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<document id="5653412001E6F824ACCEF27893A77291" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66B02FFA0AF89FCECFCD139ED" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera invenusta" docType="treatment" docVersion="9" lastPageNumber="102" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="100" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="25080EF546FFEB4394272C6F56889717">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="97693F9F8E42CA7A9D0B042CC03B797F">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="FA8946E5D5903357FCCAFACB74075086">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="C57F159BB578440B927C48EA084A38E9">Bai, Ming</mods:namePart>
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<mods:namePart id="8FF1FB203110005F2D2098CCC717E204">Yang, Xing-Ke</mods:namePart>
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<mods:namePart id="F9F71322E1EBACB2E388DD097236E095">Ahrens, Dirk</mods:namePart>
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<taxonomicName id="4C144D736B02FFA6AF89FCECFDE439EC" authority="(Moser, 1918)" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[151,570,1158,1185]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="invenusta">
<emphasis id="B960EAE26B02FFA6AF89FCECFDE439EC" bold="true" box="[151,570,1158,1185]" pageId="99" pageNumber="100">
<emphasis id="B960EAE26B02FFA6AF89FCECFE5739ED" bold="true" box="[151,393,1158,1185]" italics="true" pageId="99" pageNumber="100">Maladera invenusta</emphasis>
(
<bibRefCitation id="EF854B016B02FFA6AE86FCECFDEC39EC" author="Moser, J." box="[408,562,1158,1184]" pageId="99" pageNumber="100" pagination="209 - 247" refId="ref334395" refString="Moser, J. (1918) Neue Melolonthiden aus der Sammlung des Deutschen Entomologischen Museums zu Berlin Dahlem (Col.). Stettiner Entomologische Zeitung, 79, 209 - 247." type="journal article" year="1918">Moser, 1918</bibRefCitation>
)
</emphasis>
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<paragraph id="8BAB36F06B02FFA6AF89FCC1FEB13989" blockId="99.[151,570,1158,1221]" box="[151,367,1194,1221]" pageId="99" pageNumber="100">
<figureCitation id="132F2A756B02FFA6AF89FCC1FED33989" box="[151,269,1195,1221]" captionStart="FIGURE 21" captionStartId="100.[151,250,1909,1933]" captionTargetBox="[195,1400,192,1873]" captionTargetId="figure-19@100.[184,1402,181,1884]" captionTargetPageId="100" captionText="FIGURE 21. A–D: Maladera invenusta (Moser) (China: Guangxi: Banbaxiang); E–H: M. futschauana (Brenske, 1897) (syntype: Autosercia montivaga Moser, 1915; Vietnam: Montes Mauson); I–L: M. longruiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496360" httpUri="https://zenodo.org/record/4496360/files/figure.png" pageId="99" pageNumber="100">Figures 21</figureCitation>
A–D, 95
</paragraph>
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<subSubSection id="C30E657B6B02FFA6AF89FC85FC3B386A" pageId="99" pageNumber="100" type="reference_group">
<paragraph id="8BAB36F06B02FFA6AF89FC85FCB7384B" blockId="99.[151,997,1263,1318]" box="[151,873,1263,1287]" pageId="99" pageNumber="100">
<treatmentCitationGroup id="AB0411DE6B02FFA6AF89FC85FCB7384B" box="[151,873,1263,1287]" pageId="99" pageNumber="100">
<treatmentCitation id="0AB510E16B02FFA6AF89FC85FDF6384B" author="Moser, J." box="[151,552,1263,1287]" page="212" pageId="99" pageNumber="100" year="1918">
<taxonomicName id="4C144D736B02FFA6AF89FC85FDF6384B" authority="Moser, 1918: 212" authorityName="Moser" authorityPageNumber="212" authorityYear="1918" box="[151,552,1263,1287]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="invenusta">
<emphasis id="B960EAE26B02FFA6AF89FC85FEB2384B" box="[151,364,1263,1287]" italics="true" pageId="99" pageNumber="100">Autoserica invenusta</emphasis>
<bibRefCitation id="EF854B016B02FFA6AE6DFC9AFDF6384B" author="Moser, J." box="[371,552,1263,1287]" pageId="99" pageNumber="100" pagination="209 - 247" refId="ref334395" refString="Moser, J. (1918) Neue Melolonthiden aus der Sammlung des Deutschen Entomologischen Museums zu Berlin Dahlem (Col.). Stettiner Entomologische Zeitung, 79, 209 - 247." type="journal article" year="1918">Moser, 1918: 212</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16B02FFA6AD2CFC85FCBB384B" author="Niijima, Y. &amp; Kinoshita, E." box="[562,869,1263,1287]" page="29" pageId="99" pageNumber="100" year="1923">
<bibRefCitation id="EF854B016B02FFA6AD2CFC85FCBB384B" author="Niijima, Y. &amp; Kinoshita, E." box="[562,869,1263,1287]" pageId="99" pageNumber="100" pagination="1 - 253" refId="ref334745" refString="Niijima, Y. &amp; Kinoshita, E. (1923) Die Untersuchungen u ¨ ber japanische Melolonthiden. II. Research Bulletin of the College Experimental Forestry Hokkaido Imperial University, 2, 1 - 253." type="journal article" year="1923">Niijima &amp; Kinoshita 1923: 29</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BAB36F06B02FFA6AF89FD64FC3B386A" blockId="99.[151,997,1263,1318]" box="[151,997,1294,1318]" pageId="99" pageNumber="100">
<treatmentCitationGroup id="AB0411DE6B02FFA6AF89FD64FC3B386A" box="[151,997,1294,1318]" pageId="99" pageNumber="100">
<treatmentCitation id="0AB510E16B02FFA6AF89FD64FDE7386A" author="Nomura, S." box="[151,569,1294,1318]" page="106" pageId="99" pageNumber="100" year="1974">
<taxonomicName id="4C144D736B02FFA6AF89FD64FDE7386A" authority=": Nomura, 1974: 106" authorityName="Nomura" authorityPageNumber="106" authorityYear="1974" box="[151,569,1294,1318]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="invenusta">
<emphasis id="B960EAE26B02FFA6AF89FD64FEBC3869" box="[151,354,1294,1318]" italics="true" pageId="99" pageNumber="100">Maladera invenusta</emphasis>
:
<bibRefCitation id="EF854B016B02FFA6AE6EFD64FDE7386A" author="Nomura, S." box="[368,569,1294,1318]" pageId="99" pageNumber="100" pagination="81 - 115" refId="ref335003" refString="Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115." type="journal article" year="1974">Nomura, 1974: 106</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16B02FFA6AD5CFD64FD33386A" author="Ahrens, D." box="[578,749,1294,1318]" page="85" pageId="99" pageNumber="100" year="2002">
<bibRefCitation id="EF854B016B02FFA6AD5CFD64FD33386A" author="Ahrens, D." box="[578,749,1294,1318]" pageId="99" pageNumber="100" pagination="53 - 91" refId="ref329519" refString="Ahrens, D. (2002) Notes on distribution and synonymy of sericid beetles from Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea, Sericini). Annales Historico-Naturales Musei Nationalis Hungarici, 94, 53 - 91." type="journal article" year="2002">Ahrens 2002: 85</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16B02FFA6ADE6FD64FCB0386A" author="Ahrens, D." box="[760,878,1294,1318]" page="237" pageId="99" pageNumber="100" year="2006">
<bibRefCitation id="EF854B016B02FFA6ADE6FD64FCB0386A" author="Ahrens, D." box="[760,878,1294,1318]" pageId="99" pageNumber="100" refId="ref329946" refString="Ahrens, D. (2006 b) Sericinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248." type="book" year="2006">2006b: 237</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16B02FFA6AC66FD64FC3F386A" author="Ahrens, D." box="[888,993,1294,1318]" page="23" pageId="99" pageNumber="100" year="2007">
<bibRefCitation id="EF854B016B02FFA6AC66FD64FC3F386A" author="Ahrens, D." box="[888,993,1294,1318]" pageId="99" pageNumber="100" pagination="1 - 51" refId="ref330155" refString="Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1" type="journal article" year="2007">2007b: 23</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B02FFA6AF89FD3EFACC3B6E" pageId="99" pageNumber="100" type="materials_examined">
<paragraph id="8BAB36F06B02FFA6AF89FD3EFD403822" blockId="99.[151,1437,1364,2039]" box="[151,670,1364,1390]" pageId="99" pageNumber="100">
<emphasis id="B960EAE26B02FFA6AF89FD3EFE613822" bold="true" box="[151,447,1364,1390]" pageId="99" pageNumber="100">
<typeStatus id="54AF88526B02FFA6AF89FD3EFF0C3822" box="[151,210,1364,1390]" pageId="99" pageNumber="100">Type</typeStatus>
material examined.
</emphasis>
See
<bibRefCitation id="EF854B016B02FFA6AEEAFD3EFD443822" author="Ahrens, D." box="[500,666,1364,1390]" pageId="99" pageNumber="100" pagination="53 - 91" refId="ref329519" refString="Ahrens, D. (2002) Notes on distribution and synonymy of sericid beetles from Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea, Sericini). Annales Historico-Naturales Musei Nationalis Hungarici, 94, 53 - 91." type="journal article" year="2002">Ahrens (2002)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAB36F06B02FFA6AFD9FD12FACC3B6E" blockId="99.[151,1437,1364,2039]" pageId="99" pageNumber="100">
<emphasis id="B960EAE26B02FFA6AFD9FD12FDE938DE" bold="true" box="[199,567,1400,1426]" pageId="99" pageNumber="100">Additional material examined.</emphasis>
<materialsCitation id="3B7C3CAD6B02FFA6AD21FD12FEA638FA" ID-GBIF-Occurrence="3034555733" collectingDate="2000-06-03" collectionCode="IZAS" collectorName="Li Wenzhu" country="China" location="Fangcheng" municipality="Banbaxiang" pageId="99" pageNumber="100" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guangxi">
<emphasis id="B960EAE26B02FFA6AD21FD12FD4F38DE" bold="true" box="[575,657,1400,1426]" pageId="99" pageNumber="100">
<collectingCountry id="F30376606B02FFA6AD21FD12FD5238DE" box="[575,652,1400,1426]" name="China" pageId="99" pageNumber="100">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796B02FFA6AD84FD12FD1938DF" box="[666,711,1400,1427]" pageId="99" pageNumber="100" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D12FD796B02FFA6ADCAFD12FD2138DF" box="[724,767,1400,1427]" pageId="99" pageNumber="100" type="female">1 ♀</specimenCount>
“
<collectingMunicipality id="6BCFAC8A6B02FFA6AC0FFD12FC4238DE" box="[785,924,1400,1426]" pageId="99" pageNumber="100">Banbaxiang</collectingMunicipality>
,
<location id="8ECB602B6B02FFA6ACB7FD12FBF938DE" LSID="urn:lsid:plazi:treatment:03BD87E66B02FFA0AF89FCECFCD139ED:8ECB602B6B02FFA6ACB7FD12FBF938DE" box="[937,1063,1400,1426]" country="China" municipality="Banbaxiang" name="Fangcheng" pageId="99" pageNumber="100" stateProvince="Guangxi">Fangcheng</location>
,
<collectingRegion id="49D0F8126B02FFA6AB2DFD12FB4838DE" box="[1075,1174,1400,1426]" country="China" name="Guangxi" pageId="99" pageNumber="100">Guangxi</collectingRegion>
,
<date id="FFAA10306B02FFA6ABBDFD12FAC938DE" box="[1187,1303,1400,1426]" pageId="99" pageNumber="100" value="2000-06-03">
<collectingDate id="EFEEE9D86B02FFA6ABBDFD12FAC938DE" box="[1187,1303,1400,1426]" pageId="99" pageNumber="100" value="2000-06-03">
3.VI.
<quantity id="4CEC9B156B02FFA6ABC6FD12FAC938DE" box="[1240,1303,1400,1426]" metricMagnitude="6" metricUnit="m" metricValue="2.00025" pageId="99" pageNumber="100" unit="m" value="2000250.0">2000</quantity>
</collectingDate>
</date>
, 250m, leg.
<collectorName id="26E153266B02FFA6AF89FDF6FECA38FA" box="[151,276,1436,1462]" pageId="99" pageNumber="100">Li Wenzhu</collectorName>
” (
<collectionCode id="ED05AE356B02FFA6AE32FDF6FEB138FA" box="[300,367,1436,1462]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="99" pageNumber="100">IZAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6B02FFA6AE9DFDF6FBB938FA" ID-GBIF-Occurrence="3034556625" box="[387,1127,1436,1463]" collectingDate="1976-06-26" collectionCode="IZAS" collectorName="Huang Ju" country="China" location="Laoda" pageId="99" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Anhui">
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“
<location id="8ECB602B6B02FFA6AEA3FDF6FDD638FA" LSID="urn:lsid:plazi:treatment:03BD87E66B02FFA0AF89FCECFCD139ED:8ECB602B6B02FFA6AEA3FDF6FDD638FA" box="[445,520,1436,1462]" country="China" name="Laoda" pageId="99" pageNumber="100" stateProvince="Anhui">Laoda</location>
,
<collectingRegion id="49D0F8126B02FFA6AD0FFDF6FD8438FA" box="[529,602,1436,1462]" country="China" name="Anhui" pageId="99" pageNumber="100">Anhui</collectingRegion>
,
<date id="FFAA10306B02FFA6AD7BFDF6FD3938FA" box="[613,743,1436,1462]" pageId="99" pageNumber="100" value="1976-06-26">
<collectingDate id="EFEEE9D86B02FFA6AD7BFDF6FD3938FA" box="[613,743,1436,1462]" pageId="99" pageNumber="100" value="1976-06-26">26.VI.1976</collectingDate>
</date>
,
<collectingMethod id="52554EE76B02FFA6ADECFDF6FC8438FB" box="[754,858,1436,1463]" pageId="99" pageNumber="100">light trap</collectingMethod>
, leg.
<collectorName id="26E153266B02FFA6AC8BFDF7FBDF38FA" box="[917,1025,1436,1463]" pageId="99" pageNumber="100">Huang Ju</collectorName>
” (
<collectionCode id="ED05AE356B02FFA6AB05FDF6FB8138FA" box="[1051,1119,1436,1462]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="99" pageNumber="100">IZAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6B02FFA6AB6FFDF6FC0F3896" ID-GBIF-Occurrence="3034556792" collectingDate="1996-05-21" collectingDateMax="1996-05-22" collectingDateMin="1996-05-21" collectionCode="CP" collectorName="Pacholatko &amp; Dembicky" country="Vietnam" location="Vietnam Cuc Phuong Nat. Park" pageId="99" pageNumber="100" specimenCount="1" specimenCount-male="1">
<emphasis id="B960EAE26B02FFA6AB6FFDF6FB3E38FA" bold="true" box="[1137,1248,1436,1462]" pageId="99" pageNumber="100">
<collectingCountry id="F30376606B02FFA6AB6FFDF6FB0538FA" box="[1137,1243,1436,1462]" name="Vietnam" pageId="99" pageNumber="100">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796B02FFA6ABF8FDF6FACE38FB" box="[1254,1296,1436,1463]" pageId="99" pageNumber="100" type="male">1 ♂</specimenCount>
“N
<location id="8ECB602B6B02FFA6AA23FDF6FE4A3896" LSID="urn:lsid:plazi:treatment:03BD87E66B02FFA0AF89FCECFCD139ED:8ECB602B6B02FFA6AA23FDF6FE4A3896" country="Vietnam" name="Vietnam Cuc Phuong Nat. Park" pageId="99" pageNumber="100">Vietnam Cuc Phuong Nat. Park</location>
<date id="FFAA10306B02FFA6AE82FDAAFDE83896" box="[412,566,1472,1498]" pageId="99" pageNumber="100" value="1996-05-21" valueMax="1996-05-22" valueMin="1996-05-21">
<collectingDate id="EFEEE9D86B02FFA6AE82FDAAFDE83896" box="[412,566,1472,1498]" pageId="99" pageNumber="100" value="1996-05-21" valueMax="1996-05-22" valueMin="1996-05-21">21.-22.v.1996</collectingDate>
</date>
<collectorName id="26E153266B02FFA6AD20FDAAFD643896" box="[574,698,1472,1498]" pageId="99" pageNumber="100">Pacholátko</collectorName>
&amp;
<collectorName id="26E153266B02FFA6ADFEFDAAFC8B3896" box="[736,853,1472,1498]" pageId="99" pageNumber="100">Dembicky</collectorName>
leg.” (
<collectionCode id="ED05AE356B02FFA6ACBCFDAAFC173896" box="[930,969,1472,1498]" pageId="99" pageNumber="100">CP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6B02FFA6ACC0FDAAFCCD38B2" ID-GBIF-Occurrence="3034556382" collectingDate="1994-05-15" collectionCode="CP" collectorName="Nam Cat Tien-Nat. Park P. Pacholatko &amp; L. Dembicky" country="Vietnam" location="Vietnam" pageId="99" pageNumber="100" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD796B02FFA6ACC0FDAAFBD63897" box="[990,1032,1472,1499]" pageId="99" pageNumber="100" type="male">1 ♂</specimenCount>
“S
<location id="8ECB602B6B02FFA6AB2DFDAAFB4B3896" LSID="urn:lsid:plazi:treatment:03BD87E66B02FFA0AF89FCECFCD139ED:8ECB602B6B02FFA6AB2DFDAAFB4B3896" box="[1075,1173,1472,1498]" country="Vietnam" name="Vietnam" pageId="99" pageNumber="100">Vietnam</location>
, 1.-
<date id="FFAA10306B02FFA6ABABFDAAFAF13897" box="[1205,1327,1472,1499]" pageId="99" pageNumber="100" value="1994-05-15">
<collectingDate id="EFEEE9D86B02FFA6ABABFDAAFAF13897" box="[1205,1327,1472,1499]" pageId="99" pageNumber="100" value="1994-05-15">15.5.1994</collectingDate>
</date>
<collectorName id="26E153266B02FFA6AA29FDABFE0238B2" pageId="99" pageNumber="100">Nam Cat Tien-Nat. Park P. Pacholátko</collectorName>
&amp;
<collectorName id="26E153266B02FFA6AD1FFD8FFD4838B2" box="[513,662,1508,1535]" pageId="99" pageNumber="100">L. Dembicky</collectorName>
leg.” (
<collectionCode id="ED05AE356B02FFA6ADFDFD8EFCD438B2" box="[739,778,1508,1534]" pageId="99" pageNumber="100">CP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6B02FFA6AC00FD8EFAD03B6E" ID-GBIF-Occurrence="3034556524" collectingDate="1998-05-08" collectionCode="GPS, CP" collectorName="E. Jendek &amp; O. Sausa" country="Laos" county="Laos" elevation="20" location="Nakai" municipality="Khamousan prov." pageId="99" pageNumber="100" specimenCount="1" specimenCount-male="1">
<emphasis id="B960EAE26B02FFA6AC00FD8EFCBE38B2" bold="true" box="[798,864,1508,1534]" pageId="99" pageNumber="100">
<collectingCountry id="F30376606B02FFA6AC00FD8EFC8238B2" box="[798,860,1508,1534]" name="Laos" pageId="99" pageNumber="100">Laos</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796B02FFA6AC76FD8EFC4D38B3" box="[872,915,1508,1535]" pageId="99" pageNumber="100" type="male">1 ♂</specimenCount>
“
<collectingCounty id="62CA4E7C6B02FFA6ACBAFD8EFC0338B2" box="[932,989,1508,1534]" pageId="99" pageNumber="100">Laos</collectingCounty>
centr.,
<collectingMunicipality id="6BCFAC8A6B02FFA6AB2FFD8EFB2938B3" box="[1073,1271,1508,1535]" pageId="99" pageNumber="100">Khamousan prov.</collectingMunicipality>
<location id="8ECB602B6B02FFA6ABE0FD8EFA9F38B2" LSID="urn:lsid:plazi:treatment:03BD87E66B02FFA0AF89FCECFCD139ED:8ECB602B6B02FFA6ABE0FD8EFA9F38B2" box="[1278,1345,1508,1534]" country="Laos" county="Laos" municipality="Khamousan prov." name="Nakai" pageId="99" pageNumber="100">Nakai</location>
env., 4-
<date id="FFAA10306B02FFA6AF89FE62FF223B6F" box="[151,252,1544,1571]" pageId="99" pageNumber="100" value="1998-05-08">
<collectingDate id="EFEEE9D86B02FFA6AF89FE62FF223B6F" box="[151,252,1544,1571]" pageId="99" pageNumber="100" value="1998-05-08">8.5.1998</collectingDate>
</date>
, Route No. 8, alt. 560±
<quantity id="4CEC9B156B02FFA6AD1EFE62FDEC3B6F" box="[512,562,1544,1571]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="99" pageNumber="100" unit="m" value="20.0">
<elevation id="0039D1C36B02FFA6AD1EFE62FDEC3B6F" box="[512,562,1544,1571]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="99" pageNumber="100" unit="m" value="20.0">20m</elevation>
</quantity>
N17°42.8’, E 105°08.9’ (
<collectionCode id="ED05AE356B02FFA6AC4CFE62FC543B6E" box="[850,906,1544,1570]" pageId="99" pageNumber="100">GPS</collectionCode>
),
<collectorName id="26E153266B02FFA6AC80FE63FBD53B6E" box="[926,1035,1544,1571]" pageId="99" pageNumber="100">E. Jendek</collectorName>
&amp;
<collectorName id="26E153266B02FFA6AB30FE62FB4C3B68" box="[1070,1170,1544,1572]" pageId="99" pageNumber="100">O. Šauša</collectorName>
leg.” (
<collectionCode id="ED05AE356B02FFA6ABC0FE62FADB3B6E" box="[1246,1285,1544,1570]" pageId="99" pageNumber="100">CP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B02FFA0AFD9FE46FD793915" lastPageId="101" lastPageNumber="102" pageId="99" pageNumber="100" type="description">
<paragraph id="8BAB36F06B02FFA6AFD9FE46FA573B26" blockId="99.[151,1437,1364,2039]" pageId="99" pageNumber="100">
<emphasis id="B960EAE26B02FFA6AFD9FE46FEAB3B0A" bold="true" box="[199,373,1580,1606]" pageId="99" pageNumber="100">Redescription.</emphasis>
Length:
<quantity id="4CEC9B156B02FFA6AEC7FE46FDEE3B0B" box="[473,560,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="99" pageNumber="100" unit="mm" value="6.8">6.8 mm</quantity>
, length of elytra: 5.0 mm, width: 4.0 mm. Body oval, black to blacking brown, antenna yellow, club brown, dorsal surface dull, except some single setae on head dorsal surface nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06B02FFA6AF89FE1EFE233AE2" blockId="99.[151,1437,1364,2039]" pageId="99" pageNumber="100">
Labroclypeus moderately wide and trapezoidal, shiny, widest at base, lateral margins weakly convex and strongly convergent anteriorly, anterior angles blunt and slightly reflexed, anterior margin moderately emarginate, margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, finely and densely punctate, with numerous erect setae anteriorly; frontoclypeal suture indistinctly incised, curved; smooth area anterior to eye flat, 2.5 times as wide as long; ocular canthus long and moderately wide (nearly 1/2 of ocular diameter), finely and sparsely punctate, without terminal seta. Frons dull, with fine, dense punctures and a few single erect setae beside eyes. Eyes large, ratio diameter/ interocular width: 0.84. Antenna with ten antennomeres; club with three antennomeres and weakly reflexed, slightly longer than remaining antennomeres combined.
<collectingRegion id="49D0F8126B02FFA6ABC4FF1BFAE73AC7" box="[1242,1337,1905,1931]" country="Myanmar" name="Chin" pageId="99" pageNumber="100">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06B02FFA0AFD9FFD3FB723DB5" blockId="99.[151,1437,1364,2039]" lastBlockId="101.[151,1437,151,1185]" lastPageId="101" lastPageNumber="102" pageId="99" pageNumber="100">Pronotum moderately transverse, widest at posterior third, lateral margins evenly convex and convergent anteriorly and posteriorly, anterior angles moderately produced and sharply pointed, posterior angles convex; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures, otherwise glabrous; lateral and anterior margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, sparse punctures.</paragraph>
<caption id="DF6B66786B05FFA1AF89FF1FFDA53AB5" ID-DOI="http://doi.org/10.5281/zenodo.4496360" ID-Zenodo-Dep="4496360" httpUri="https://zenodo.org/record/4496360/files/figure.png" pageId="100" pageNumber="101" startId="100.[151,250,1909,1933]" targetBox="[195,1400,192,1873]" targetPageId="100">
<paragraph id="8BAB36F06B05FFA1AF89FF1FFDA53AB5" blockId="100.[151,1436,1909,2041]" pageId="100" pageNumber="101">
<emphasis id="B960EAE26B05FFA1AF89FF1FFE813AC1" bold="true" box="[151,351,1909,1933]" pageId="100" pageNumber="101">FIGURE 21. A–D:</emphasis>
<taxonomicName id="4C144D736B05FFA1AE78FF1FFD543AC1" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[358,650,1909,1933]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="invenusta">
<emphasis id="B960EAE26B05FFA1AE78FF1FFDEC3AC1" box="[358,562,1909,1933]" italics="true" pageId="100" pageNumber="101">Maladera invenusta</emphasis>
(Moser)
</taxonomicName>
(China: Guangxi: Banbaxiang);
<emphasis id="B960EAE26B05FFA1ACC6FF1FFBCC3AC1" bold="true" box="[984,1042,1909,1933]" pageId="100" pageNumber="101">E–H:</emphasis>
<taxonomicName id="4C144D736B05FFA1AB07FF1CFABF3AC1" authority="(Brenske, 1897)" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[1049,1377,1909,1933]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="futschauana">
<emphasis id="B960EAE26B05FFA1AB07FF1CFB693AC1" box="[1049,1207,1909,1933]" italics="true" pageId="100" pageNumber="101">M. futschauana</emphasis>
(
<bibRefCitation id="EF854B016B05FFA1ABDBFF1FFA843AC1" author="Brenske, E." box="[1221,1370,1909,1933]" pageId="100" pageNumber="101" pagination="345 - 468" refId="ref331423" refString="Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310" type="journal article" year="1897">Brenske, 1897</bibRefCitation>
)
</taxonomicName>
(syntype:
<taxonomicName id="4C144D736B05FFA1AFCEFFF3FDE83AFD" authority="Moser, 1915" authorityName="Moser" authorityYear="1915" box="[208,566,1945,1969]" class="Insecta" family="Melolonthidae" genus="Autosercia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="montivaga">
<emphasis id="B960EAE26B05FFA1AFCEFFF3FE703AFD" box="[208,430,1945,1969]" italics="true" pageId="100" pageNumber="101">Autosercia montivaga</emphasis>
Moser, 1915
</taxonomicName>
; Vietnam: Montes Mauson);
<emphasis id="B960EAE26B05FFA1AC42FFF3FC553AFD" bold="true" box="[860,907,1945,1969]" pageId="100" pageNumber="101">I–L:</emphasis>
<taxonomicName id="4C144D736B05FFA1AC8CFFF0FAC73AFD" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[914,1305,1945,1969]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="longruiensis" status="sp. nov.">
<emphasis id="B960EAE26B05FFA1AC8CFFF0FBEE3AFD" box="[914,1072,1945,1969]" italics="true" pageId="100" pageNumber="101">M. longruiensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B05FFA1AA3CFFF3FA873AFC" box="[1314,1369,1945,1968]" pageId="100" pageNumber="101" rank="species">sp. n.</taxonomicNameLabel>
(holotype).
<emphasis id="B960EAE26B05FFA1AFC9FFD7FEFB3A99" bold="true" box="[215,293,1981,2005]" pageId="100" pageNumber="101">A, E, I:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26B05FFA1AD73FFD7FD193A99" bold="true" box="[621,711,1981,2005]" pageId="100" pageNumber="101">C, G, K:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26B05FFA1AB03FFD7FBB73A99" bold="true" box="[1053,1129,1981,2005]" pageId="100" pageNumber="101">B, F, J:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26B05FFA1AA7AFFD7FF6E3AB5" bold="true" pageId="100" pageNumber="101">D, H, L:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BAB36F06B04FFA0AFD9F969FE9A3CE2" blockId="101.[151,1437,151,1185]" pageId="101" pageNumber="102">Elytra widest shortly behind middle, striae distinctly impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, odd intervals with a few single short setae; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra not membraneous, without visible rim of microtrichomes (at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06B04FFA0AFD9F9DDFECA3FC9" blockId="101.[151,1437,151,1185]" pageId="101" pageNumber="102">Ventral surface dull, coarsely and moderately densely punctate, metasternum sparsely covered with sparse fine setae; metacoxa with a few longer setae laterally, otherwise glabrous. Abdominal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.6. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few longer setae on apical half.</paragraph>
<paragraph id="8BAB36F06B04FFA0AFD9FAE5FAC03979" blockId="101.[151,1437,151,1185]" pageId="101" pageNumber="102">Legs moderately wide and short, weakly shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, with anterior margin acute, with continuous serrated line behind anterior margin, setae of anterior row very short; posterior margin smooth and moderately widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at apex, ratio of width/length: 1/2.4, sharply carinate dorsally, with two groups of spines, basal group at four sixths, apical group at five sixths of metatibial length, with a few short robust setae basally and a serrated line beside dorsal margin in basal half; lateral face weakly longitudinally convex, moderately densely punctate, with minute setae in punctures; ventral margin finely serrate, with four robust equidistant setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06B04FFA0AFD9FC55FD793915" blockId="101.[151,1437,151,1185]" box="[199,679,1087,1113]" pageId="101" pageNumber="102">
Aedeagus:
<figureCitation id="132F2A756B04FFA0AE5AFC55FE4D3915" box="[324,403,1087,1113]" captionStart="FIGURE 21" captionStartId="100.[151,250,1909,1933]" captionTargetBox="[195,1400,192,1873]" captionTargetId="figure-19@100.[184,1402,181,1884]" captionTargetPageId="100" captionText="FIGURE 21. A–D: Maladera invenusta (Moser) (China: Guangxi: Banbaxiang); E–H: M. futschauana (Brenske, 1897) (syntype: Autosercia montivaga Moser, 1915; Vietnam: Montes Mauson); I–L: M. longruiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496360" httpUri="https://zenodo.org/record/4496360/files/figure.png" pageId="101" pageNumber="102">Fig. 21</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756B04FFA0AD23FC55FD7D3915" box="[573,675,1087,1113]" captionStart="FIGURE 21" captionStartId="100.[151,250,1909,1933]" captionTargetBox="[195,1400,192,1873]" captionTargetId="figure-19@100.[184,1402,181,1884]" captionTargetPageId="100" captionText="FIGURE 21. A–D: Maladera invenusta (Moser) (China: Guangxi: Banbaxiang); E–H: M. futschauana (Brenske, 1897) (syntype: Autosercia montivaga Moser, 1915; Vietnam: Montes Mauson); I–L: M. longruiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496360" httpUri="https://zenodo.org/record/4496360/files/figure.png" pageId="101" pageNumber="102">Fig. 21D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B04FFA0AFD9FC08FCD139ED" pageId="101" pageNumber="102" type="distribution">
<paragraph id="8BAB36F06B04FFA0AFD9FC08FCD139ED" blockId="101.[151,1437,151,1185]" pageId="101" pageNumber="102">
<emphasis id="B960EAE26B04FFA0AFD9FC08FEBF3930" bold="true" box="[199,353,1122,1148]" pageId="101" pageNumber="102">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B04FFA0AEC8FC09FDF93931" box="[470,551,1122,1149]" captionStart="FIGURE 95" captionStartId="349.[151,250,1031,1055]" captionTargetBox="[210,1378,187,1002]" captionTargetId="figure-20@349.[201,1386,181,1008]" captionTargetPageId="349" captionText="FIGURE 95. Distribution of the Maladera teizoana group: Maladera fereobscurata sp. n., M. futschauana (Brenske, 1897), M. invenusta (Moser, 1918), Maladera piceola (Moser, 1915) (not precisely localized collection site indicated by a “?”)." figureDoi="http://doi.org/10.5281/zenodo.4496572" httpUri="https://zenodo.org/record/4496572/files/figure.png" pageId="101" pageNumber="102">Fig. 95</figureCitation>
) and
<tableCitation id="C696034B6B04FFA0AD7BFC09FD673931" box="[613,697,1123,1149]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="101" pageNumber="102">Table 1</tableCitation>
. This species was originally described from
<collectingCountry id="F30376606B04FFA0ABB4FC09FB213931" box="[1194,1279,1123,1149]" name="Taiwan" pageId="101" pageNumber="102">Taiwan</collectingCountry>
, it is recorded for mainland
<collectingCountry id="F30376606B04FFA0AE30FCEDFEAA39ED" box="[302,372,1159,1185]" name="China" pageId="101" pageNumber="102">China</collectingCountry>
,
<collectingCountry id="F30376606B04FFA0AE61FCEDFE6B39ED" box="[383,437,1159,1185]" name="Laos" pageId="101" pageNumber="102">Laos</collectingCountry>
and
<collectingCountry id="F30376606B04FFA0AEF5FCEDFD9439ED" box="[491,586,1159,1185]" name="Vietnam" pageId="101" pageNumber="102">Vietnam</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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