treatments-xml/data/03/BD/87/03BD87B5FF94FFB825B94F57FC4FFF2A.xml

<document id="61BC7F6600FA2FFDEC1D0F89618B7039" ID-CLB-Dataset="45899" ID-DOI="10.5281/zenodo.279301" ID-GBIF-Dataset="67e7ec16-df1a-4241-a454-8dfb4372336b" ID-ISSN="1175-5326" ID-Zenodo-Dep="279301" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460414038779" checkinUser="plazi" docAuthor="Ospina-Sarria, Jhon Jairo, Mendez-Narvaez, Javier, Burbano-Yandi, Carlos &amp; Bolívar-G, Wilmar" docDate="2011" docId="03BD87B5FF94FFB825B94F57FC4FFF2A" docLanguage="en" docName="zt03111p048.pdf" docOrigin="Zootaxa 3111" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pristimantis quicato Ospina-Sarria, Mendez-Narvaez, Burbano-Yandi &amp; Bolívar-G, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="46" masterDocId="FF84FFCDFF95FFB1252E4C0BFFF7FFFE" masterDocTitle="A new species of Pristimantis (Amphibia: Craugastoridae) with cranial crests from the Colombian Andes" masterLastPageNumber="48" masterPageNumber="37" pageNumber="38" updateTime="1698328335530" updateUser="ExternalLinkService">
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<mods:title id="990DDE954A346967EBF416FBCF8B8285">A new species of Pristimantis (Amphibia: Craugastoridae) with cranial crests from the Colombian Andes</mods:title>
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<mods:namePart id="078B67BDFBCB159887D4C3D9548FDC3F">Ospina-Sarria, Jhon Jairo</mods:namePart>
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<mods:namePart id="2ECBCEEB70E36E15CB9A41679B040E53">Mendez-Narvaez, Javier</mods:namePart>
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<mods:namePart id="58CC0718144DA7101CA93017B248D401">Burbano-Yandi, Carlos</mods:namePart>
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<mods:namePart id="0528B3D0D1D399FC0D0EF912039EDF27">Bolívar-G, Wilmar</mods:namePart>
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<treatment id="03BD87B5FF94FFB825B94F57FC4FFF2A" ID-DOI="http://doi.org/10.5281/zenodo.5678602" ID-GBIF-Taxon="119424470" ID-Zenodo-Dep="5678602" LSID="urn:lsid:plazi:treatment:03BD87B5FF94FFB825B94F57FC4FFF2A" httpUri="http://treatment.plazi.org/id/03BD87B5FF94FFB825B94F57FC4FFF2A" lastPageId="9" lastPageNumber="46" pageId="1" pageNumber="38">
<subSubSection id="C30E6528FF94FFB025B94F57FE3FFC68" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="8BAB36A3FF94FFB025B94F57FE18FC8B" blockId="1.[151,495,859,918]" box="[151,495,859,886]" pageId="1" pageNumber="38">
<heading id="D0E381CFFF94FFB025B94F57FE18FC8B" bold="true" box="[151,495,859,886]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="B960EAB1FF94FFB025B94F57FE18FC8B" bold="true" box="[151,495,859,886]" pageId="1" pageNumber="38">
<taxonomicName id="4C144D20FF94FFB025B94F57FE7CFC88" box="[151,395,860,886]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="38" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF94FFB025B94F57FE7CFC88" bold="true" box="[151,395,860,886]" italics="true" pageId="1" pageNumber="38">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF94FFB024BC4F50FE18FC8B" box="[402,495,859,885]" pageId="1" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB36A3FF94FFB025B94F75FE3FFC68" blockId="1.[151,495,859,918]" box="[151,456,894,918]" pageId="1" pageNumber="38">
(
<figureCitation id="132F2A26FF94FFB025B04F75FF09FC68" box="[158,254,894,918]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="1" pageNumber="38">Figure 2</figureCitation>
A, 3A, 4, 5A, 7A)
</paragraph>
</subSubSection>
<subSubSection id="C30E6528FF94FFB025B94FCCFAFBFA94" pageId="1" pageNumber="38" type="materials_examined">
<paragraph id="8BAB36A3FF94FFB025B94FCCFD87FBD8" blockId="1.[151,1436,966,2034]" pageId="1" pageNumber="38">
<typeStatus id="54AF8801FF94FFB025B94FCCFEF3FC21" box="[151,260,967,991]" pageId="1" pageNumber="38" type="holotype">
<emphasis id="B960EAB1FF94FFB025B94FCCFEF3FC21" bold="true" box="[151,260,967,991]" pageId="1" pageNumber="38">Holotype</emphasis>
</typeStatus>
. 
<collectionCode id="ED05AE66FF94FFB0243A4FCDFEB3FC20" box="[276,324,966,990]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="1" pageNumber="38">ICN</collectionCode>
55409, adult female, collected on 
<date id="FFAA1063FF94FFB027FA4FCDFC7DFC20" box="[724,906,966,990]" pageId="1" pageNumber="38" value="2011-04-30">30th April 2011</date>
by Wilmar Bolívar-G. at Reserva Natural “La Sirena” (3º 31', 
<geoCoordinate id="EE205064FF94FFB0247E4FE1FE33FBFC" box="[336,452,1002,1026]" direction="west" orientation="longitude" pageId="1" pageNumber="38" precision="925" value="-76.1">76º 06' W</geoCoordinate>
, 
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), Vereda la Nevera, Corregimiento de Tenjo, Municipality of Palmira, Departamento Valle del Cauca, 
<collectingCountry id="F3037633FF94FFB024D44805FD9BFBD8" box="[506,620,1038,1062]" name="Colombia" pageId="1" pageNumber="38">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="8BAB36A3FF94FFB025E94838FD3BFB90" blockId="1.[151,1436,966,2034]" pageId="1" pageNumber="38">
<emphasis id="B960EAB1FF94FFB025E94838FE84FBB5" bold="true" box="[199,371,1075,1099]" pageId="1" pageNumber="38">Paratopotypes</emphasis>
. 
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55406–08, 55410–11, all adult males, collected on 
<date id="FFAA1063FF94FFB026DC4839FB57FBB4" box="[1010,1184,1074,1098]" pageId="1" pageNumber="38" value="2011-04-31">31st April 2011</date>
by Jhon Jairo Ospina- Sarria, Javier Mendez, and Carlos Burbano-Yandi.
</paragraph>
<paragraph id="8BAB36A3FF94FFB025E94871FE80FB48" blockId="1.[151,1436,966,2034]" pageId="1" pageNumber="38">
<emphasis id="B960EAB1FF94FFB025E94871FE4AFB6D" bold="true" box="[199,445,1146,1171]" pageId="1" pageNumber="38">Referred specimens.</emphasis>
UV–CD 
<date id="FFAA1063FF94FFB027014871FD63FB6C" box="[559,660,1146,1170]" pageId="1" pageNumber="38" value="1917" valueMax="1921">1917–21</date>
, Juveniles males and UV–CD 
<date id="FFAA1063FF94FFB026D24871FB96FB6C" box="[1020,1121,1146,1170]" pageId="1" pageNumber="38" value="1922" valueMax="1924">1922–24</date>
, subadults males, collected with the 
<typeStatus id="54AF8801FF94FFB025D74894FEDEFB49" box="[249,297,1183,1207]" pageId="1" pageNumber="38">type</typeStatus>
series.
</paragraph>
<paragraph id="8BAB36A3FF94FFB025E948C9FAFBFA94" blockId="1.[151,1436,966,2034]" pageId="1" pageNumber="38">
<emphasis id="B960EAB1FF94FFB025E948C9FE5DFB25" bold="true" box="[199,426,1218,1243]" pageId="1" pageNumber="38">Generic allocation.</emphasis>
Currently there are no morphological characters supporting the monophyly of the genus 
<taxonomicName id="4C144D20FF94FFB025B948ECFDE4FB00" authority="Hedges et al. 2008" authorityName="Hedges et al." authorityYear="2008" box="[151,531,1254,1279]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B960EAB1FF94FFB025B948ECFED7FB00" box="[151,288,1255,1278]" italics="true" pageId="1" pageNumber="38">Pristimantis</emphasis>
(
<bibRefCitation id="EF854B52FF94FFB0240148EDFDFCFB00" author="Hedges" box="[303,523,1254,1279]" pageId="1" pageNumber="38" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New world direct-developing frogs (Anura: Terrarana): Molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737, 1 - 182." type="journal article" year="2008">
Hedges 
<emphasis id="B960EAB1FF94FFB024A348E3FE48FB00" box="[397,447,1254,1279]" italics="true" pageId="1" pageNumber="38">et al</emphasis>
. 2008
</bibRefCitation>
)
</taxonomicName>
. However, 
<taxonomicName id="4C144D20FF94FFB027BD48ECFCEBFB00" box="[659,796,1255,1278]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B960EAB1FF94FFB027BD48ECFCEBFB00" box="[659,796,1255,1278]" italics="true" pageId="1" pageNumber="38">Pristimantis</emphasis>
</taxonomicName>
is mostly composed of small, slender, direct-developing frogs from South 
<collectingCountry id="F3037633FF94FFB0244F4901FE32FADC" box="[353,453,1290,1314]" name="United States of America" pageId="1" pageNumber="38">America</collectingCountry>
, and all species resembling the new taxon have been placed in the genus 
<taxonomicName id="4C144D20FF94FFB020224907FA62FADD" box="[1292,1429,1292,1315]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B960EAB1FF94FFB020224907FA62FADD" box="[1292,1429,1292,1315]" italics="true" pageId="1" pageNumber="38">Pristimantis</emphasis>
</taxonomicName>
. Thus, despite we assume that additional phylogenetic evidence might eventually lead to changes in the generic allocation of the new species, assigning it to 
<taxonomicName id="4C144D20FF94FFB027A2495FFCE2FA95" box="[652,789,1364,1387]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B960EAB1FF94FFB027A2495FFCE2FA95" box="[652,789,1364,1387]" italics="true" pageId="1" pageNumber="38">Pristimantis</emphasis>
</taxonomicName>
is the most prudent and conservative option.
</paragraph>
</subSubSection>
<subSubSection id="C30E6528FF94FFB225E9497DFB31F8D9" lastPageId="3" lastPageNumber="40" pageId="1" pageNumber="38" type="diagnosis">
<paragraph id="8BAB36A3FF94FFB325E9497DFD40FE52" blockId="1.[151,1436,966,2034]" lastBlockId="2.[151,1437,151,428]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
<emphasis id="B960EAB1FF94FFB025E9497DFECDFA71" bold="true" box="[199,314,1398,1423]" pageId="1" pageNumber="38">Diagnosis</emphasis>
. 
<taxonomicName id="4C144D20FF94FFB02464497CFDC7FA70" box="[330,560,1399,1422]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="38" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF94FFB02464497CFDC7FA70" box="[330,560,1399,1422]" italics="true" pageId="1" pageNumber="38">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF94FFB02717497CFD62FA71" box="[569,661,1399,1423]" pageId="1" pageNumber="38" rank="species">
<emphasis id="B960EAB1FF94FFB02717497CFD62FA71" bold="true" box="[569,661,1399,1423]" pageId="1" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
presents the following characteristics: (1) dorsal skin granular with some conical tubercles distributed uniformly, and forming ridges (H-shaped) on shoulder; ventral skin aerolate; discoidal fold present; dorsolateral folds present, fragmented into a series of tubercles from the posterolateral corner of the orbit, and reaching the anterior half of body; (2) tympanic membrane differentiated; tympanic annulus prominent, its length 1/3–1/4 eye length, with upper edge obscured by supratympanic fold; (3) snout subacuminate in dorsal view (more rounded in females), rounded in profile; canthus rostralis slightly concave; (4) upper eyelid with small round tubercles, and one large, conical tubercle; upper eyelid narrower than IOD; low cranial crests present; (5) choanae small and partly concealed by palatal shelf of maxilla; dentigerous processes of vomers oblique, positioned posterior to level of choanae and separated medially by distance less than width of the visible vomerine process, each process bearing 6–7 teeth; (6) males lacking vocal slits, but having glandular, white nuptial pads; (7) Finger I shorter than Finger II, discs on outer fingers broad, about twice the width of digit proximal to pad; (8) fingers with broad lateral fringes; outer palmar tubercle fully divided, elliptical and equal in size; round supernumerary tubercles, subarticular tubercles larger than supernumerary tubercles, tubercles round and projecting; (9) ulnar tubercles present, subconical, folds absent; (10) heel bearing small tubercles, and one large, conical tubercle; outer edge of tarsus bearing subconical tubercles; an indistinct inner tarsal fold is present on the distal quarter of tarsus; (11) inner metatarsal tubercle oval, six to seven times the size of the oval outer metatarsal tubercle; supernumerary plantar tubercles scarce, low, subarticular tubercles larger than supernumerary tubercles, of shape round and projecting; (12) toes with broad lateral fringes; webbing absent; Toe V much longer than Toe III (reaching the base of the distal subarticular tubercle of Toe IV), discs of toes slightly smaller than those of outer fingers; (13) coloration in life highly variable, dorsally brown to yellow with reddish brown markings on green background (sometimes white, see variation); venter white with dark markings, throat brown with a white inverted triangle; flanks brown to grayish brown, with black spots; two vertical brown to black bars on the upper lips, postorbital and canthal stripe dark brown; loreal region cream with green spots; interobital blotch dark; dorsal surfaces of limbs green with reddish markings; posterior surfaces of thighs white with dark bars; anterior surfaces of thighs and groin cream with dark spots; iris reddish orange with black reticulum; (14) SVL in males 
<quantity id="4CEC9B46FF97FFB321834D64FAA7FE79" box="[1197,1360,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" metricValueMax="3.01" metricValueMin="2.31" pageId="2" pageNumber="39" unit="mm" value="26.6" valueMax="30.1" valueMin="23.1">23.1–30.1 mm</quantity>
(mean = 26.4 ± 2.5, n = 5), in a single female 
<quantity id="4CEC9B46FF97FFB327654D9FFD45FE52" box="[587,690,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="2" pageNumber="39" unit="mm" value="37.3">37.3 mm</quantity>
.
</paragraph>
<caption id="DF6B662BFF97FFB325B94BABFA88F82B" httpUri="https://zenodo.org/record/279302/files/figure.png" pageId="2" pageNumber="39" targetBox="[181,1405,470,1922]" targetPageId="2">
<paragraph id="8BAB36A3FF97FFB325B94BABFA88F82B" blockId="2.[151,1436,1952,2005]" pageId="2" pageNumber="39">
<emphasis id="B960EAB1FF97FFB325B94BABFEE4F848" bold="true" box="[151,275,1952,1975]" pageId="2" pageNumber="39">FIGURE 1.</emphasis>
(A): Holotype of 
<taxonomicName id="4C144D20FF97FFB324E74BA9FC86F849" authority="Boulenger 1908" authorityName="Boulenger" authorityYear="1908" box="[457,881,1952,1975]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="B960EAB1FF97FFB324E74BA9FD41F848" box="[457,694,1952,1975]" italics="true" pageId="2" pageNumber="39">Pristimantis calcaratus</emphasis>
(Boulenger 1908)
</taxonomicName>
, adult male, BMNH 1947.2.15.53, SVL 17.6 mm. (B): Dorsolateral and ventral view, and drawings of snout shape in dorsal and lateral view (original drawings by John D. Lynch).
</paragraph>
</caption>
<caption id="DF6B662BFF96FFB225B94FABFDF6FBEC" httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40" targetBox="[151,1436,193,906]" targetPageId="3">
<paragraph id="8BAB36A3FF96FFB225B94FABFDF6FBEC" blockId="3.[151,1437,928,1042]" pageId="3" pageNumber="40">
<emphasis id="B960EAB1FF96FFB225B94FABFEE5FC48" bold="true" box="[151,274,928,951]" pageId="3" pageNumber="40">FIGURE 2.</emphasis>
Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of 
<taxonomicName id="4C144D20FF96FFB221E04FA9FA6BFC49" box="[1230,1436,930,951]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF96FFB221E04FA9FA6BFC49" box="[1230,1436,930,951]" italics="true" pageId="3" pageNumber="40">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF96FFB225B94FCBFF11FC2B" box="[151,230,960,981]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
(ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of 
<taxonomicName id="4C144D20FF96FFB226AC4FCBFB98FC2B" box="[898,1135,959,981]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="B960EAB1FF96FFB226AC4FCBFB98FC2B" box="[898,1135,959,981]" italics="true" pageId="3" pageNumber="40">Pristimantis calcaratus</emphasis>
</taxonomicName>
(ICN 36940, SVL 27.7 mm). (C): Holotype of 
<taxonomicName id="4C144D20FF96FFB224684FD4FDD6FC0A" box="[326,545,990,1012]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="kelephus">
<emphasis id="B960EAB1FF96FFB224684FD4FDD6FC0A" box="[326,545,990,1012]" italics="true" pageId="3" pageNumber="40">Pristimantis kelephus</emphasis>
</taxonomicName>
(ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of 
<taxonomicName id="4C144D20FF96FFB221BB4FD4FA95FC0A" box="[1173,1378,990,1012]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="jubatus">
<emphasis id="B960EAB1FF96FFB221BB4FD4FA95FC0A" box="[1173,1378,990,1012]" italics="true" pageId="3" pageNumber="40">Pristimantis jubatus</emphasis>
</taxonomicName>
(ICN 52478, Adult male, SVL 22.1 mm).
</paragraph>
</caption>
<paragraph id="8BAB36A3FF96FFB225EB484BFB31F8D9" blockId="3.[151,1436,1087,2011]" pageId="3" pageNumber="40">
<taxonomicName id="4C144D20FF96FFB225EB484BFE5CFBA9" box="[197,427,1088,1111]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="quicato">
<emphasis id="B960EAB1FF96FFB225EB484BFE5CFBA9" box="[197,427,1088,1111]" italics="true" pageId="3" pageNumber="40">Pristimantis quicato</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C144D20FF96FFB2276B484BFD2EFBA9" box="[581,729,1087,1111]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="B960EAB1FF96FFB2276B484BFD2EFBA9" box="[581,729,1087,1111]" italics="true" pageId="3" pageNumber="40">P. calcaratus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D20FF96FFB227C7484BFC9EFBA9" box="[745,873,1087,1111]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="kelephus">
<emphasis id="B960EAB1FF96FFB227C7484BFC9EFBA9" box="[745,873,1087,1111]" italics="true" pageId="3" pageNumber="40">P. kelephus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C144D20FF96FFB2268D484BFBE3FBA9" box="[931,1044,1087,1111]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="jubatus">
<emphasis id="B960EAB1FF96FFB2268D484BFBE3FBA9" box="[931,1044,1087,1111]" italics="true" pageId="3" pageNumber="40">P. jubatus</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A26FF96FFB2210A4834FB7FFBA9" box="[1060,1160,1087,1111]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
, 
<figureCitation id="132F2A26FF96FFB221B84834FB52FBA9" box="[1174,1189,1087,1111]" captionStart="FIGURE 3" captionStartId="4.[151,250,1931,1954]" captionTargetBox="[151,1436,193,1908]" captionTargetId="figure@4.[151,1436,193,1910]" captionTargetPageId="4" captionText="FIGURE 3. Relationship in length Toe IV vs Toe V. (A): Holotype of Pristimantis quicato sp. nov. (ICN 55409, SVL 37 mm). (B): Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279304/files/figure.png" pageId="3" pageNumber="40">3</figureCitation>
, 
<figureCitation id="132F2A26FF96FFB2219A4834FB34FBA9" box="[1204,1219,1087,1111]" captionStart="FIGURE 5" captionStartId="6.[151,250,1349,1372]" captionTargetBox="[151,1436,481,1328]" captionTargetId="figure@6.[151,1436,481,1328]" captionTargetPageId="6" captionText="FIGURE 5. (A): Pristimantis quicato sp. nov. (ICN 55410, paratype, adult male, SVL 30.1 mm); (B): Pristimantis calcaratus (not collected, Reserva Forestal Bitaco, Valle del Cauca); (C): Pristimantis jubatus (not collected, PNN Munchique, Cauca, Photo: J. C. García); (D): Pristimantis kelephus (not collected, Serranía de los Paraguas, Valle del Cauca, Photo: Wilmar Bolívar-G)." httpUri="https://zenodo.org/record/279306/files/figure.png" pageId="3" pageNumber="40">5</figureCitation>
, 
<figureCitation id="132F2A26FF96FFB221FC4834FB15FBA9" box="[1234,1250,1087,1111]" captionStart="FIGURE 7" captionStartId="9.[151,250,1714,1737]" captionTargetBox="[151,1436,842,1692]" captionTargetId="figure@9.[151,1436,842,1693]" captionTargetPageId="9" captionText="FIGURE 7. Ornamentation of frontoparietals in: (A) Pristimantis quicato sp. nov. (ICN 55409, SVL 37 mm); (B) P. jubatus (ICN 52482, SVL 33.3 mm) and (C) P. kelephus (ICN 39639, SVL 30 mm). (D) Ornamentation of frontoparietals is absent in P. calcaratus (ICN 39640, SVL 27.2 mm)." httpUri="https://zenodo.org/record/279308/files/figure.png" pageId="3" pageNumber="40">7</figureCitation>
). It differs from those species by a combination of traits that are described as follows (characters in parenthesis for 
<taxonomicName id="4C144D20FF96FFB221F0486EFABAFB82" box="[1246,1357,1125,1148]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="quicato">
<emphasis id="B960EAB1FF96FFB221F0486EFABAFB82" box="[1246,1357,1125,1148]" italics="true" pageId="3" pageNumber="40">P. quicato</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C144D20FF96FFB2204A486EFE99FB61" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="B960EAB1FF96FFB2204A486EFE99FB61" italics="true" pageId="3" pageNumber="40">Pristimantis calcaratus</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A26FF96FFB22452488CFE2BFB61" box="[380,476,1159,1183]" captionStart="FIGURE 1" captionStartId="2.[151,250,1952,1975]" captionTargetBox="[181,1405,470,1922]" captionTargetId="figure@2.[181,1405,461,1922]" captionTargetPageId="2" captionText="FIGURE 1. (A): Holotype of Pristimantis calcaratus (Boulenger 1908), adult male, BMNH 1947.2. 15.53, SVL 17.6 mm. (B): Dorsolateral and ventral view, and drawings of snout shape in dorsal and lateral view (original drawings by John D. Lynch)." httpUri="https://zenodo.org/record/279302/files/figure.png" pageId="3" pageNumber="40">Figure 1</figureCitation>
, 
<figureCitation id="132F2A26FF96FFB224C4488CFE0EFB61" box="[490,505,1159,1183]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">2</figureCitation>
B, 3B, 5B, 7D) has short vocal slits in males (absent), lacks cranial crests (present), and is smaller: snout-vent length 19.0– 
<quantity id="4CEC9B46FF96FFB2276248A0FD47FB3D" box="[588,688,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="3" pageNumber="40" unit="mm" value="23.8">23.8 mm</quantity>
in adult males (n=18), 
<quantity id="4CEC9B46FF96FFB2269948A0FBADFB3D" box="[951,1114,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="2.945" metricValueMax="3.12" metricValueMin="2.77" pageId="3" pageNumber="40" unit="mm" value="29.45" valueMax="31.2" valueMin="27.7">27.7–31.2 mm</quantity>
in females (n=3) (
<quantity id="4CEC9B46FF96FFB2200448A0FF34FB19" metricMagnitude="-2" metricUnit="m" metricValue="2.66" metricValueMax="3.01" metricValueMin="2.31" pageId="3" pageNumber="40" unit="mm" value="26.6" valueMax="30.1" valueMin="23.1">23.1–30.1 mm</quantity>
in adult males and 
<quantity id="4CEC9B46FF96FFB224B448C4FE0AFB19" box="[410,509,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="3" pageNumber="40" unit="mm" value="37.3">37.3 mm</quantity>
in one female). 
<taxonomicName id="4C144D20FF96FFB2279E48DBFC55FB19" box="[688,930,1231,1255]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="kelephus">
<emphasis id="B960EAB1FF96FFB2279E48DBFC55FB19" box="[688,930,1231,1255]" italics="true" pageId="3" pageNumber="40">Pristimantis kelephus</emphasis>
</taxonomicName>
has the dorsal skin with many round pustules forming a distinct pattern above shoulders and transverse ridges above sacrum (the tubercles are distributed uniformly across the dorsum and forming ridges only on shoulder) (
<figureCitation id="132F2A26FF96FFB226A4491CFC1AFAD1" box="[906,1005,1303,1327]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
A, 2C); the venter is mostly black or brown with white spots (venter white with brown markings), and it lacks pale spots or bars on the posterior surfaces of the thighs (posterior surfaces of thighs white with dark bars) (
<figureCitation id="132F2A26FF96FFB226804954FBF9FA89" box="[942,1038,1375,1399]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
A, 2C). Also, 
<taxonomicName id="4C144D20FF96FFB22183496BFADCFA89" box="[1197,1323,1375,1399]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="kelephus">
<emphasis id="B960EAB1FF96FFB22183496BFADCFA89" box="[1197,1323,1375,1399]" italics="true" pageId="3" pageNumber="40">P. kelephus</emphasis>
</taxonomicName>
has vocal slits (absent), the snout shape is round in dorsal view (subacuminate in males) (
<figureCitation id="132F2A26FF96FFB221374988FB8CFA65" box="[1049,1147,1411,1435]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
C, 3C, 5D, 7C), the distal tip of Toe V reaches between the penultimate and distal subarticular tubercle of Toe IV (the distal end of Toe V reaches the distal subarticular tubercle of Toe IV) (
<figureCitation id="132F2A26FF96FFB227F949C0FCC0FA1D" box="[727,823,1483,1507]" captionStart="FIGURE 3" captionStartId="4.[151,250,1931,1954]" captionTargetBox="[151,1436,193,1908]" captionTargetId="figure@4.[151,1436,193,1910]" captionTargetPageId="4" captionText="FIGURE 3. Relationship in length Toe IV vs Toe V. (A): Holotype of Pristimantis quicato sp. nov. (ICN 55409, SVL 37 mm). (B): Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279304/files/figure.png" pageId="3" pageNumber="40">Figure 3</figureCitation>
C), and is smaller in size: snout-vent length 
<quantity id="4CEC9B46FF96FFB2200249C0FF34F9F9" metricMagnitude="-2" metricUnit="m" metricValue="2.035" metricValueMax="2.19" metricValueMin="1.88" pageId="3" pageNumber="40" unit="mm" value="20.35" valueMax="21.9" valueMin="18.8">18.8–21.9 mm</quantity>
in adult males (n=22), 
<quantity id="4CEC9B46FF96FFB224FC49E4FD80F9F9" box="[466,631,1519,1543]" metricMagnitude="-2" metricUnit="m" metricValue="2.895" metricValueMax="3.02" metricValueMin="2.77" pageId="3" pageNumber="40" unit="mm" value="28.95" valueMax="30.2" valueMin="27.7">27.7–30.2 mm</quantity>
in females (n=16) (
<quantity id="4CEC9B46FF96FFB2267249E4FBF5F9F9" box="[860,1026,1519,1543]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" metricValueMax="3.01" metricValueMin="2.31" pageId="3" pageNumber="40" unit="mm" value="26.6" valueMax="30.1" valueMin="23.1">23.1–30.1 mm</quantity>
in adult males and 
<quantity id="4CEC9B46FF96FFB221C849E4FABBF9F9" box="[1254,1356,1519,1543]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="3" pageNumber="40" unit="mm" value="37.3">37.3 mm</quantity>
in one female). 
<taxonomicName id="4C144D20FF96FFB225D54A1EFE29F9D2" box="[251,478,1556,1580]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="jubatus">
<emphasis id="B960EAB1FF96FFB225D54A1EFE29F9D2" box="[251,478,1556,1580]" italics="true" pageId="3" pageNumber="40">Pristimantis jubatus</emphasis>
</taxonomicName>
has pustules above the sacrum, while the rest of the dorsum is smooth, and has subconical tubercles scattered on head and close tympanum (tubercles are uniformly distributed across the dorsum and form ridges only on shoulder) (
<figureCitation id="132F2A26FF96FFB224D94A50FDAFF98D" box="[503,600,1627,1651]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
A, 2D); it lacks sub-anal tubercles and a conical interocular tubercle (both are present) (
<figureCitation id="132F2A26FF96FFB224074A74FE7DF969" box="[297,394,1663,1687]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
A, 2D, 5A, 5C); the snout shape is round in both dorsal and lateral profiles (snout subacuminate in dorsal view) (
<figureCitation id="132F2A26FF96FFB224AC4AA8FE14F945" box="[386,483,1699,1723]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
D, 3D, 5C, 7B); the distal tip of Toe V reaches the point between the penultimate and distal subarticular tubercle on Toe IV (the distal end of Toe V reaches the distal subarticular tubercles of Toe IV) (
<figureCitation id="132F2A26FF96FFB225B04AE0FF08F8FD" box="[158,255,1771,1795]" captionStart="FIGURE 3" captionStartId="4.[151,250,1931,1954]" captionTargetBox="[151,1436,193,1908]" captionTargetId="figure@4.[151,1436,193,1910]" captionTargetPageId="4" captionText="FIGURE 3. Relationship in length Toe IV vs Toe V. (A): Holotype of Pristimantis quicato sp. nov. (ICN 55409, SVL 37 mm). (B): Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279304/files/figure.png" pageId="3" pageNumber="40">Figure 3</figureCitation>
D); and adult males and females are slightly smaller in snout-vent length, 
<quantity id="4CEC9B46FF96FFB221644AE0FB19F8FD" box="[1098,1262,1771,1795]" metricMagnitude="-2" metricUnit="m" metricValue="2.195" metricValueMax="2.41" metricValueMin="1.98" pageId="3" pageNumber="40" unit="mm" value="21.95" valueMax="24.1" valueMin="19.8">19.8–24.1 mm</quantity>
in adult males, 
<quantity id="4CEC9B46FF96FFB225B94B04FECDF8D9" box="[151,314,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="3.46" metricValueMin="3.2399999999999998" pageId="3" pageNumber="40" unit="mm" value="33.5" valueMax="34.6" valueMin="32.4">32.4–34.6 mm</quantity>
in females (SVL in adult males 
<quantity id="4CEC9B46FF96FFB2278A4B04FCB0F8D9" box="[676,839,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" metricValueMax="3.01" metricValueMin="2.31" pageId="3" pageNumber="40" unit="mm" value="26.6" valueMax="30.1" valueMin="23.1">23.1–30.1 mm</quantity>
and in a single female 
<quantity id="4CEC9B46FF96FFB221644B04FB42F8D9" box="[1098,1205,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="3" pageNumber="40" unit="mm" value="37.3">37.3 mm</quantity>
.).
</paragraph>
</subSubSection>
<subSubSection id="C30E6528FF96FFB625EB4B38FE01F902" lastPageId="7" lastPageNumber="44" pageId="3" pageNumber="40" type="description">
<paragraph id="8BAB36A3FF96FFB725EB4B38FC6FFE52" blockId="3.[151,1436,1087,2011]" lastBlockId="6.[151,1436,151,428]" lastPageId="6" lastPageNumber="43" pageId="3" pageNumber="40">
<emphasis id="B960EAB1FF96FFB225EB4B38FCFDF8B2" bold="true" box="[197,778,1843,1868]" pageId="3" pageNumber="40">
Description of the 
<typeStatus id="54AF8801FF96FFB224864B3FFDF8F8B2" box="[424,527,1844,1868]" pageId="3" pageNumber="40" type="holotype">holotype</typeStatus>
(
<figureCitation id="132F2A26FF96FFB227314B38FD70F8B2" box="[543,647,1843,1868]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[151,1436,193,906]" captionTargetId="figure@3.[151,1436,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral and ventral view of the species mentioned in the text. (A): Preserved holotype of Pristimantis quicato sp. nov. (ICN 55409, Adult female, SVL 37.0 mm). (B): Adult female of Pristimantis calcaratus (ICN 36940, SVL 27.7 mm). (C): Holotype of Pristimantis kelephus (ICN 39637, Adult female, SVL 30.2 mm). (D): Holotype of Pristimantis jubatus (ICN 52478, Adult male, SVL 22.1 mm)." httpUri="https://zenodo.org/record/279303/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
A, 3A, 7A)
</emphasis>
. An adult female, 
<quantity id="4CEC9B46FF96FFB226CB4B38FBBCF8B5" box="[997,1099,1843,1867]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="3" pageNumber="40" unit="mm" value="37.3">37.3 mm</quantity>
SVL. Head as wide as body, head width 41% of SVL; slightly wider than long, head length 35.9% of SVL. Snout slightly longer, rounded in dorsal and lateral view; canthus rostralis slightly concave; loreal region concave with small tubercles; lips not flared; nostrils protuberant, directed laterally; internarial distance 72.3% of eye-to-nostril distance; eye-to-nostril distance 102.1% of eye-length. Upper eyelid width slightly narrower than IOD, 50.9% of IOD; upper eyelid with small tubercles, and one large, conical tubercle on each eyelid, and low cranial crest. Tympanic annulus evident, its length 41.3% of eye diameter; supratympanic fold originating at corner of eye, encroaching on the top edge of the tympanic membrane; postrictal tubercles conical. Choanae small, oval and partly concealed by palatal shelf of maxilla; dentigerous processes of vomers slightly larger than choanae, oblique, positioned posterior to level of choanae and separated medially by distance less than width of odontophore, each process bearing 7 teeth; tongue longer than wide, posterior border notched, posterior 2/3 free. Skin on dorsum granular with some conical tubercles distributed uniformly, and forming ridges (H-shaped) on shoulders; dorsolateral folds on anterior half of body; ventral skin areolate, discoidal fold evident, throat like venter but with a diffuse black inverted triangle; skin on flanks areolate; cloacal sheath and tubercles present; ulnar tubercles present, not forming a distinct fold; thenar tubercle oval, smaller than divided palmar tubercles; supernumerary palmar tubercles present; subarticular tubercles prominent; fingers bearing lateral keels; Finger I shorter than Finger II; tips of digits round to subtruncated; inner digits bearing much narrower discs; all fingers bearing pads on digital tips.
</paragraph>
<caption id="DF6B662BFF91FFB525B94B80FCB7F821" httpUri="https://zenodo.org/record/279304/files/figure.png" pageId="4" pageNumber="41" targetBox="[151,1436,193,1908]" targetPageId="4">
<paragraph id="8BAB36A3FF91FFB525B94B80FCB7F821" blockId="4.[151,1437,1931,2015]" pageId="4" pageNumber="41">
<emphasis id="B960EAB1FF91FFB525B94B80FEE5F85F" bold="true" box="[151,274,1931,1954]" pageId="4" pageNumber="41">FIGURE 3.</emphasis>
Relationship in length Toe IV vs Toe V. (A): Holotype of 
<taxonomicName id="4C144D20FF91FFB526714B86FBDAF85C" box="[863,1069,1933,1954]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF91FFB526714B86FBDAF85C" box="[863,1069,1933,1954]" italics="true" pageId="4" pageNumber="41">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF91FFB5211D4B87FB73F85F" box="[1075,1156,1932,1953]" pageId="4" pageNumber="41" rank="species">
<emphasis id="B960EAB1FF91FFB5211D4B87FB73F85F" bold="true" box="[1075,1156,1932,1953]" pageId="4" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
(ICN 55409, SVL 37 mm). (B): 
<taxonomicName id="4C144D20FF91FFB525E84BA0FE44F83E" box="[198,435,1962,1984]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="B960EAB1FF91FFB525E84BA0FE44F83E" box="[198,435,1962,1984]" italics="true" pageId="4" pageNumber="41">Pristimantis calcaratus</emphasis>
</taxonomicName>
(ICN 36940, SVL 27.7 mm). (C): Holotype of 
<taxonomicName id="4C144D20FF91FFB526BB4BA0FB98F83E" box="[917,1135,1962,1984]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="kelephus">
<emphasis id="B960EAB1FF91FFB526BB4BA0FB98F83E" box="[917,1135,1962,1984]" italics="true" pageId="4" pageNumber="41">Pristimantis kelephus</emphasis>
</taxonomicName>
(ICN 39637, SVL 30.2 mm). (D): Holotype of 
<taxonomicName id="4C144D20FF91FFB524694BC1FDE3F821" box="[327,532,1993,2015]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="jubatus">
<emphasis id="B960EAB1FF91FFB524694BC1FDE3F821" box="[327,532,1993,2015]" italics="true" pageId="4" pageNumber="41">Pristimantis jubatus</emphasis>
</taxonomicName>
(ICN 52478, SVL 22.1 mm).
</paragraph>
</caption>
<caption id="DF6B662BFF90FFB425B94ACAFEE8F8CA" httpUri="https://zenodo.org/record/279305/files/figure.png" pageId="5" pageNumber="42" targetBox="[151,1436,193,1708]" targetPageId="5">
<paragraph id="8BAB36A3FF90FFB425B94ACAFEE8F8CA" blockId="5.[151,1437,1729,1844]" pageId="5" pageNumber="42">
<emphasis id="B960EAB1FF90FFB425B94ACAFEE4F929" bold="true" box="[151,275,1729,1752]" pageId="5" pageNumber="42">FIGURE 4.</emphasis>
<taxonomicName id="4C144D20FF90FFB424344AC8FE1EF926" box="[282,489,1731,1752]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF90FFB424344AC8FE1EF926" box="[282,489,1731,1752]" italics="true" pageId="5" pageNumber="42">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF90FFB424DE4AC9FDB6F926" box="[496,577,1730,1752]" pageId="5" pageNumber="42" rank="species">
<emphasis id="B960EAB1FF90FFB424DE4AC9FDCCF929" bold="true" box="[496,571,1730,1751]" pageId="5" pageNumber="42">sp. nov</emphasis>
.
</taxonomicNameLabel>
Intrapopulation variation in dorsal coloration in preservative (all adult males). (A): ICN 55406, SVL 26.0 mm. (B): ICN 55411, SVL 23.1 mm. (C): ICN 55408, SVL 26.7 mm. (D) ICN 55410, SVL 30.1 mm. (E) ICN 55407, SVL 26.1 mm. (F-G): Polymorphic patterns, “Head Cap” (ICN 55406) and Dorsoconcolor + Vertebral Dark Line (ICN 55411).
</paragraph>
</caption>
<caption id="DF6B662BFF93FFB725B9494EFF16FA46" httpUri="https://zenodo.org/record/279306/files/figure.png" pageId="6" pageNumber="43" targetBox="[151,1436,481,1328]" targetPageId="6">
<paragraph id="8BAB36A3FF93FFB725B9494EFF16FA46" blockId="6.[151,1436,1349,1464]" pageId="6" pageNumber="43">
<emphasis id="B960EAB1FF93FFB725B9494EFEE4FAA5" bold="true" box="[151,275,1349,1372]" pageId="6" pageNumber="43">FIGURE 5.</emphasis>
(A): 
<taxonomicName id="4C144D20FF93FFB72464494CFDEFFAA2" box="[330,536,1351,1372]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF93FFB72464494CFDEFFAA2" box="[330,536,1351,1372]" italics="true" pageId="6" pageNumber="43">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF93FFB72731494DFD87FAA2" box="[543,624,1350,1372]" pageId="6" pageNumber="43" rank="species">
<emphasis id="B960EAB1FF93FFB72731494DFD9DFAA5" bold="true" box="[543,618,1350,1371]" pageId="6" pageNumber="43">sp. nov</emphasis>
.
</taxonomicNameLabel>
(ICN 55410, paratype, adult male, SVL 30.1 mm); (B): 
<taxonomicName id="4C144D20FF93FFB72181494CFA6BFAA2" box="[1199,1436,1350,1372]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="B960EAB1FF93FFB72181494CFA6BFAA2" box="[1199,1436,1350,1372]" italics="true" pageId="6" pageNumber="43">Pristimantis calcaratus</emphasis>
</taxonomicName>
(not collected, Reserva Forestal Bitaco, Valle del Cauca); (C): 
<taxonomicName id="4C144D20FF93FFB7260B496DFC02FA85" box="[805,1013,1381,1403]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="jubatus">
<emphasis id="B960EAB1FF93FFB7260B496DFC02FA85" box="[805,1013,1381,1403]" italics="true" pageId="6" pageNumber="43">Pristimantis jubatus</emphasis>
</taxonomicName>
(not collected, PNN Munchique, Cauca, Photo: J. C. García); (D): 
<taxonomicName id="4C144D20FF93FFB724B1498FFD8DFA67" box="[415,634,1411,1433]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="kelephus">
<emphasis id="B960EAB1FF93FFB724B1498FFD8DFA67" box="[415,634,1411,1433]" italics="true" pageId="6" pageNumber="43">Pristimantis kelephus</emphasis>
</taxonomicName>
(not collected, Serranía de los Paraguas, Valle del Cauca, Photo: Wilmar Bolívar-G).
</paragraph>
</caption>
<paragraph id="8BAB36A3FF93FFB725EB49EFFBDEF906" blockId="6.[151,1436,1508,2037]" pageId="6" pageNumber="43">
<collectingCountry id="F3037633FF93FFB725EB49EFFF0AFA02" box="[197,253,1508,1532]" name="India" pageId="6" pageNumber="43">Hind</collectingCountry>
limbs moderate, heels overlap when held at right angles to sagittal plane; tibia long 59.5% of SVL; foot long 58% of SVL. Relative length of adpressed toes IV&gt; V&gt; III&gt; II&gt; I; tip of Toe V extending to base of distal subarticular tubercle of Toe IV; tip of Toe III extending to distal border of penultimate subarticular tubercle on Toe IV; discs of toes slightly smaller to those on outer fingers, bearing broad lateral fringes; webbing absent. Heel bearing small tubercles, and one large, conical tubercle; subconical tubercles along outer border of tarsus, inner edge of tarsus bearing a tubercle on distal third of tarsus; indistinct inner tarsal fold on distal 1/4 of tarsus; inner metatarsal tubercle twice as long as wide (six times larger than the subconical outer metatarsal tubercle); few supernumerary plantar tubercles present, at bases of toes II–V; protuberant subarticular tubercles.
</paragraph>
<paragraph id="8BAB36A3FF93FFB625E94B0FFC06FE31" blockId="6.[151,1436,1508,2037]" lastBlockId="7.[151,1436,151,463]" lastPageId="7" lastPageNumber="44" pageId="6" pageNumber="43">
<emphasis id="B960EAB1FF93FFB725E94B0FFECCF8E3" bold="true" box="[199,315,1796,1821]" pageId="6" pageNumber="43">Variation.</emphasis>
Males are smaller than females; measurements and proportions are presented in 
<tableCitation id="C6960318FF93FFB721F84B0FFADCF8E2" box="[1238,1323,1796,1820]" captionStart="TABLE 1" captionStartId="7.[151,235,512,533]" captionTargetBox="[151,1436,597,1404]" captionTargetPageId="7" captionText="TABLE 1. Morphometrics and proportions (in percentages) of Pristimantis quicato sp. nov. Mean and standard deviation in parentheses follow ranges." httpUri="http://table.plazi.org/id/DF6B662BFF92FFB625B94E0BFE52FDC7" pageId="6" pageNumber="43" tableUuid="DF6B662BFF92FFB625B94E0BFE52FDC7">Table 1</tableCitation>
. The skin on the dorsum and upper surfaces of the limbs have conical tubercles, which are usually smaller on the limbs than on the dorsum of the body in females, and are uniformly distributed in males. Juveniles differ from conspecific adults by having more tubercles, but together with adult present pustules in the skin on flanks and posterior surfaces of shanks. Cranial crest with dermal ridge (ICN 55406, ICN 55408, 
<figureCitation id="132F2A26FF93FFB726FF4B9FFBC7F852" box="[977,1072,1940,1964]" captionStart="FIGURE 4" captionStartId="5.[151,250,1729,1752]" captionTargetBox="[151,1436,193,1708]" captionTargetId="figure@5.[151,1436,193,1709]" captionTargetPageId="5" captionText="FIGURE 4. Pristimantis quicato sp. nov. Intrapopulation variation in dorsal coloration in preservative (all adult males). (A): ICN 55406, SVL 26.0 mm. (B): ICN 55411, SVL 23.1 mm. (C): ICN 55408, SVL 26.7 mm. (D) ICN 55410, SVL 30.1 mm. (E) ICN 55407, SVL 26.1 mm. (F-G): Polymorphic patterns, “ Head Cap ” (ICN 55406) and Dorsoconcolor + Vertebral Dark Line (ICN 55411)." httpUri="https://zenodo.org/record/279305/files/figure.png" pageId="6" pageNumber="43">Figure 4</figureCitation>
A, 4C) and without dermal ridge (ICN 55407, ICN 55410–11, 
<figureCitation id="132F2A26FF93FFB724F04BB3FDCAF82E" box="[478,573,1976,2000]" captionStart="FIGURE 4" captionStartId="5.[151,250,1729,1752]" captionTargetBox="[151,1436,193,1708]" captionTargetId="figure@5.[151,1436,193,1709]" captionTargetPageId="5" captionText="FIGURE 4. Pristimantis quicato sp. nov. Intrapopulation variation in dorsal coloration in preservative (all adult males). (A): ICN 55406, SVL 26.0 mm. (B): ICN 55411, SVL 23.1 mm. (C): ICN 55408, SVL 26.7 mm. (D) ICN 55410, SVL 30.1 mm. (E) ICN 55407, SVL 26.1 mm. (F-G): Polymorphic patterns, “ Head Cap ” (ICN 55406) and Dorsoconcolor + Vertebral Dark Line (ICN 55411)." httpUri="https://zenodo.org/record/279305/files/figure.png" pageId="6" pageNumber="43">Figure 4</figureCitation>
B, 4D, 4E). The coloration in 
<taxonomicName id="4C144D20FF93FFB726A64BB1FB9BF82F" box="[904,1132,1978,2001]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="quicato">
<emphasis id="B960EAB1FF93FFB726A64BB1FB9BF82F" box="[904,1132,1978,2001]" italics="true" pageId="6" pageNumber="43">Pristimantis quicato</emphasis>
</taxonomicName>
is highly variable, presenting various shades of brown or gray dorsally with diffuse dark marks, which consist of spots and longitudinal stripes (ICN 55407–08). This species has also two patterns of polymorphisms according to the scheme proposed by 
<bibRefCitation id="EF854B52FF92FFB625B94CB7FE62FF2A" author="Carrillo-Rivera" box="[151,405,188,212]" pageId="7" pageNumber="44" refString="Carrillo-Rivera, N. (2006) Patrones polimorficos en especies Colombianas del genero Eleutherodactylus (Amphibia: Leptodactylidae). Tesis de pregrado Universidad de Antioquia, 67 pp." type="book" year="2006">Carrillo-Rivera (2006)</bibRefCitation>
for 
<taxonomicName id="4C144D20FF92FFB624EA4CB7FD79FF2A" box="[452,654,188,212]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B960EAB1FF92FFB624EA4CB7FD79FF2A" box="[452,654,188,212]" italics="true" pageId="7" pageNumber="44">Eleutherodactylus</emphasis>
</taxonomicName>
: (1) “Head Cap” (ICN 55406), which describes a dark area that covers the head and ends up forming a A or a inverted U shape on the back, the rest of the body white with brown spots (
<figureCitation id="132F2A26FF92FFB625B04D0FFF09FEE2" box="[158,254,260,284]" captionStart="FIGURE 4" captionStartId="5.[151,250,1729,1752]" captionTargetBox="[151,1436,193,1708]" captionTargetId="figure@5.[151,1436,193,1709]" captionTargetPageId="5" captionText="FIGURE 4. Pristimantis quicato sp. nov. Intrapopulation variation in dorsal coloration in preservative (all adult males). (A): ICN 55406, SVL 26.0 mm. (B): ICN 55411, SVL 23.1 mm. (C): ICN 55408, SVL 26.7 mm. (D) ICN 55410, SVL 30.1 mm. (E) ICN 55407, SVL 26.1 mm. (F-G): Polymorphic patterns, “ Head Cap ” (ICN 55406) and Dorsoconcolor + Vertebral Dark Line (ICN 55411)." httpUri="https://zenodo.org/record/279305/files/figure.png" pageId="7" pageNumber="44">Figure 4</figureCitation>
F) and (2) Dorsoconcolor + Vertebral Dark Line (ICN 55411), where the dorsal surface of the snout, eyelids, head, and dorsum, varies from cream to brown tones differing from the rest of the darker body (
<figureCitation id="132F2A26FF92FFB620384D2CFA8FFEC1" box="[1302,1400,295,319]" captionStart="FIGURE 4" captionStartId="5.[151,250,1729,1752]" captionTargetBox="[151,1436,193,1708]" captionTargetId="figure@5.[151,1436,193,1709]" captionTargetPageId="5" captionText="FIGURE 4. Pristimantis quicato sp. nov. Intrapopulation variation in dorsal coloration in preservative (all adult males). (A): ICN 55406, SVL 26.0 mm. (B): ICN 55411, SVL 23.1 mm. (C): ICN 55408, SVL 26.7 mm. (D) ICN 55410, SVL 30.1 mm. (E) ICN 55407, SVL 26.1 mm. (F-G): Polymorphic patterns, “ Head Cap ” (ICN 55406) and Dorsoconcolor + Vertebral Dark Line (ICN 55411)." httpUri="https://zenodo.org/record/279305/files/figure.png" pageId="7" pageNumber="44">Figure 4</figureCitation>
G). 
<taxonomicName id="4C144D20FF92FFB625B94D46FE8DFE9A" box="[151,378,333,356]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="quicato">
<emphasis id="B960EAB1FF92FFB625B94D46FE8DFE9A" box="[151,378,333,356]" italics="true" pageId="7" pageNumber="44">Pristimantis quicato</emphasis>
</taxonomicName>
presents variation in size and pigmentation of testis: two adult males had testis of equal size to kidneys (ICN 55407, ICN 055411), and three had size approximate to 2/3 of kidneys (ICN 55406, ICN 55408, ICN 55410); for juveniles and subadults, testis size was ranged between 1/3–1/2 of that of kidneys (UV–CD 
<date id="FFAA1063FF92FFB620074D9FFA67FE52" box="[1321,1424,404,428]" pageId="7" pageNumber="44" value="1917" valueMax="1924">1917–24</date>
); only one adult male was found with melanophores in a testicle (ICN 55410).
</paragraph>
<caption id="DF6B662BFF92FFB625B94E0BFE52FDC7" ID-Table-UUID="DF6B662BFF92FFB625B94E0BFE52FDC7" httpUri="http://table.plazi.org/id/DF6B662BFF92FFB625B94E0BFE52FDC7" pageId="7" pageNumber="44" targetBox="[151,1436,597,1404]" targetIsTable="true" targetPageId="7">
<paragraph id="8BAB36A3FF92FFB625B94E0BFE52FDC7" blockId="7.[151,1436,512,569]" pageId="7" pageNumber="44">
<emphasis id="B960EAB1FF92FFB625B94E0BFEF1FDEB" bold="true" box="[151,262,512,533]" pageId="7" pageNumber="44">TABLE 1.</emphasis>
Morphometrics and proportions (in percentages) of 
<taxonomicName id="4C144D20FF92FFB6260A4E0AFC03FDE8" box="[804,1012,513,534]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF92FFB6260A4E0AFC03FDE8" box="[804,1012,513,534]" italics="true" pageId="7" pageNumber="44">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF92FFB626D24E0BFBB9FDE8" box="[1020,1102,512,534]" pageId="7" pageNumber="44" rank="species">
<emphasis id="B960EAB1FF92FFB626D24E0BFBBFFDEB" bold="true" box="[1020,1096,512,533]" pageId="7" pageNumber="44">sp. nov</emphasis>
.
</taxonomicNameLabel>
Mean and standard deviation in parentheses follow ranges.
</paragraph>
</caption>
<paragraph id="8BAB36A3FF92FFB6275C4E5EFBC1FA82" pageId="7" pageNumber="44">
<table id="F914C403FF92004E25B94E5EFA6BFA82" box="[151,1436,597,1404]" gridcols="3" gridrows="20" pageId="7" pageNumber="44">
<tr id="352434E1FF92004E25B94E5EFA6BFD95" box="[151,1436,597,619]" gridrow="0" pageId="7" pageNumber="44" rowspan-0="1">
<th id="76F55D9DFF92004E275C4E5EFD4AFD95" box="[626,701,597,619]" gridcol="1" gridrow="0" pageId="7" pageNumber="44">Female</th>
<th id="76F55D9DFF92004E26FF4E5EFA6BFD95" box="[977,1436,597,619]" gridcol="2" gridrow="0" pageId="7" pageNumber="44">Males</th>
</tr>
<tr id="352434E1FF92004E25B94E75FA6BFD6A" box="[151,1436,638,660]" gridrow="1" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94E75FE42FD6A" box="[151,437,638,660]" gridcol="0" gridrow="1" pageId="7" pageNumber="44">SVL</th>
<td id="76F55D9DFF92004E275C4E75FD4AFD6A" box="[626,701,638,660]" gridcol="1" gridrow="1" pageId="7" pageNumber="44">37.3</td>
<td id="76F55D9DFF92004E26FF4E75FA6BFD6A" box="[977,1436,638,660]" gridcol="2" gridrow="1" pageId="7" pageNumber="44">23.1–30.1 (26.4 ± 2.5)</td>
</tr>
<tr id="352434E1FF92004E25B94EA3FA6BFD40" box="[151,1436,680,702]" gridrow="2" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94EA3FE42FD40" box="[151,437,680,702]" gridcol="0" gridrow="2" pageId="7" pageNumber="44">HW</th>
<td id="76F55D9DFF92004E275C4EA3FD4AFD40" box="[626,701,680,702]" gridcol="1" gridrow="2" pageId="7" pageNumber="44">15.5</td>
<td id="76F55D9DFF92004E26FF4EA3FA6BFD40" box="[977,1436,680,702]" gridcol="2" gridrow="2" pageId="7" pageNumber="44">10–12.8 (11.2 ± 1.0)</td>
</tr>
<tr id="352434E1FF92004E25B94EDAFA6BFD16" box="[151,1436,721,744]" gridrow="3" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94EDAFE42FD16" box="[151,437,721,744]" gridcol="0" gridrow="3" pageId="7" pageNumber="44">HL</th>
<td id="76F55D9DFF92004E275C4EDAFD4AFD16" box="[626,701,721,744]" gridcol="1" gridrow="3" pageId="7" pageNumber="44">13.4</td>
<td id="76F55D9DFF92004E26FF4EDAFA6BFD16" box="[977,1436,721,744]" gridcol="2" gridrow="3" pageId="7" pageNumber="44">8.6–10.4 (9.5 ± 0.7)</td>
</tr>
<tr id="352434E1FF92004E25B94EF1FA6BFCEE" box="[151,1436,762,784]" gridrow="4" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94EF1FE42FCEE" box="[151,437,762,784]" gridcol="0" gridrow="4" pageId="7" pageNumber="44">IOD</th>
<td id="76F55D9DFF92004E275C4EF1FD4AFCEE" box="[626,701,762,784]" gridcol="1" gridrow="4" pageId="7" pageNumber="44">7.7</td>
<td id="76F55D9DFF92004E26FF4EF1FA6BFCEE" box="[977,1436,762,784]" gridcol="2" gridrow="4" pageId="7" pageNumber="44">5.2–6.4 (5.5 ± 0.5)</td>
</tr>
<tr id="352434E1FF92004E25B94F28FA6BFCC4" box="[151,1436,803,826]" gridrow="5" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94F28FE42FCC4" box="[151,437,803,826]" gridcol="0" gridrow="5" pageId="7" pageNumber="44">UEW</th>
<td id="76F55D9DFF92004E275C4F28FD4AFCC4" box="[626,701,803,826]" gridcol="1" gridrow="5" pageId="7" pageNumber="44">3.2</td>
<td id="76F55D9DFF92004E26FF4F28FA6BFCC4" box="[977,1436,803,826]" gridcol="2" gridrow="5" pageId="7" pageNumber="44">2.4–2.9 (2.6 ± 0.2)</td>
</tr>
<tr id="352434E1FF92004E25B94F46FA6BFC9D" box="[151,1436,845,867]" gridrow="6" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94F46FE42FC9D" box="[151,437,845,867]" gridcol="0" gridrow="6" pageId="7" pageNumber="44">ED</th>
<td id="76F55D9DFF92004E275C4F46FD4AFC9D" box="[626,701,845,867]" gridcol="1" gridrow="6" pageId="7" pageNumber="44">4.6</td>
<td id="76F55D9DFF92004E26FF4F46FA6BFC9D" box="[977,1436,845,867]" gridcol="2" gridrow="6" pageId="7" pageNumber="44">2.7–3.9 (3.3 ± 0.5)</td>
</tr>
<tr id="352434E1FF92004E25B94F7DFA6BFC72" box="[151,1436,886,908]" gridrow="7" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94F7DFE42FC72" box="[151,437,886,908]" gridcol="0" gridrow="7" pageId="7" pageNumber="44">END</th>
<td id="76F55D9DFF92004E275C4F7DFD4AFC72" box="[626,701,886,908]" gridcol="1" gridrow="7" pageId="7" pageNumber="44">4.7</td>
<td id="76F55D9DFF92004E26FF4F7DFA6BFC72" box="[977,1436,886,908]" gridcol="2" gridrow="7" pageId="7" pageNumber="44">2.5–3.9 (3.4 ± 0.5)</td>
</tr>
<tr id="352434E1FF92004E25B94FABFA6BFC48" box="[151,1436,928,950]" gridrow="8" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94FABFE42FC48" box="[151,437,928,950]" gridcol="0" gridrow="8" pageId="7" pageNumber="44">IND</th>
<td id="76F55D9DFF92004E275C4FABFD4AFC48" box="[626,701,928,950]" gridcol="1" gridrow="8" pageId="7" pageNumber="44">3.4</td>
<td id="76F55D9DFF92004E26FF4FABFA6BFC48" box="[977,1436,928,950]" gridcol="2" gridrow="8" pageId="7" pageNumber="44">2.1–3.2 (2.7 ± 0.4)</td>
</tr>
<tr id="352434E1FF92004E25B94FC2FA6BFC1E" box="[151,1436,969,992]" gridrow="9" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94FC2FE42FC1E" box="[151,437,969,992]" gridcol="0" gridrow="9" pageId="7" pageNumber="44">TD</th>
<td id="76F55D9DFF92004E275C4FC2FD4AFC1E" box="[626,701,969,992]" gridcol="1" gridrow="9" pageId="7" pageNumber="44">1.9</td>
<td id="76F55D9DFF92004E26FF4FC2FA6BFC1E" box="[977,1436,969,992]" gridcol="2" gridrow="9" pageId="7" pageNumber="44">1.0–1.5 (1.2 ± 0.2)</td>
</tr>
<tr id="352434E1FF92004E25B94FF9FA6BFBF7" box="[151,1436,1010,1033]" gridrow="10" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94FF9FE42FBF7" box="[151,437,1010,1033]" gridcol="0" gridrow="10" pageId="7" pageNumber="44">TL</th>
<td id="76F55D9DFF92004E275C4FF9FD4AFBF7" box="[626,701,1010,1033]" gridcol="1" gridrow="10" pageId="7" pageNumber="44">22.2</td>
<td id="76F55D9DFF92004E26FF4FF9FA6BFBF7" box="[977,1436,1010,1033]" gridcol="2" gridrow="10" pageId="7" pageNumber="44">15.0–17.7 (16.1 ± 1.0)</td>
</tr>
<tr id="352434E1FF92004E25B94810FA6BFBCC" box="[151,1436,1051,1074]" gridrow="11" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94810FE42FBCC" box="[151,437,1051,1074]" gridcol="0" gridrow="11" pageId="7" pageNumber="44">FL</th>
<td id="76F55D9DFF92004E275C4810FD4AFBCC" box="[626,701,1051,1074]" gridcol="1" gridrow="11" pageId="7" pageNumber="44">22</td>
<td id="76F55D9DFF92004E26FF4810FA6BFBCC" box="[977,1436,1051,1074]" gridcol="2" gridrow="11" pageId="7" pageNumber="44">13.4–18.3 (15.2 ± 1.8)</td>
</tr>
<tr id="352434E1FF92004E25B9484EFA6BFBA5" box="[151,1436,1093,1115]" gridrow="12" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B9484EFE42FBA5" box="[151,437,1093,1115]" gridcol="0" gridrow="12" pageId="7" pageNumber="44">Head width/head length</th>
<td id="76F55D9DFF92004E275C484EFD4AFBA5" box="[626,701,1093,1115]" gridcol="1" gridrow="12" pageId="7" pageNumber="44">114</td>
<td id="76F55D9DFF92004E26FF484EFA6BFBA5" box="[977,1436,1093,1115]" gridcol="2" gridrow="12" pageId="7" pageNumber="44">113.1–123.1</td>
</tr>
<tr id="352434E1FF92004E25B94865FA6BFB7A" box="[151,1436,1134,1156]" gridrow="13" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94865FE42FB7A" box="[151,437,1134,1156]" gridcol="0" gridrow="13" pageId="7" pageNumber="44">Head width/SVL</th>
<td id="76F55D9DFF92004E275C4865FD4AFB7A" box="[626,701,1134,1156]" gridcol="1" gridrow="13" pageId="7" pageNumber="44">41</td>
<td id="76F55D9DFF92004E26FF4865FA6BFB7A" box="[977,1436,1134,1156]" gridcol="2" gridrow="13" pageId="7" pageNumber="44">41.5–43.4</td>
</tr>
<tr id="352434E1FF92004E25B94893FA6BFB50" box="[151,1436,1176,1198]" gridrow="14" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94893FE42FB50" box="[151,437,1176,1198]" gridcol="0" gridrow="14" pageId="7" pageNumber="44">Eye-to-nostril distance/HW</th>
<td id="76F55D9DFF92004E275C4893FD4AFB50" box="[626,701,1176,1198]" gridcol="1" gridrow="14" pageId="7" pageNumber="44">30</td>
<td id="76F55D9DFF92004E26FF4893FA6BFB50" box="[977,1436,1176,1198]" gridcol="2" gridrow="14" pageId="7" pageNumber="44">25–32.7</td>
</tr>
<tr id="352434E1FF92004E25B948CAFA6BFB29" box="[151,1436,1217,1239]" gridrow="15" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B948CAFE42FB29" box="[151,437,1217,1239]" gridcol="0" gridrow="15" pageId="7" pageNumber="44">Interorbital distance /HW</th>
<td id="76F55D9DFF92004E275C48CAFD4AFB29" box="[626,701,1217,1239]" gridcol="1" gridrow="15" pageId="7" pageNumber="44">50</td>
<td id="76F55D9DFF92004E26FF48CAFA6BFB29" box="[977,1436,1217,1239]" gridcol="2" gridrow="15" pageId="7" pageNumber="44">47.3–53</td>
</tr>
<tr id="352434E1FF92004E25B948E1FA6BFAFE" box="[151,1436,1258,1280]" gridrow="16" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B948E1FE42FAFE" box="[151,437,1258,1280]" gridcol="0" gridrow="16" pageId="7" pageNumber="44">Internarial distance /IOD</th>
<td id="76F55D9DFF92004E275C48E1FD4AFAFE" box="[626,701,1258,1280]" gridcol="1" gridrow="16" pageId="7" pageNumber="44">44</td>
<td id="76F55D9DFF92004E26FF48E1FA6BFAFE" box="[977,1436,1258,1280]" gridcol="2" gridrow="16" pageId="7" pageNumber="44">39.6–53.8</td>
</tr>
<tr id="352434E1FF92004E25B94918FA6BFAD4" box="[151,1436,1299,1322]" gridrow="17" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94918FE42FAD4" box="[151,437,1299,1322]" gridcol="0" gridrow="17" pageId="7" pageNumber="44">Tympanum diameter /ED</th>
<td id="76F55D9DFF92004E275C4918FD4AFAD4" box="[626,701,1299,1322]" gridcol="1" gridrow="17" pageId="7" pageNumber="44">41</td>
<td id="76F55D9DFF92004E26FF4918FA6BFAD4" box="[977,1436,1299,1322]" gridcol="2" gridrow="17" pageId="7" pageNumber="44">33.3–38.7</td>
</tr>
<tr id="352434E1FF92004E25B94936FA6BFAAD" box="[151,1436,1341,1363]" gridrow="18" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B94936FE42FAAD" box="[151,437,1341,1363]" gridcol="0" gridrow="18" pageId="7" pageNumber="44">Tibia length/SVL</th>
<td id="76F55D9DFF92004E275C4936FD4AFAAD" box="[626,701,1341,1363]" gridcol="1" gridrow="18" pageId="7" pageNumber="44">59</td>
<td id="76F55D9DFF92004E26FF4936FA6BFAAD" box="[977,1436,1341,1363]" gridcol="2" gridrow="18" pageId="7" pageNumber="44">58.8–64.9</td>
</tr>
<tr id="352434E1FF92004E25B9496DFA6BFA82" box="[151,1436,1382,1404]" gridrow="19" pageId="7" pageNumber="44">
<th id="76F55D9DFF92004E25B9496DFE42FA82" box="[151,437,1382,1404]" gridcol="0" gridrow="19" pageId="7" pageNumber="44">Foot length/SVL</th>
<td id="76F55D9DFF92004E275C496DFD4AFA82" box="[626,701,1382,1404]" gridcol="1" gridrow="19" pageId="7" pageNumber="44">59</td>
<td id="76F55D9DFF92004E26FF496DFA6BFA82" box="[977,1436,1382,1404]" gridcol="2" gridrow="19" pageId="7" pageNumber="44">57.3–60.8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAB36A3FF92FFB625EB49C8FE01F902" blockId="7.[151,1437,1475,2004]" pageId="7" pageNumber="44">
<emphasis id="B960EAB1FF92FFB625EB49C8FDFFFA22" bold="true" box="[197,520,1475,1500]" pageId="7" pageNumber="44">
Coloration of the 
<typeStatus id="54AF8801FF92FFB624B449CFFDF4FA22" box="[410,515,1476,1500]" pageId="7" pageNumber="44" type="holotype">holotype</typeStatus>
.
</emphasis>
In life, dorsal color is light brown to green with dark markings; canthal and postorbital stripe black; interorbital blotch black; well-defined dark brown labial bars from eye to upper lip; posterior surfaces of thighs dark brown with black markings. Belly, anterior surfaces of thighs and concealed surfaces of tibia with black mottling on white background; throat brown with a white inverted triangle; flanks grayish brown, with black spots. Palms and soles grayish brown and iris reddish orange. In preservative, dorsal surfaces are grayish brown with darker spots, dark brown around dorsolateral folds; brown canthal stripe; dark labial bars; black supratympanic stripe; black cloacal patch; limbs brown with dark oblique bars; posterior surfaces of thighs grayish brown, groin, anterior surfaces of thighs white with brown blotches. Belly white with dark markings; throat brown with an inverted white triangle.
</paragraph>
</subSubSection>
<subSubSection id="C30E6528FF92FFB625EB4B0CFEF2F82A" pageId="7" pageNumber="44" type="distribution">
<paragraph id="8BAB36A3FF92FFB625EB4B0CFEF2F82A" blockId="7.[151,1437,1475,2004]" pageId="7" pageNumber="44">
<emphasis id="B960EAB1FF92FFB625EB4B0CFDA3F8DE" bold="true" box="[197,596,1799,1824]" pageId="7" pageNumber="44">Distribution and natural history.</emphasis>
The species is only known from the 
<typeStatus id="54AF8801FF92FFB621294B02FBC0F8DF" box="[1031,1079,1801,1825]" pageId="7" pageNumber="44">type</typeStatus>
locality (
<figureCitation id="132F2A26FF92FFB621884B0CFAFCF8E1" box="[1190,1291,1799,1823]" captionStart="FIGURE 6" captionStartId="8.[151,250,1975,1998]" captionTargetBox="[240,1343,207,1937]" captionTargetId="figure@8.[205,1374,169,1952]" captionTargetPageId="8" captionText="FIGURE 6. Map of Colombia showing distribution of species mentioned in the text, and with an square signaling the type locality of Pristimantis quicato sp. nov." httpUri="https://zenodo.org/record/279307/files/figure.png" pageId="7" pageNumber="44">Figure 6</figureCitation>
). Specimens were found between 1900 h and 2300 h in primary cloud forest in association with plants along streams, at heights above 
<quantity id="4CEC9B46FF92FFB625CF4B5BFED2F896" box="[225,293,1872,1896]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="44" unit="m" value="1.5">1.5 m</quantity>
. The new species was not found in zones of habitat intervention (pastures, edges of road), which suggests little or no tolerance to habitat disturbance. 
<taxonomicName id="4C144D20FF92FFB627FB4B7EFC4CF872" box="[725,955,1909,1932]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="quicato">
<emphasis id="B960EAB1FF92FFB627FB4B7EFC4CF872" box="[725,955,1909,1932]" italics="true" pageId="7" pageNumber="44">Pristimantis quicato</emphasis>
</taxonomicName>
was found in sympatry with 
<taxonomicName id="4C144D20FF92FFB6203D4B7EFEE7F84E" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B960EAB1FF92FFB6203D4B7EFEE7F84E" italics="true" pageId="7" pageNumber="44">Pristimantis boulengeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D20FF92FFB6240C4B92FE7EF84E" box="[290,393,1945,1968]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="piceus">
<emphasis id="B960EAB1FF92FFB6240C4B92FE7EF84E" box="[290,393,1945,1968]" italics="true" pageId="7" pageNumber="44">P. piceus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D20FF92FFB624B54B92FDA5F84E" box="[411,594,1944,1968]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="thectopternus">
<emphasis id="B960EAB1FF92FFB624B54B92FDA5F84E" box="[411,594,1944,1968]" italics="true" pageId="7" pageNumber="44">P. thectopternus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D20FF92FFB6274B4B92FCF5F84E" box="[613,770,1944,1968]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="B960EAB1FF92FFB6274B4B92FCF5F84E" box="[613,770,1944,1968]" italics="true" pageId="7" pageNumber="44">P. uranobates</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D20FF92FFB6263A4B93FBC6F84E" box="[788,1073,1944,1968]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="larinopygion">
<emphasis id="B960EAB1FF92FFB6263A4B93FBC6F84E" box="[788,1073,1944,1968]" italics="true" pageId="7" pageNumber="44">Hyloscirtus larinopygion</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D20FF92FFB6216D4B93FB12F84E" box="[1091,1253,1944,1968]" class="Amphibia" family="Bufonidae" genus="Osornophryne" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B960EAB1FF92FFB6216D4B93FB12F84E" box="[1091,1253,1944,1968]" italics="true" pageId="7" pageNumber="44">Osornophryne</emphasis>
</taxonomicName>
sp. and 
<taxonomicName id="4C144D20FF92FFB620604B93FF28F82A" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B960EAB1FF92FFB620604B93FF28F82A" italics="true" pageId="7" pageNumber="44">Bolitoglossa</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
<caption id="DF6B662BFF9DFFB925B94BBCFDDEF812" httpUri="https://zenodo.org/record/279307/files/figure.png" pageId="8" pageNumber="45" targetBox="[240,1343,207,1937]" targetPageId="8">
<paragraph id="8BAB36A3FF9DFFB925B94BBCFDDEF812" blockId="8.[151,1436,1975,2028]" pageId="8" pageNumber="45">
<emphasis id="B960EAB1FF9DFFB925B94BBCFEE3F833" bold="true" box="[151,276,1975,1998]" pageId="8" pageNumber="45">FIGURE 6.</emphasis>
Map of Colombia showing distribution of species mentioned in the text, and with an square signaling the type locality of 
<taxonomicName id="4C144D20FF9DFFB9242A4BDCFE25F812" box="[260,466,2007,2028]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="quicato" status="sp. nov.">
<emphasis id="B960EAB1FF9DFFB9242A4BDCFE25F812" box="[260,466,2007,2028]" italics="true" pageId="8" pageNumber="45">Pristimantis quicato</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357CAFF9DFFB924F74BDCFDDEF812" box="[473,553,2007,2028]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C30E6528FF9CFFB825E94C93FC4FFF2A" pageId="9" pageNumber="46" type="etymology">
<paragraph id="8BAB36A3FF9CFFB825E94C93FC4FFF2A" blockId="9.[151,1436,151,212]" pageId="9" pageNumber="46">
<emphasis id="B960EAB1FF9CFFB825E94C93FEB1FF4E" bold="true" box="[199,326,152,176]" pageId="9" pageNumber="46">Etymology</emphasis>
. The specific epithet is used as a noun in apposition. The name 
<taxonomicName id="4C144D20FF9CFFB8213B4C9CFB9FFF51" box="[1045,1128,151,175]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="quicato">quicato</taxonomicName>
is commonly used by local from the departments of Cauca and Valle del Cauca to refer to children.
</paragraph>
</subSubSection>
</treatment>
</document>