" pageId="7" pageNumber="44">GBIFCH00592381</specimenCode>
; KSP, 
<collectionCode box="[1198,1271,591,619]" collectionName="MZB" pageId="7" pageNumber="44">MZB</collectionCode>
; 
<quantity box="[1286,1352,590,619]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="7" pageNumber="44" unit="in" value="14.0">14 in</quantity>
alcohol; 
<specimenCode box="[272,521,640,669]" pageId="7" pageNumber="44">GBIFCH00515503</specimenCode>
; 
<collectionCode box="[536,598,640,669]" pageId="7" pageNumber="44">KSP</collectionCode>
, 
<collectionCode box="[612,681,641,669]" collectionName="MZB" pageId="7" pageNumber="44">MZB</collectionCode>
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="3397333302" collectingDate="2012-08-09" collectingDateMax="2012-08-13" collectingDateMin="2012-08-09" collectionCode="MZB" collectorName="N. Kluge &amp; L. Sheyko" country="Indonesia" elevation="160" latitude="-2.5686111" location="Waena" longLatPrecision="21" longitude="140.64749" municipality="Jayapura" pageId="7" pageNumber="44" specimenCount="49" specimenCount-female="4" specimenCount-larva="22" specimenCount-male="15" stateProvince="Papua Province">
<specimenCount box="[712,762,640,669]" pageId="7" pageNumber="44" type="male">4 ♂</specimenCount>
and 
<specimenCount box="[828,878,640,669]" pageId="7" pageNumber="44" type="female">2 ♀</specimenCount>
imagos with individually associated larval and subimaginal exuviae, 
<specimenCount box="[579,628,691,720]" pageId="7" pageNumber="44" type="female">1 ♀</specimenCount>
subimago with its larval exuviae, 
<specimenCount box="[1076,1125,691,720]" pageId="7" pageNumber="44" type="female">1 ♀</specimenCount>
imago, 
<specimenCount box="[1232,1297,691,720]" pageId="7" pageNumber="44" type="male">11 ♂</specimenCount>
and 
<specimenCount box="[159,293,742,771]" pageId="7" pageNumber="44">8 ♀ larvae</specimenCount>
ready to molt to subimagoes, 
<specimenCount box="[680,802,742,771]" pageId="7" pageNumber="44" type="larva">22 larvae</specimenCount>
; same locality as holotype; 
<date box="[1158,1348,742,771]" pageId="7" pageNumber="44" value="2012-08-09" valueMax="2012-08-13" valueMin="2012-08-09">
<collectingDate box="[1158,1348,742,771]" pageId="7" pageNumber="44" value="2012-08-09" valueMax="2012-08-13" valueMin="2012-08-09">9–13.viii.2012</collectingDate>
</date>
; leg. 
<collectorName box="[214,508,793,822]" pageId="7" pageNumber="44">N. Kluge &amp; L. Sheyko</collectorName>
; SaintJPetersburg State University.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>