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<document id="430B4E7B0E5112B594F28A22440FB46F" ID-CLB-Dataset="29745" ID-DOI="10.11646/zootaxa.4442.4.5" ID-GBIF-Dataset="404b799f-8422-476d-af0d-74decce43f47" ID-ISSN="1175-5326" ID-Zenodo-Dep="1304888" ID-ZooBank="DBE99173-309F-4363-BBCE-F355476203F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530694481366" checkinUser="plazi" docAuthor="Chen, Zhi-Teng" docDate="2018" docId="03BD87A3E55FFF91FF5C8147FE515655" docLanguage="en" docName="zootaxa.4442.4.5.pdf" docOrigin="Zootaxa 4442 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Largusoperla difformitatem Chen 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="575" masterDocId="FF84FFDBE55EFF92FFCB837DFFE45027" masterDocTitle="A new fossil species of Largusoperla (Plecoptera: Perlidae) and its evolutionary implications" masterLastPageNumber="578" masterPageNumber="572" pageNumber="573" updateTime="1698552620967" updateUser="ExternalLinkService">
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<mods:title id="04305A07B46CB511BDC7B9A696C404CE">A new fossil species of Largusoperla (Plecoptera: Perlidae) and its evolutionary implications</mods:title>
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<mods:namePart id="70B6E31832C32806C39579499FF59828">Chen, Zhi-Teng</mods:namePart>
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<mods:date id="4F0E8A03BAC094C583CC7DD8E9104559">2018</mods:date>
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<treatment id="03BD87A3E55FFF91FF5C8147FE515655" ID-DOI="http://doi.org/10.5281/zenodo.5967862" ID-GBIF-Taxon="150804846" ID-Zenodo-Dep="5967862" LSID="urn:lsid:plazi:treatment:03BD87A3E55FFF91FF5C8147FE515655" httpUri="http://treatment.plazi.org/id/03BD87A3E55FFF91FF5C8147FE515655" lastPageId="3" lastPageNumber="575" pageId="1" pageNumber="573">
<subSubSection id="C30E653EE55FFF93FF5C8147FEE25251" pageId="1" pageNumber="573" type="nomenclature">
<paragraph id="8BAB36B5E55FFF93FF5C8147FD785273" blockId="1.[151,668,570,630]" box="[151,668,570,596]" pageId="1" pageNumber="573">
<heading id="D0E381D9E55FFF93FF5C8147FD785273" bold="true" box="[151,668,570,596]" fontSize="11" level="1" pageId="1" pageNumber="573" reason="1">
<emphasis id="B960EAA7E55FFF93FF5C8147FD785273" bold="true" box="[151,668,570,596]" pageId="1" pageNumber="573">
<taxonomicName id="4C144D36E55FFF93FF5C8147FDD55273" ID-CoL="8NGVM" authority="Chen" authorityName="Chen" authorityYear="2018" box="[151,561,570,596]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="difformitatem" status="sp. nov.">
<emphasis id="B960EAA7E55FFF93FF5C8147FE025273" bold="true" box="[151,486,570,596]" italics="true" pageId="1" pageNumber="573">Largusoperla difformitatem</emphasis>
Chen
</taxonomicName>
,
<taxonomicNameLabel id="A25357DCE55FFF93FDF58147FD785273" box="[574,668,570,596]" pageId="1" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB36B5E55FFF93FF5C8120FEE25251" blockId="1.[151,668,570,630]" box="[151,262,605,630]" pageId="1" pageNumber="573">
<figureCitation id="132F2A30E55FFF93FF5C8120FEE65251" box="[151,258,605,630]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 34" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 78" captionStartId-0="1.[151,250,2011,2033]" captionStartId-1="2.[151,250,1328,1350]" captionStartId-2="2.[151,250,2013,2035]" captionStartId-3="3.[151,250,1420,1442]" captionStartId-4="4.[151,250,1967,1989]" captionTargetBox-0="[326,1271,1253,1999]" captionTargetBox-1="[313,1231,733,1307]" captionTargetBox-4="[151,1417,1191,1956]" captionTargetId-0="figure@1.[326,1271,1253,1999]" captionTargetId-1="figure@2.[313,1273,733,1307]" captionTargetId-4="figure@4.[151,1436,1191,1956]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-4="4" captionText-0="FIGURE 12. Largusoperla difformitatem, sp. nov. Adult habituS, 1, dorSal view; 2,ventral view." captionText-1="FIGURE 34. Largusoperla difformitatem, sp. nov. 3, Head and pronotum, dorSal view; 4, ReconStruction of head and pronotum, dorSal view" captionText-2="FIGURE 5. Largusoperla difformitatem, sp. nov. Right forewing, dorSal view" captionText-3="FIGURE 6. Largusoperla difformitatem, sp. nov. Terminalia, dorSal view" captionText-4="FIGURE 78. Largusoperla difformitatem, sp. nov. 7, Terminalia, ventral view; 8, ReconStruction of terminalia, ventral view." httpUri-0="https://zenodo.org/record/1304890/files/figure.png" httpUri-1="https://zenodo.org/record/1304892/files/figure.png" httpUri-4="https://zenodo.org/record/1304894/files/figure.png" pageId="1" pageNumber="573">Figs. 18</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E653EE55FFF90FF5C81D8FABC5167" lastPageId="2" lastPageNumber="574" pageId="1" pageNumber="573" type="description">
<paragraph id="8BAB36B5E55FFF93FF5C81D8FDAC52C5" blockId="1.[151,1437,677,1206]" pageId="1" pageNumber="573">
<emphasis id="B960EAA7E55FFF93FF5C81D8FECD5299" bold="true" box="[151,297,677,702]" pageId="1" pageNumber="573">Description.</emphasis>
Macropterous; body length (eXcluding antennae and cerci) ca.
<quantity id="4CEC9B50E55FFF93FC2581D8FBA2529A" box="[1006,1094,677,702]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="573" unit="mm" value="9.5">9.5 mm</quantity>
, slender, generally yellow and patterned with dark brown (
<figureCitation id="132F2A30E55FFF93FE0581B4FDDF52C5" box="[462,571,713,738]" captionStart="FIGURE 12" captionStartId="1.[151,250,2011,2033]" captionTargetBox="[326,1271,1253,1999]" captionTargetId="figure@1.[326,1271,1253,1999]" captionTargetPageId="1" captionText="FIGURE 12. Largusoperla difformitatem, sp. nov. Adult habituS, 1, dorSal view; 2,ventral view." httpUri="https://zenodo.org/record/1304890/files/figure.png" pageId="1" pageNumber="573">Figs. 1, 2</figureCitation>
).
</paragraph>
<paragraph id="8BAB36B5E55FFF93FF0C8190FD2E5369" blockId="1.[151,1437,677,1206]" pageId="1" pageNumber="573">
<emphasis id="B960EAA7E55FFF93FF0C8190FEE35321" bold="true" box="[199,263,749,774]" pageId="1" pageNumber="573">Head</emphasis>
(
<figureCitation id="132F2A30E55FFF93FEDD8190FE675321" box="[278,387,749,774]" captionStart="FIGURE 34" captionStartId="2.[151,250,1328,1350]" captionTargetBox="[313,1231,733,1307]" captionTargetId="figure@2.[313,1273,733,1307]" captionTargetPageId="2" captionText="FIGURE 34. Largusoperla difformitatem, sp. nov. 3, Head and pronotum, dorSal view; 4, ReconStruction of head and pronotum, dorSal view" httpUri="https://zenodo.org/record/1304892/files/figure.png" pageId="1" pageNumber="573">Figs. 3, 4</figureCitation>
): Head rounded, with a dark stigma covering the three ocelli, compound eyes dark. Antennae brown and hairy, only about 10 basal segments preserved. MaXillary palps slender, with a shortened apical segment, about two times longer than labial palps.
</paragraph>
<paragraph id="8BAB36B5E55FFF93FF0C8024FB8B539D" blockId="1.[151,1437,677,1206]" pageId="1" pageNumber="573">
<emphasis id="B960EAA7E55FFF93FF0C8024FEA45355" bold="true" box="[199,320,857,882]" pageId="1" pageNumber="573">Pronotum</emphasis>
(
<figureCitation id="132F2A30E55FFF93FE998024FE205355" box="[338,452,857,882]" captionStart="FIGURE 34" captionStartId="2.[151,250,1328,1350]" captionTargetBox="[313,1231,733,1307]" captionTargetId="figure@2.[313,1273,733,1307]" captionTargetPageId="2" captionText="FIGURE 34. Largusoperla difformitatem, sp. nov. 3, Head and pronotum, dorSal view; 4, ReconStruction of head and pronotum, dorSal view" httpUri="https://zenodo.org/record/1304892/files/figure.png" pageId="1" pageNumber="573">Figs. 3, 4</figureCitation>
): Pronotum rugose, about as wide as head, mostly brown eXcept for the pale median band, near trapezoidal with obtuse anterior corners. MesothoraX and metathoraX sclerotized, nearly as wide as pronotum. Legs brown; first two tarsal segments shortest with conspicuous euplantulae.
</paragraph>
<paragraph id="8BAB36B5E55FFF93FF0C80B8FB535491" blockId="1.[151,1437,677,1206]" pageId="1" pageNumber="573">
<emphasis id="B960EAA7E55FFF93FF0C80B8FEF753F9" bold="true" box="[199,275,965,990]" pageId="1" pageNumber="573">Wings</emphasis>
(
<figureCitation id="132F2A30E55FFF93FEEA80B8FE8A53F9" box="[289,366,965,990]" captionStart="FIGURE 12" captionStartId="1.[151,250,2011,2033]" captionTargetBox="[326,1271,1253,1999]" captionTargetId="figure@1.[326,1271,1253,1999]" captionTargetPageId="1" captionText="FIGURE 12. Largusoperla difformitatem, sp. nov. Adult habituS, 1, dorSal view; 2,ventral view." httpUri="https://zenodo.org/record/1304890/files/figure.png" pageId="1" pageNumber="573">Figs. 1</figureCitation>
,
<figureCitation id="132F2A30E55FFF93FEB780B8FE6853F9" box="[380,396,965,990]" captionStart="FIGURE 5" captionStartId="2.[151,250,2013,2035]" captionText="FIGURE 5. Largusoperla difformitatem, sp. nov. Right forewing, dorSal view" pageId="1" pageNumber="573">5</figureCitation>
): Wings hyaline, veins brown, a subapical angle presents in all wings. Right forewing: length ca.
<quantity id="4CEC9B50E55FFF93FF088094FEC55426" box="[195,289,1001,1026]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="573" unit="mm" value="9.5">9.5 mm</quantity>
; ScP about three-fourth of forewing length, reaching ScP+RA beyond ra-rp; apical area between anterior margin and RA with one crossvein; over 13 crossveins present between ScP and the anterior margin; RP originated from middle of RA and with three branches; M forked at near apical one-third of the wing length; ra-rp not connected with the opposite crossvein rp-ma; CuA forked basal to the fork of M, with three branches; CuP simple; area between M and CuA with siX crossveins; area between CuA and CuP with four crossveins; AA1 simple, AA2 not visible. Hind wings length ca. 8.0 mm, apical veins similar to right forewing.
</paragraph>
<caption id="DF6B663DE55FFF93FF5C84A6FB9357D6" box="[151,1143,2011,2033]" httpUri="https://zenodo.org/record/1304890/files/figure.png" pageId="1" pageNumber="573" startId="1.[151,250,2011,2033]" targetBox="[326,1271,1253,1999]" targetPageId="1">
<paragraph id="8BAB36B5E55FFF93FF5C84A6FB9357D6" blockId="1.[151,1143,2011,2033]" box="[151,1143,2011,2033]" pageId="1" pageNumber="573">
<emphasis id="B960EAA7E55FFF93FF5C84A6FEC857D7" bold="true" box="[151,300,2011,2033]" pageId="1" pageNumber="573">FIGURE 12.</emphasis>
<taxonomicName id="4C144D36E55FFF93FEF984A6FDAF57D6" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[306,587,2011,2033]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="difformitatem" status="sp. nov.">
<emphasis id="B960EAA7E55FFF93FEF984A6FDAF57D6" box="[306,587,2011,2033]" italics="true" pageId="1" pageNumber="573">Largusoperla difformitatem</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25357DCE55FFF93FD9384A6FD4C57D7" box="[600,680,2011,2032]" pageId="1" pageNumber="573" rank="species">
<emphasis id="B960EAA7E55FFF93FD9384A6FD4C57D7" bold="true" box="[600,680,2011,2032]" pageId="1" pageNumber="573">sp. nov.</emphasis>
</taxonomicNameLabel>
Adult habituS, 1, dorSal view; 2,ventral view.
</paragraph>
</caption>
<paragraph id="8BAB36B5E55CFF90FF0C83EAFABC5167" blockId="2.[151,1436,151,680]" pageId="2" pageNumber="574">
<emphasis id="B960EAA7E55CFF90FF0C83EAFED85097" bold="true" box="[199,316,151,176]" pageId="2" pageNumber="574">Abdomen</emphasis>
(
<figureCitation id="132F2A30E55CFF90FE8783EAFE505097" box="[332,436,151,176]" captionStart="FIGURE 12" captionStartId="1.[151,250,2011,2033]" captionTargetBox="[326,1271,1253,1999]" captionTargetId="figure@1.[326,1271,1253,1999]" captionTargetPageId="1" captionText="FIGURE 12. Largusoperla difformitatem, sp. nov. Adult habituS, 1, dorSal view; 2,ventral view." httpUri="https://zenodo.org/record/1304890/files/figure.png" pageId="2" pageNumber="574">Figs. 1-2</figureCitation>
,
<figureCitation id="132F2A30E55CFF90FE0A83EAFE0E5097" box="[449,490,151,176]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 78" captionStartId-0="3.[151,250,1420,1442]" captionStartId-1="4.[151,250,1967,1989]" captionTargetBox-1="[151,1417,1191,1956]" captionTargetId-1="figure@4.[151,1436,1191,1956]" captionTargetPageId-1="4" captionText-0="FIGURE 6. Largusoperla difformitatem, sp. nov. Terminalia, dorSal view" captionText-1="FIGURE 78. Largusoperla difformitatem, sp. nov. 7, Terminalia, ventral view; 8, ReconStruction of terminalia, ventral view." httpUri-1="https://zenodo.org/record/1304894/files/figure.png" pageId="2" pageNumber="574">6-8</figureCitation>
): Abdomen near half the length of the body, with dark lateral marks. Posterolateral margins of tergum 9 eXtended. Sternum 9 eXtended backwards, posteromedially with a transverse lobe. Tergum 10 not well-preserved but apparently cleft, without projecting hemiterga. Paraproct broad and weakly sclerotized, length two times longer than width; left paraproct subtriangular with sharp apeX, widened at mid-length; right paraproct subtriangular with sharp apeX, constricted at mid-length. Cerci hairy and short, segments not clear.
</paragraph>
</subSubSection>
<subSubSection id="C30E653EE55CFF90FF0C8231FD0A518A" pageId="2" pageNumber="574" type="materials_examined">
<paragraph id="8BAB36B5E55CFF90FF0C8231FD0A518A" blockId="2.[151,1436,151,680]" pageId="2" pageNumber="574">
<emphasis id="B960EAA7E55CFF90FF0C8231FE935142" bold="true" box="[199,375,332,357]" pageId="2" pageNumber="574">Type material.</emphasis>
Holotype male, No. CZT-PLE-MA2. This specimen is mostly wizened; apical halves of left foreleg and left midleg and apeX of antennae are not preserved; dorsal aspect of abdomen is not clear; wings are overlapped so only the right forewing is clear visible.
</paragraph>
</subSubSection>
<subSubSection id="C30E653EE55CFF90FF0C82CAFDFD51D2" pageId="2" pageNumber="574" type="etymology">
<paragraph id="8BAB36B5E55CFF90FF0C82CAFA9851E8" blockId="2.[151,1436,151,680]" box="[199,1404,439,464]" pageId="2" pageNumber="574">
<emphasis id="B960EAA7E55CFF90FF0C82CAFEAA51F7" bold="true" box="[199,334,439,464]" pageId="2" pageNumber="574">Etymology.</emphasis>
The specific epithet means “asymmetry” in Latin, which refers to the asymmetrical paraprocts.
</paragraph>
<paragraph id="8BAB36B5E55CFF90FF0C82A1FDFD51D2" blockId="2.[151,1436,151,680]" box="[199,537,476,501]" pageId="2" pageNumber="574">
<emphasis id="B960EAA7E55CFF90FF0C82A1FE9551D2" bold="true" box="[199,369,476,501]" pageId="2" pageNumber="574">Syninclusions.</emphasis>
Plant remains.
</paragraph>
</subSubSection>
<subSubSection id="C30E653EE55CFF91FF0C8282FE515655" lastPageId="3" lastPageNumber="575" pageId="2" pageNumber="574" type="discussion">
<paragraph id="8BAB36B5E55CFF90FF0C8282FED25280" blockId="2.[151,1436,151,680]" pageId="2" pageNumber="574">
<emphasis id="B960EAA7E55CFF90FF0C8282FEDF523F" bold="true" box="[199,315,511,536]" pageId="2" pageNumber="574">Remarks.</emphasis>
Similar body color, wing venation and terminalia structures of the new species indicates its taXonomic inclusion in
<taxonomicName id="4C144D36E55CFF90FE6B8159FDD3521B" authorityName="Chen, Wang &amp; Du" authorityYear="2018" box="[416,567,548,572]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAA7E55CFF90FE6B8159FDD3521B" box="[416,567,548,572]" italics="true" pageId="2" pageNumber="574">Largusoperla</emphasis>
</taxonomicName>
. The new species can be distinguished from other three known congeners by shape of the asymmetrical paraprocts and the subapical angles of wings. The paraproct shape of
<taxonomicName id="4C144D36E55CFF90FB078134FA665247" authorityName="Chen" authorityYear="2018" box="[1228,1410,584,608]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="difformitatem">
<emphasis id="B960EAA7E55CFF90FB078134FA665247" box="[1228,1410,584,608]" italics="true" pageId="2" pageNumber="574">L. difformitatem</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C144D36E55CFF90FE8F8110FE7152A3" authorityName="Chen, Wang &amp; Du" authorityYear="2018" box="[324,405,620,644]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="flata">
<emphasis id="B960EAA7E55CFF90FE8F8110FE7152A3" box="[324,405,620,644]" italics="true" pageId="2" pageNumber="574">L. flata</emphasis>
</taxonomicName>
, however, the paraprocts of
<taxonomicName id="4C144D36E55CFF90FD1B8110FCC452A3" authorityName="Chen, Wang &amp; Du" authorityYear="2018" box="[720,800,620,644]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="flata">
<emphasis id="B960EAA7E55CFF90FD1B8110FCC452A3" box="[720,800,620,644]" italics="true" pageId="2" pageNumber="574">L. flata</emphasis>
</taxonomicName>
are laterally thickened, less divergent and shorter than
<taxonomicName id="4C144D36E55CFF90FA4E8110FECB528F" authorityName="Chen" authorityYear="2018" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="difformitatem">
<emphasis id="B960EAA7E55CFF90FA4E8110FECB528F" italics="true" pageId="2" pageNumber="574">L. difformitatem</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF6B663DE55CFF90FF5C864DFE9F5542" httpUri="https://zenodo.org/record/1304892/files/figure.png" pageId="2" pageNumber="574" startId="2.[151,250,1328,1350]" targetBox="[313,1231,733,1307]" targetPageId="2">
<paragraph id="8BAB36B5E55CFF90FF5C864DFE9F5542" blockId="2.[151,1435,1328,1381]" pageId="2" pageNumber="574">
<emphasis id="B960EAA7E55CFF90FF5C864DFED55562" bold="true" box="[151,305,1328,1350]" pageId="2" pageNumber="574">FIGURE 34.</emphasis>
<taxonomicName id="4C144D36E55CFF90FEF7864DFDBE5561" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[316,602,1328,1350]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="difformitatem" status="sp. nov.">
<emphasis id="B960EAA7E55CFF90FEF7864DFDBE5561" box="[316,602,1328,1350]" italics="true" pageId="2" pageNumber="574">Largusoperla difformitatem</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25357DCE55CFF90FDA0864DFD245562" box="[619,704,1328,1349]" pageId="2" pageNumber="574" rank="species">
<emphasis id="B960EAA7E55CFF90FDA0864DFD245562" bold="true" box="[619,704,1328,1349]" pageId="2" pageNumber="574">sp. nov.</emphasis>
</taxonomicNameLabel>
3, Head and pronotum, dorSal view; 4, ReconStruction of head and pronotum, dorSal view
</paragraph>
</caption>
<caption id="DF6B663DE55CFF90FF5C84A0FC5657D5" box="[151,946,2012,2035]" pageId="2" pageNumber="574">
<paragraph id="8BAB36B5E55CFF90FF5C84A0FC5657D5" blockId="2.[151,946,2012,2035]" box="[151,946,2012,2035]" pageId="2" pageNumber="574">
<emphasis id="B960EAA7E55CFF90FF5C84A0FEF657D5" bold="true" box="[151,274,2013,2035]" pageId="2" pageNumber="574">FIGURE 5.</emphasis>
<taxonomicName id="4C144D36E55CFF90FED284A1FDD657D5" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[281,562,2012,2034]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="difformitatem" status="sp. nov.">
<emphasis id="B960EAA7E55CFF90FED284A1FDD657D5" box="[281,562,2012,2034]" italics="true" pageId="2" pageNumber="574">Largusoperla difformitatem</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25357DCE55CFF90FDF584A0FD6B57D5" box="[574,655,2013,2034]" pageId="2" pageNumber="574" rank="species">
<emphasis id="B960EAA7E55CFF90FDF584A0FD6B57D5" bold="true" box="[574,655,2013,2034]" pageId="2" pageNumber="574">sp. nov.</emphasis>
</taxonomicNameLabel>
Right forewing, dorSal view
</paragraph>
</caption>
<caption id="DF6B663DE55DFF91FF5C86F1FC615585" box="[151,901,1420,1442]" pageId="3" pageNumber="575">
<paragraph id="8BAB36B5E55DFF91FF5C86F1FC615585" blockId="3.[151,901,1420,1442]" box="[151,901,1420,1442]" pageId="3" pageNumber="575">
<emphasis id="B960EAA7E55DFF91FF5C86F1FEF65586" bold="true" box="[151,274,1420,1442]" pageId="3" pageNumber="575">FIGURE 6.</emphasis>
<taxonomicName id="4C144D36E55DFF91FED286F1FDD65585" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[281,562,1420,1442]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="difformitatem" status="sp. nov.">
<emphasis id="B960EAA7E55DFF91FED286F1FDD65585" box="[281,562,1420,1442]" italics="true" pageId="3" pageNumber="575">Largusoperla difformitatem</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25357DCE55DFF91FDF586F1FD6B5586" box="[574,655,1420,1441]" pageId="3" pageNumber="575" rank="species">
<emphasis id="B960EAA7E55DFF91FDF586F1FD6B5586" bold="true" box="[574,655,1420,1441]" pageId="3" pageNumber="575">sp. nov.</emphasis>
</taxonomicNameLabel>
Terminalia, dorSal view
</paragraph>
</caption>
<paragraph id="8BAB36B5E55DFF91FF0C86B4FE515655" blockId="3.[151,1437,1481,1650]" pageId="3" pageNumber="575">
The asymmetrical paraprocts of
<taxonomicName id="4C144D36E55DFF91FDF886B7FD0D55C6" authorityName="Chen" authorityYear="2018" box="[563,745,1481,1505]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="difformitatem">
<emphasis id="B960EAA7E55DFF91FDF886B7FD0D55C6" box="[563,745,1481,1505]" italics="true" pageId="3" pageNumber="575">L. difformitatem</emphasis>
</taxonomicName>
were not formed by the fossilization process. The outlines of the paraprocts are clearly visible and the differences between the two paraprocts are distinctive. In addition, asymmetrical paraprocts also occurs in other species of
<taxonomicName id="4C144D36E55DFF91FCEB856CFC53560E" authorityName="Chen, Wang &amp; Du" authorityYear="2018" box="[800,951,1553,1577]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAA7E55DFF91FCEB856CFC53560E" box="[800,951,1553,1577]" italics="true" pageId="3" pageNumber="575">Largusoperla</emphasis>
</taxonomicName>
being described by colleagues elsewhere. The asymmetry of genitalia is a possible plesiomorphic character of the genus
<taxonomicName id="4C144D36E55DFF91FBD58548FB51566A" authorityName="Chen, Wang &amp; Du" authorityYear="2018" box="[1054,1205,1589,1613]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAA7E55DFF91FBD58548FB51566A" box="[1054,1205,1589,1613]" italics="true" pageId="3" pageNumber="575">Largusoperla</emphasis>
</taxonomicName>
and perhaps for the
<taxonomicName id="4C144D36E55DFF91FF5C8524FED35655" box="[151,311,1625,1650]" class="Insecta" family="Perlidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="subFamily" subFamily="Acroneuriinae">Acroneuriinae</taxonomicName>
in general.
</paragraph>
</subSubSection>
</treatment>
</document>