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<document id="233D8EEE63563133706C928B27DB6895" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66B86FF2DAF89FA00FD133F81" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera shiwandashanensis Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="233" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="232" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="448336E15271D6BAFF51745792B73B9A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5BC57400EA55ABA41633984CA0254D1F">
<mods:title id="7AD1EDEACA1B49E6828AB9651919B386">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="77E6B6B3C94C0DB3E1E035341362A853" type="personal">
<mods:role id="7EC7AAE8FF6A170E7885EEE22724DCF1">
<mods:roleTerm id="2D88B3E07C09C71521076A2E0AA7AE22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09DD9A5692522BE1E3C36A8C8AF6C3EF">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="2B0115805C0E9608894B04C46D0CB86D" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="AB754941C19123136582BBB7C083B3AD" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="D51EF419E51A607BAC97F32391FC8799" type="personal">
<mods:role id="019F23E849884B01DE53BD98B8CABAAB">
<mods:roleTerm id="D7DC66C86D83D66E3752CCF1B0851CAF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E70E1373DD5EDA13A32616F70C93F4FF">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="9ED972FBEF6AB6985D80921618CD95FB" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="4B424BE1777B49EA4BC94841A7CC2593" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="0A7B8CBFBFC0CE5B1BE1E062E48EB735" type="personal">
<mods:role id="A9E79B0BFE57124C5FDDA9063A706773">
<mods:roleTerm id="F6C618BD501B0DED31FC21170BB2062B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5668C9CFFA9F3CDC6739F5D2FC0A0014">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="68E7A0DB586549D1FA1D6F5709F7A4E9" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="10B3E0635BAA8A8822CA4F708ECA9BA3" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="7879AC79056182908452387B82B15271" type="personal">
<mods:role id="98355EC9874223CCF2E1CC8F0F277C5F">
<mods:roleTerm id="49736D5908F6CBB2994BD7468F983AEB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3209BA6431C70CA9AFF4FEC99D2265E8">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="ABD8B22DA9632B028A3EAB01A7B48065" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="5759B1B8BB9ED5707AD90C13EC0F4D9F" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="360BCFBD2C34090EF2B66B7E4B4FA50B" type="personal">
<mods:role id="00F5474610FB413F496E3D8DA6011909">
<mods:roleTerm id="EEE571C23A150BF7D159BAF3E2073FE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="451B749448F8DB5434BA24B6AFB7F329">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="DC798455187775779CA136CA77F2A533" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="3C7AE6538109B416CD7FB8069A3301F2" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:typeOfResource id="0C88A70CD60A5D343887884646CF6B0F">text</mods:typeOfResource>
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<mods:title id="EC233AA879C289CE31D26D2C4304A464">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="00565AC9EE589F56E65D37BFCE836719">
<mods:date id="B7B927D81C8047A5ED1651A8C0907F65">2021</mods:date>
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<mods:number id="E988B6F930A46667DB98F4870F7438A7">2021-02-03</mods:number>
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<mods:number id="4B6800D8BC4D29F91943CAC08E531451">4922</mods:number>
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<mods:classification id="E26F05D907C1FF98DDFA25165E2A5785">journal article</mods:classification>
<mods:identifier id="8A828F2E4473FC882FCDBC9F2CCCEC4B" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="839C65E728329FF74F89CC065FD4AFA6" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="40BCD1EBD8E83C39C4619ED6B8CF4C30" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
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<mods:identifier id="153339ED93A56D751C8C96281F2378AC" type="Zenodo-Dep">4496316</mods:identifier>
<mods:identifier id="8387FFE966DE9370504739B5E4E86889" type="ZooBank">D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C</mods:identifier>
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<treatment id="03BD87E66B86FF2DAF89FA00FD133F81" ID-DOI="http://doi.org/10.5281/zenodo.4534765" ID-GBIF-Taxon="177323620" ID-Zenodo-Dep="4534765" LSID="urn:lsid:plazi:treatment:03BD87E66B86FF2DAF89FA00FD133F81" httpUri="http://treatment.plazi.org/id/03BD87E66B86FF2DAF89FA00FD133F81" lastPageId="232" lastPageNumber="233" pageId="231" pageNumber="232">
<subSubSection id="C30E657B6B86FF22AF89FA00FCB13FC9" box="[151,879,618,645]" pageId="231" pageNumber="232" type="nomenclature">
<paragraph id="8BAB36F06B86FF22AF89FA00FCB13FC9" blockId="231.[151,879,618,681]" box="[151,879,618,645]" pageId="231" pageNumber="232">
<heading id="D0E3819C6B86FF22AF89FA00FCB13FC9" bold="true" box="[151,879,618,645]" fontSize="11" level="2" pageId="231" pageNumber="232" reason="4">
<emphasis id="B960EAE26B86FF22AF89FA00FCB13FC9" bold="true" box="[151,879,618,645]" pageId="231" pageNumber="232">
<taxonomicName id="4C144D736B86FF22AF89FA00FCFE3FC8" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,800,618,645]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="231" pageNumber="232" phylum="Arthropoda" rank="species" species="shiwandashanensis" status="sp. nov.">
<emphasis id="B960EAE26B86FF22AF89FA00FE213FC9" bold="true" box="[151,511,618,645]" italics="true" pageId="231" pageNumber="232">Maladera shiwandashanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B86FF22AC32FA01FCB13FC9" box="[812,879,619,645]" pageId="231" pageNumber="232" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B86FF22AF89FAE5FEB23FE5" box="[151,364,654,681]" pageId="231" pageNumber="232" type="description">
<paragraph id="8BAB36F06B86FF22AF89FAE5FEB23FE5" blockId="231.[151,879,618,681]" box="[151,364,654,681]" pageId="231" pageNumber="232">
<figureCitation id="132F2A756B86FF22AF89FAE5FED03FE5" box="[151,270,654,681]" captionStart="FIGURE 53" captionStartId="230.[151,250,1891,1915]" captionTargetBox="[178,1406,195,1858]" captionTargetId="figure-19@230.[164,1423,181,1868]" captionTargetPageId="230" captionText="FIGURE 53. A–E: Maladera bikouensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. shiwandashanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. wupingensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, C, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496452" httpUri="https://zenodo.org/record/4496452/files/figure.png" pageId="231" pageNumber="232">Figures 53</figureCitation>
F–I, 117
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B86FF22AF89FABDFF0D3EA1" pageId="231" pageNumber="232" type="materials_examined">
<paragraph id="8BAB36F06B86FF22AF89FABDFF0D3EA1" blockId="231.[151,1437,726,2013]" pageId="231" pageNumber="232">
<emphasis id="B960EAE26B86FF22AF89FABDFE1C3FBC" bold="true" box="[151,450,726,753]" pageId="231" pageNumber="232">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6B86FF22AED4FABDFE5D3E75" ID-GBIF-Occurrence="3034556842" collectingDate="2013-04-05" collectionCode="NMPC" collectorName="M. Fikacek &amp; J. Hajek &amp; J. Ruzicka" country="China" elevation="325" latitude="21.906666" location="Shiwandashan National Forest Park" longLatPrecision="125" longitude="107.903336" municipality="Shiwandashan National Forest" pageId="231" pageNumber="232" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54AF88526B86FF22AED4FABDFDE93FBD" box="[458,567,727,753]" pageId="231" pageNumber="232" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796B86FF22AD5CFABDFD893FBD" box="[578,599,727,753]" pageId="231" pageNumber="232" type="male">♂</specimenCount>
“
<collectingCountry id="F30376606B86FF22AD74FABDFD6D3FBD" box="[618,691,727,753]" name="China" pageId="231" pageNumber="232">China</collectingCountry>
: Guanxi A.R.,
<date id="FFAA10306B86FF22AC42FABCFC1B3FBD" box="[860,965,726,753]" pageId="231" pageNumber="232" value="2013-04-05">
<collectingDate id="EFEEE9D86B86FF22AC42FABCFC1B3FBD" box="[860,965,726,753]" pageId="231" pageNumber="232" value="2013-04-05">5.iv.2013</collectingDate>
</date>
<location id="8ECB602B6B86FF22ACD3FABDFABF3FBD" LSID="urn:lsid:plazi:treatment:03BD87E66B86FF2DAF89FA00FD133F81:8ECB602B6B86FF22ACD3FABDFABF3FBD" box="[973,1377,727,753]" country="China" latitude="21.906666" longLatPrecision="125" longitude="107.903336" municipality="Shiwandashan National Forest" name="Shiwandashan National Forest Park" pageId="231" pageNumber="232">
<collectingMunicipality id="6BCFAC8A6B86FF22ACD3FABDFAFB3FBD" box="[973,1317,727,753]" pageId="231" pageNumber="232">Shiwandashan National Forest</collectingMunicipality>
Park
</location>
(forested river valley, at light)
<geoCoordinate id="EE2050376B86FF22AEDCFA90FDE53E59" box="[450,571,762,789]" degrees="21" direction="north" minutes="54.4" orientation="latitude" pageId="231" pageNumber="232" precision="92" value="21.906666">21°54.4’N</geoCoordinate>
,
<geoCoordinate id="EE2050376B86FF22AD58FA90FD173E59" box="[582,713,762,789]" degrees="107" direction="east" minutes="54.2" orientation="longitude" pageId="231" pageNumber="232" precision="92" value="107.903336">107°54.2’E</geoCoordinate>
,
<quantity id="4CEC9B156B86FF22ADCAFA91FC8F3E59" box="[724,849,763,789]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="231" pageNumber="232" unit="m" value="325.0" valueMax="360.0" valueMin="290.0">
<elevation id="0039D1C36B86FF22ADCAFA91FC8F3E59" box="[724,849,763,789]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="231" pageNumber="232" unit="m" value="325.0" valueMax="360.0" valueMin="290.0">290-360 m</elevation>
</quantity>
,
<collectorName id="26E153266B86FF22AC42FA91FC013E59" box="[860,991,763,789]" pageId="231" pageNumber="232">M. Fikáček</collectorName>
,
<collectorName id="26E153266B86FF22ACF4FA91FB963E59" box="[1002,1096,763,789]" pageId="231" pageNumber="232">J. Hájek</collectorName>
,
<collectorName id="26E153266B86FF22AB4DFA91FB163E59" box="[1107,1224,762,789]" pageId="231" pageNumber="232">J. Růžicka</collectorName>
leg./ 1079
<taxonomicName id="4C144D736B86FF22AA5BFA91FA423E59" box="[1349,1436,763,789]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="231" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356B86FF22AE3BFB75FEA53E75" box="[293,379,799,825]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="231" pageNumber="232">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6B86FF22AE93FB75FAF13E11" ID-GBIF-Occurrence="3034555384" collectingDate="2013-04-05" collectionCode="ZFMK" collectorName="M. Fikacek &amp; J. Hajek &amp; J. Ruzicka" country="China" elevation="325" latitude="21.906666" location="Shiwandashan National Forest" longLatPrecision="125" longitude="107.903336" pageId="231" pageNumber="232" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54AF88526B86FF22AE93FB75FE203E75" box="[397,510,799,825]" pageId="231" pageNumber="232" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D12FD796B86FF22AD17FB75FDED3E75" box="[521,563,799,825]" pageId="231" pageNumber="232" type="male">1 ♂</specimenCount>
“
<collectingCountry id="F30376606B86FF22AD5AFB75FD533E75" box="[580,653,799,825]" name="China" pageId="231" pageNumber="232">China</collectingCountry>
: Guanxi A.R.,
<date id="FFAA10306B86FF22AC2DFB74FC453E75" box="[819,923,798,825]" pageId="231" pageNumber="232" value="2013-04-05">
<collectingDate id="EFEEE9D86B86FF22AC2DFB74FC453E75" box="[819,923,798,825]" pageId="231" pageNumber="232" value="2013-04-05">5.iv.2013</collectingDate>
</date>
<location id="8ECB602B6B86FF22ACBCFB75FB263E75" LSID="urn:lsid:plazi:treatment:03BD87E66B86FF2DAF89FA00FD133F81:8ECB602B6B86FF22ACBCFB75FB263E75" box="[930,1272,799,825]" country="China" latitude="21.906666" longLatPrecision="125" longitude="107.903336" name="Shiwandashan National Forest" pageId="231" pageNumber="232">Shiwandashan National Forest</location>
Park (forested river valley, at light)
<geoCoordinate id="EE2050376B86FF22AE9CFB28FE253E11" box="[386,507,834,861]" degrees="21" direction="north" minutes="54.4" orientation="latitude" pageId="231" pageNumber="232" precision="92" value="21.906666">21°54.4’N</geoCoordinate>
,
<geoCoordinate id="EE2050376B86FF22AD18FB28FD573E11" box="[518,649,834,861]" degrees="107" direction="east" minutes="54.2" orientation="longitude" pageId="231" pageNumber="232" precision="92" value="107.903336">107°54.2’E</geoCoordinate>
,
<quantity id="4CEC9B156B86FF22AD8BFB29FCCF3E11" box="[661,785,835,861]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="231" pageNumber="232" unit="m" value="325.0" valueMax="360.0" valueMin="290.0">
<elevation id="0039D1C36B86FF22AD8BFB29FCCF3E11" box="[661,785,835,861]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="231" pageNumber="232" unit="m" value="325.0" valueMax="360.0" valueMin="290.0">290-360 m</elevation>
</quantity>
,
<collectorName id="26E153266B86FF22AC03FB29FC7E3E11" box="[797,928,835,861]" pageId="231" pageNumber="232">M. Fikáček</collectorName>
,
<collectorName id="26E153266B86FF22ACB4FB29FBD73E11" box="[938,1033,835,861]" pageId="231" pageNumber="232">J. Hájek</collectorName>
,
<collectorName id="26E153266B86FF22AB0AFB29FB573E11" box="[1044,1161,834,861]" pageId="231" pageNumber="232">J. Růžicka</collectorName>
leg.” (
<collectionCode id="ED05AE356B86FF22ABCAFB29FAF63E11" box="[1236,1320,835,861]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="231" pageNumber="232" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6B86FF22AA24FB29FB073ECD" ID-GBIF-Occurrence="3034556255" collectingDate="1998-05-01" collectingDateMax="1998-05-07" collectingDateMin="1998-05-01" collectionCode="ZFMK" collectorName="Y. Arita" country="Vietnam" location="Asia" pageId="231" pageNumber="232" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D12FD796B86FF22AA24FB29FABA3E11" box="[1338,1380,835,861]" pageId="231" pageNumber="232" type="male">1 ♂</specimenCount>
“N.-
<collectingCountry id="F30376606B86FF22AF89FB0DFF283ECD" box="[151,246,871,897]" name="Vietnam" pageId="231" pageNumber="232">Vietnam</collectingCountry>
Vinh Phu-Prov., Tam Dao
<date id="FFAA10306B86FF22AD3BFB0DFD1B3ECD" box="[549,709,871,897]" pageId="231" pageNumber="232" value="1998-05-01" valueMax="1998-05-07" valueMin="1998-05-01">
<collectingDate id="EFEEE9D86B86FF22AD3BFB0DFD1B3ECD" box="[549,709,871,897]" pageId="231" pageNumber="232" value="1998-05-01" valueMax="1998-05-07" valueMin="1998-05-01">01.-07.V.1998</collectingDate>
</date>
<collectorName id="26E153266B86FF22ADD5FB0DFCFC3ECD" box="[715,802,871,897]" pageId="231" pageNumber="232">Y. Arita</collectorName>
leg./ 375
<taxonomicName id="4C144D736B86FF22AC8CFB0DFC373ECD" box="[914,1001,871,897]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="231" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
<location id="8ECB602B6B86FF22ACF1FB0DFBFC3ECD" LSID="urn:lsid:plazi:treatment:03BD87E66B86FF2DAF89FA00FD133F81:8ECB602B6B86FF22ACF1FB0DFBFC3ECD" box="[1007,1058,871,897]" country="Vietnam" name="Asia" pageId="231" pageNumber="232">Asia</location>
spec.” (
<collectionCode id="ED05AE356B86FF22AB60FB0DFB0C3ECD" box="[1150,1234,871,897]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="231" pageNumber="232" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6B86FF22ABFBFB0DFB0D3EE9" ID-GBIF-Occurrence="3034555345" collectionCode="ZFMK" collectorName="V. Vit. Kuban" country="Vietnam" location="Vinh Phu" municipality="Tam Dao" pageId="231" pageNumber="232" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D12FD796B86FF22ABFBFB0DFAD13ECD" box="[1253,1295,871,897]" pageId="231" pageNumber="232" type="male">1 ♂</specimenCount>
“N
<collectingCountry id="F30376606B86FF22AA20FB0DFA433ECD" box="[1342,1437,871,897]" name="Vietnam" pageId="231" pageNumber="232">Vietnam</collectingCountry>
(Tonkin) pr.
<location id="8ECB602B6B86FF22AE3DFBE1FE533EE9" LSID="urn:lsid:plazi:treatment:03BD87E66B86FF2DAF89FA00FD133F81:8ECB602B6B86FF22AE3DFBE1FE533EE9" box="[291,397,907,933]" country="Vietnam" municipality="Tam Dao" name="Vinh Phu" pageId="231" pageNumber="232">Vinh Phu</location>
1990
<collectingMunicipality id="6BCFAC8A6B86FF22AECCFBE1FDE43EE9" box="[466,570,907,933]" pageId="231" pageNumber="232">Tam Dao</collectingMunicipality>
6.-9.
<collectorName id="26E153266B86FF22AD6EFBE1FCD33EE9" box="[624,781,907,933]" pageId="231" pageNumber="232">V. Vit. Kuban</collectorName>
leg./ 1050
<taxonomicName id="4C144D736B86FF22AC92FBE1FC3D3EE9" box="[908,995,907,933]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="231" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356B86FF22AB69FBE1FB153EE9" box="[1143,1227,907,933]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="231" pageNumber="232" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6B86FF22ABC0FBE1FF0E3EA1" ID-GBIF-Occurrence="3034556437" collectingDate="1996-05-09" collectingDateMax="1996-05-19" collectingDateMin="1996-05-09" collectionCode="CP" country="Vietnam" elevation="1200" latitude="21.27" location="Pacholatko" longLatPrecision="758" longitude="105.39" pageId="231" pageNumber="232" specimenCount="1" specimenCount-male="1" stateProvince="Ha Noi" typeStatus="paratype">
<specimenCount id="9D12FD796B86FF22ABC0FBE1FAD63EE9" box="[1246,1288,907,933]" pageId="231" pageNumber="232" type="male">1 ♂</specimenCount>
“N
<collectingCountry id="F30376606B86FF22AA28FBE1FA463EE9" box="[1334,1432,907,933]" name="Vietnam" pageId="231" pageNumber="232">Vietnam</collectingCountry>
,
<geoCoordinate id="EE2050376B86FF22AF89FBC5FF343E85" box="[151,234,943,969]" degrees="21.27" direction="north" orientation="latitude" pageId="231" pageNumber="232" precision="555" value="21.27">21.27N</geoCoordinate>
<geoCoordinate id="EE2050376B86FF22AFEAFBC4FE8C3E85" box="[244,338,942,969]" degrees="105.39" direction="east" orientation="longitude" pageId="231" pageNumber="232" precision="555" value="105.39">105.39E</geoCoordinate>
<locationDeviation id="9A192FD26B86FF22AE42FBC5FDF33E85" box="[348,557,943,969]" location="Hanoi" pageId="231" pageNumber="232">
<quantity id="4CEC9B156B86FF22AE42FBC5FE7B3E85" box="[348,421,943,969]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="231" pageNumber="232" unit="km" value="70.0">70 km</quantity>
NW
<collectingRegion id="49D0F8126B86FF22AEF8FBC5FDF33E85" box="[486,557,943,969]" country="Vietnam" name="Ha Noi" pageId="231" pageNumber="232">Hanoi</collectingRegion>
</locationDeviation>
, Tam Dao
<date id="FFAA10306B86FF22ADB0FBC5FCE33E85" box="[686,829,943,969]" pageId="231" pageNumber="232" value="1996-05-09" valueMax="1996-05-19" valueMin="1996-05-09">
<collectingDate id="EFEEE9D86B86FF22ADB0FBC5FCE33E85" box="[686,829,943,969]" pageId="231" pageNumber="232" value="1996-05-09" valueMax="1996-05-19" valueMin="1996-05-09">9.-19.v.1996</collectingDate>
</date>
, 900-
<quantity id="4CEC9B156B86FF22AC60FBC5FC083E85" box="[894,982,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="231" pageNumber="232" unit="m" value="1200.0">
<elevation id="0039D1C36B86FF22AC60FBC5FC083E85" box="[894,982,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="231" pageNumber="232" unit="m" value="1200.0">1200 m</elevation>
</quantity>
<location id="8ECB602B6B86FF22ACC1FBC5FB853E85" LSID="urn:lsid:plazi:treatment:03BD87E66B86FF2DAF89FA00FD133F81:8ECB602B6B86FF22ACC1FBC5FB853E85" box="[991,1115,943,969]" country="Vietnam" latitude="21.27" longLatPrecision="758" longitude="105.39" name="Pacholatko" pageId="231" pageNumber="232" stateProvince="Ha Noi">Pacholátko</location>
&amp; Dembicky leg./ TS 112” (
<collectionCode id="ED05AE356B86FF22AFBEFBB9FF193EA1" box="[160,199,979,1005]" pageId="231" pageNumber="232">CP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B86FF2DAFD9FB9CFCB73CB9" lastPageId="232" lastPageNumber="233" pageId="231" pageNumber="232" type="description">
<paragraph id="8BAB36F06B86FF22AFD9FB9CFB593979" blockId="231.[151,1437,726,2013]" pageId="231" pageNumber="232">
<emphasis id="B960EAE26B86FF22AFD9FB9CFE87395C" bold="true" box="[199,345,1014,1040]" pageId="231" pageNumber="232">Description.</emphasis>
Length:
<quantity id="4CEC9B156B86FF22AEA2FB9DFDFF395D" box="[444,545,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="231" pageNumber="232" unit="mm" value="10.8">10.8 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156B86FF22ADFDFB9DFCE4395D" box="[739,826,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="231" pageNumber="232" unit="mm" value="8.1">8.1 mm</quantity>
, width:
<quantity id="4CEC9B156B86FF22AC8DFB9DFC35395D" box="[915,1003,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="231" pageNumber="232" unit="mm" value="7.2">7.2 mm</quantity>
. Body wide, oval, dark brown, antenna yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous.
</paragraph>
<paragraph id="8BAB36F06B86FF22AFD9FC55FEAD38D1" blockId="231.[151,1437,726,2013]" pageId="231" pageNumber="232">
Labroclypeus wide, lateral margins weakly convex and moderately convergent, lateral margins produce with ocular canthus a blunt angle, anterior angles convex, anterior margin shallowly emarginate medially, margins weakly reflexed; surface moderately shiny, at base narrowly dull, surface weakly convex, finely and densely punctate, distance between punctures smaller their diameter, with numerous long erect setae; smooth area in front of eyes 2.5 times as wide as long; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, with one or two short terminal setae. Eyes moderately large, ratio diameter/ interocular width: 0.59. Frons with fine and moderately dense punctures, with a few single setae beside eyes and behind frontoclypeal suture. Antenna with ten antennomeres, club with three antennomeres and straight, slightly longer than remaining antennomeres combined.
<collectingRegion id="49D0F8126B86FF22AB29FD35FB483835" box="[1079,1174,1375,1401]" country="Myanmar" name="Chin" pageId="231" pageNumber="232">Mentum</collectingRegion>
convexly elevated and flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06B86FF22AFD9FDCDFCD13B1D" blockId="231.[151,1437,726,2013]" pageId="231" pageNumber="232">Pronotum widest at base, lateral margins evenly convex and convergent anteriorly, and also posteriorly, anterior angles sharp and distinctly produced, posterior angles moderately rounded; anterior margin convex, with fine complete marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum, impunctate on basal midline.</paragraph>
<paragraph id="8BAB36F06B86FF22AFD9FE31FDA03BAD" blockId="231.[151,1437,726,2013]" pageId="231" pageNumber="232">Elytra convex, widest at posterior third, striae distinctly impressed, finely punctate, intervals convex, finely and densely punctate, punctures concentrated along striae, with minute setae in punctures, odd intervals with a few short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes.</paragraph>
<paragraph id="8BAB36F06B86FF22AFD9FE81FBBF3AD9" blockId="231.[151,1437,726,2013]" pageId="231" pageNumber="232">Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Distance between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.76. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta; penultimate sternite medially retracted, ultimate sternite 1.5 times as long as penultimate. Pygidium strongly convex, coarsely and densely punctate, on apex with a few long setae.</paragraph>
<paragraph id="8BAB36F06B86FF2DAFD9FFF5FBA93C9D" blockId="231.[151,1437,726,2013]" lastBlockId="232.[151,1437,151,717]" lastPageId="232" lastPageNumber="233" pageId="231" pageNumber="232">Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae present; posterior ventral margin almost straight, not widened in apical half, posterior margin ventrally in apical half and dorsally completely serrated, glabrous. Metatibia moderately long and wide, dorsal and ventral margin subparallel, ratio width/length: ½.8, sharply carinate dorsally, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, basally with a few robust and short setae; lateral face weakly longitudinally convex, with coarse and moderately dense punctures; ventral margin with two widely separated spines; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres finely densely punctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere as long as following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate. Anterior claws symmetrical.</paragraph>
<paragraph id="8BAB36F06B89FF2DAFD9F9B1FCB73CB9" blockId="232.[151,1437,151,717]" box="[199,873,474,501]" pageId="232" pageNumber="233">
Aedeagus:
<figureCitation id="132F2A756B89FF2DAE5AF9B1FE4A3CB9" box="[324,404,474,501]" captionStart="FIGURE 53" captionStartId="230.[151,250,1891,1915]" captionTargetBox="[178,1406,195,1858]" captionTargetId="figure-19@230.[164,1423,181,1868]" captionTargetPageId="230" captionText="FIGURE 53. A–E: Maladera bikouensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. shiwandashanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. wupingensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, C, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496452" httpUri="https://zenodo.org/record/4496452/files/figure.png" pageId="232" pageNumber="233">Fig. 53</figureCitation>
F–H. Habitus:
<figureCitation id="132F2A756B89FF2DAD27F9B1FD4B3CB9" box="[569,661,474,501]" captionStart="FIGURE 53" captionStartId="230.[151,250,1891,1915]" captionTargetBox="[178,1406,195,1858]" captionTargetId="figure-19@230.[164,1423,181,1868]" captionTargetPageId="230" captionText="FIGURE 53. A–E: Maladera bikouensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. shiwandashanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. wupingensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, C, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496452" httpUri="https://zenodo.org/record/4496452/files/figure.png" pageId="232" pageNumber="233">Fig. 53I</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B89FF2DAFD9F994FB483F71" pageId="232" pageNumber="233" type="diagnosis">
<paragraph id="8BAB36F06B89FF2DAFD9F994FB483F71" blockId="232.[151,1437,151,717]" pageId="232" pageNumber="233">
<emphasis id="B960EAE26B89FF2DAFD9F994FE9C3F54" bold="true" box="[199,322,510,536]" pageId="232" pageNumber="233">Diagnosis.</emphasis>
<taxonomicName id="4C144D736B89FF2DAE56F995FC483F55" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[328,918,511,537]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="232" pageNumber="233" phylum="Arthropoda" rank="species" species="shiwandashanensis" status="sp. nov.">
<emphasis id="B960EAE26B89FF2DAE56F995FD4B3F55" box="[328,661,511,537]" italics="true" pageId="232" pageNumber="233">Maladera shiwandashanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B89FF2DACBFF995FC023F55" box="[929,988,511,537]" pageId="232" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
differs from the two previous species in having the phallobase with a wide and moderately deep dorsomedian apical emargination..
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B89FF2DAFD9FA2DFD873FC9" pageId="232" pageNumber="233" type="etymology">
<paragraph id="8BAB36F06B89FF2DAFD9FA2DFD873FC9" blockId="232.[151,1437,151,717]" pageId="232" pageNumber="233">
<emphasis id="B960EAE26B89FF2DAFD9FA2DFE903F2D" bold="true" box="[199,334,583,609]" pageId="232" pageNumber="233">Etymology.</emphasis>
The name of the new species (adjective in nominative singular case) is derived from its
<typeStatus id="54AF88526B89FF2DAA38FA2DFA883F2D" box="[1318,1366,583,609]" pageId="232" pageNumber="233">type</typeStatus>
locality, Shiwandashan National Forest Park.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B89FF2DAFD9FAE4FBAC3FE5" box="[199,1138,654,681]" pageId="232" pageNumber="233" type="description">
<paragraph id="8BAB36F06B89FF2DAFD9FAE4FBAC3FE5" blockId="232.[151,1437,151,717]" box="[199,1138,654,681]" pageId="232" pageNumber="233">
<emphasis id="B960EAE26B89FF2DAFD9FAE4FEE13FE4" bold="true" box="[199,319,654,680]" pageId="232" pageNumber="233">Variation.</emphasis>
Length:
<quantity id="4CEC9B156B89FF2DAEBAFAE5FDE53FE5" box="[420,571,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.08" metricValueMin="0.98" pageId="232" pageNumber="233" unit="mm" value="10.3" valueMax="10.8" valueMin="9.8">9.8–10.8 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156B89FF2DAC1EFAE5FC573FE5" box="[768,905,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.1" metricValueMin="7.4" pageId="232" pageNumber="233" unit="mm" value="7.75" valueMax="8.1" valueMin="7.4">7.4–8.1 mm</quantity>
, width:
<quantity id="4CEC9B156B89FF2DACFDFAE4FBB33FE5" box="[995,1133,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.2" metricValueMin="6.5" pageId="232" pageNumber="233" unit="mm" value="6.85" valueMax="7.2" valueMin="6.5">6.5–7.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B89FF2DAFD9FAD8FD133F81" box="[199,717,690,717]" pageId="232" pageNumber="233" type="distribution">
<paragraph id="8BAB36F06B89FF2DAFD9FAD8FD133F81" blockId="232.[151,1437,151,717]" box="[199,717,690,717]" pageId="232" pageNumber="233">
<emphasis id="B960EAE26B89FF2DAFD9FAD8FEBF3F80" bold="true" box="[199,353,690,716]" pageId="232" pageNumber="233">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B89FF2DAEC8FAD9FDEB3F81" box="[470,565,691,717]" captionStart="FIGURE 117" captionStartId="360.[151,250,1034,1058]" captionTargetBox="[210,1379,187,1002]" captionTargetId="figure-19@360.[202,1385,181,1010]" captionTargetPageId="360" captionText="FIGURE 117. Distribution of the Maladera liwenzhui group: Maladera bikouensis sp. n., M. liwenzhui sp. n., M. shiwandashanensis sp. n., M. wupingensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496636" httpUri="https://zenodo.org/record/4496636/files/figure.png" pageId="232" pageNumber="233">Fig. 117</figureCitation>
) and
<tableCitation id="C696034B6B89FF2DAD6DFAD9FD193F81" box="[627,711,691,717]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="232" pageNumber="233">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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