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<document id="7032DE5A8541BC45BD045AEDEFD5B664" ID-DOI="10.11646/zootaxa.4040.1.5" ID-GBIF-Dataset="c486d3d3-9944-4850-af11-c1fbc61bdaac" ID-ISSN="1175-5326" ID-Zenodo-Dep="289985" ID-ZooBank="703E3F2B-69F7-4781-ACDE-51C6291544BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333711373" checkinUser="plazi" docAuthor="Henao, Efrain R., Carneiro, Eduardo, Mielke, Olaf H. H. &amp; Casagrade, Mirna M." docDate="2015" docId="4A5E87B9FFB29A16FF19B6E16B40B4F6" docLanguage="en" docName="zt04040p073.pdf" docOrigin="Zootaxa 4040 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wahydra nieblensis Steinhauser 1991" docType="treatment" docVersion="6" lastPageNumber="62" masterDocId="B667FFC1FFB19A15FF8EB75B680AB72F" masterDocTitle="Taxonomic notes on Wahydra Steinhauser (Hesperiidae, Hesperiinae, Anthoptini) with description of four new species" masterLastPageNumber="73" masterPageNumber="59" pageNumber="62" updateTime="1698618892071" updateUser="plazi">
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<mods:title id="63C45EFCF37232CF7C7BC25CEE7B9E64">Taxonomic notes on Wahydra Steinhauser (Hesperiidae, Hesperiinae, Anthoptini) with description of four new species</mods:title>
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<mods:namePart id="2CD7441623221349585A0F36E950ABFE">Henao, Efrain R.</mods:namePart>
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<mods:namePart id="4B876F92D28ED67CB910B14CEE274DB4">Carneiro, Eduardo</mods:namePart>
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<mods:namePart id="09CA958D4F56B8CB48DEE150A9C436E8">Mielke, Olaf H. H.</mods:namePart>
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<mods:namePart id="A1A9864EBE3FBE20D0C748409E719165">Casagrade, Mirna M.</mods:namePart>
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<mods:date id="EA6DB71A58775EAA5FF6BE972DC17755">2015</mods:date>
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<treatment id="4A5E87B9FFB29A16FF19B6E16B40B4F6" ID-DOI="http://doi.org/10.5281/zenodo.6110607" ID-GBIF-Taxon="127694073" ID-Zenodo-Dep="6110607" LSID="urn:lsid:plazi:treatment:4A5E87B9FFB29A16FF19B6E16B40B4F6" httpUri="http://treatment.plazi.org/id/4A5E87B9FFB29A16FF19B6E16B40B4F6" lastPageNumber="62" pageId="3" pageNumber="62">
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<paragraph id="C24836AFFFB29A16FF19B6E16A6EB6FA" blockId="3.[151,612,442,502]" box="[151,612,442,469]" pageId="3" pageNumber="62">
<heading id="990081C3FFB29A16FF19B6E16A6EB6FA" bold="true" box="[151,612,442,469]" fontSize="11" level="1" pageId="3" pageNumber="62" reason="1">
<taxonomicName id="05F74D2CFFB29A16FF19B6E16A6EB6FA" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[151,612,442,469]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="nieblensis">
<emphasis id="F083EABDFFB29A16FF19B6E16A6EB6FA" bold="true" box="[151,612,442,469]" pageId="3" pageNumber="62">
<emphasis id="F083EABDFFB29A16FF19B6E1698EB6FB" bold="true" box="[151,388,442,468]" italics="true" pageId="3" pageNumber="62">Wahydra nieblensis</emphasis>
<bibRefCitation id="A6664B5EFFB29A16FE04B6E06A6EB6FA" author="Steinhauser" box="[394,612,443,469]" pageId="3" pageNumber="62" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article" year="1991">Steinhauser, 1991</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C24836AFFFB29A16FF19B6866955B6D9" blockId="3.[151,612,442,502]" box="[151,351,477,502]" pageId="3" pageNumber="62">
(
<figureCitation id="5ACC2A2AFFB29A16FF11B6866909B6D9" box="[159,259,477,502]" captionStart="FIGURES 1 26" captionStartId="4.[151,264,1812,1834]" captionTargetBox="[188,1391,195,1811]" captionTargetId="figure@4.[188,1398,195,1907]" captionTargetPageId="4" captionText="FIGURES 1 26. Dorsal and ventral views of Wahydra Steinhauser, 1991 species. Figs 1 2: Wahydra kenava (Butler, 1870); Figs 3 4: Wahydra bella (Hayward, 1939); Figs 5 6: Wahydra nieblensis Steinhauser, 1991; Figs 7 8: Wahydra ekka (Evans, 1955) comb. nov.; Figs 9 10: Wahydra subhebetis Steinhauser, 1991; Figs 11 12: Wahydra tassa (Evans, 1955); Figs 13 16: Wahydra vola (Evans, 1955) (male left, female right); Figs 17 18: Wahydra trinitas Henao, Carneiro, Mielke &amp; Casagrande sp. nov. (Holotype).; Figs 19 20: Wahydra curtis Mielke, Henao, Carneiro &amp; Casagrande sp. nov. (Paratypus); Figs 21 22: Wahydra shueyi Carneiro, Henao, Mielke &amp; Casagrande sp. nov. (Paratypus).; Figs 23 24: Wahydra mapiriensis Casagrande, Henao, Carneiro &amp; Mielke sp. nov. (Holotype).; Figs 25 26: Wahydra sp. (unidentified female)." httpUri="https://zenodo.org/record/289986/files/figure.png" pageId="3" pageNumber="62">Figs 5, 6</figureCitation>
,
<figureCitation id="5ACC2A2AFFB29A16FE81B6856927B6D9" box="[271,301,478,502]" captionStart="FIGURES 27 40" captionStartId="5.[151,264,1801,1823]" captionTargetBox="[197,1344,199,1760]" captionTargetId="figure@5.[191,1359,193,1770]" captionTargetPageId="5" captionText="FIGURES 27 40. Right dorsal forewing of the male of Wahydra Steinhauser, 1991 species, showing general aspect of brands (27 39). Fig. 27: Wahydra kenava (Butler, 1870); Fig. 28: Wahydra bella (Hayward, 1939); Fig. 29: Wahydra nieblensis Steinhauser, 1991; Fig. 30: Wahydra ekka (Evans, 1955) comb. nov.; Fig. 31: Wahydra subhebetis Steinhauser, 1991; Fig. 32: Wahydra tassa (Evans, 1955); Fig. 33: Wahydra vola (Evans, 1955); Fig. 34: Wahydra trinitas Carneiro, Mielke &amp; Casagrande sp. nov. (Holotype); Fig. 35: Wahydra curtis Mielke, Henao, Carneiro &amp; Casagrande sp. nov. (Paratype); Fig. 36: Wahydra shueyi Carneiro, Henao, Mielke &amp; Casagrande sp. nov. (Holotype).; Fig. 37: Wahydra mapiriensis Casagrande, Henao, Carneiro &amp; Mielke sp. nov. (Holotype); Fig. 38: Wahydra dores (Bell, 1959); Fig. 39: Wahydra thisbe (Hayward, 1942); Fig. 40: Wahydra sp. (unidentified female) dorsal discal spots." httpUri="https://zenodo.org/record/289987/files/figure.png" pageId="3" pageNumber="62">29</figureCitation>
,
<figureCitation id="5ACC2A2AFFB29A16FEB7B685695CB6D9" box="[313,342,478,502]" captionStart="FIGURES 41 46" captionStartId="6.[151,264,1696,1718]" captionTargetBox="[163,1390,204,1633]" captionTargetId="figure@6.[151,1436,193,1675]" captionTargetPageId="6" captionText="FIGURES 41 46. Male genitalia of Wahydra Steinhauser, 1991 species. Fig. 41: Wahydra kenava (Butler, 1870); Fig. 42: Wahydra bella (Hayward, 1939); Fig. 43: Wahydra nieblensis Steinhauser, 1991; Fig. 44: Wahydra ekka (Evans, 1955) comb. nov.; Fig. 45: Wahydra subhebetis Steinhauser, 1991; Fig. 46: Wahydra tassa (Evans, 1955)." httpUri="https://zenodo.org/record/289988/files/figure.png" pageId="3" pageNumber="62">43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB29A16FF19B57E6C83B599" pageId="3" pageNumber="62" type="reference_group">
<paragraph id="C24836AFFFB29A16FF19B57E6C83B599" blockId="3.[151,1436,549,694]" pageId="3" pageNumber="62">
<taxonomicName id="05F74D2CFFB29A16FF19B57E6A17B514" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[151,541,549,571]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="nieblensis">
<emphasis id="F083EABDFFB29A16FF19B57E6957B514" box="[151,349,549,571]" italics="true" pageId="3" pageNumber="62">Wahydra nieblensis</emphasis>
<bibRefCitation id="A6664B5EFFB29A16FEEBB57E6A17B514" author="Steinhauser" box="[357,541,549,571]" pageId="3" pageNumber="62" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article" year="1991">Steinhauser, 1991</bibRefCitation>
</taxonomicName>
.
<emphasis id="F083EABDFFB29A16FDA9B57E6B0DB514" bold="true" box="[551,775,549,571]" pageId="3" pageNumber="62">Bull. Allyn Mus. 132</emphasis>
: 35, figs 4245 (male, female d, v), 89 (male gen.), 110 (female gen.);
<typeStatus id="1D4C880DFFB29A16FE89B51F696AB575" box="[263,352,580,602]" pageId="3" pageNumber="62" type="holotype">holotype</typeStatus>
male,
<date id="B649106FFFB29A16FE2AB51F6A07B575" box="[420,525,580,602]" pageId="3" pageNumber="62" value="1971-08">VIII-1971</date>
, Niebli, NW slope Vol. Pichincha, Pichincha,
<collectingCountry id="BAE0763FFFB29A16FC6EB51F6C3CB575" box="[992,1078,580,602]" name="Ecuador" pageId="3" pageNumber="62">Ecuador</collectingCountry>
,
<quantity id="050F9B4AFFB29A16FBCEB51F6C85B575" box="[1088,1167,580,602]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="62" unit="m" value="3500.0">3500 m</quantity>
, R. de Lafebre leg.; AME, transferred to MGCL.—
<bibRefCitation id="A6664B5EFFB29A16FE31B5386A45B556" author="Bridges" box="[447,591,611,633]" pageId="3" pageNumber="62" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Hesperioidea (Lepidoptera) of the world. Urbana, 598 pp." type="book" year="1994">Bridges, 1994</bibRefCitation>
.
<emphasis id="F083EABDFFB29A16FDD6B5386D43B556" bold="true" box="[600,1353,611,633]" pageId="3" pageNumber="62">Cat. Fam.-Group, Gen.-Group, Sp. Group Nam. Hesp. (Lep.) World 8</emphasis>
, p. 154;
<emphasis id="F083EABDFFB29A16FF49B5DA68DEB5B8" bold="true" box="[199,212,641,663]" pageId="3" pageNumber="62">9</emphasis>
, p. 754.—
<bibRefCitation id="A6664B5EFFB29A16FEC8B5DA69DFB5B8" author="Mielke" box="[326,469,641,663]" pageId="3" pageNumber="62" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, pp. 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
. Hesperioidea, p. 84,
<emphasis id="F083EABDFFB29A16FD31B5DA6ADEB5B9" bold="true" box="[703,724,641,662]" italics="true" pageId="3" pageNumber="62">in</emphasis>
Lamas (ed.).
<emphasis id="F083EABDFFB29A16FCE5B5DA6D7CB5B8" bold="true" box="[875,1398,641,663]" pageId="3" pageNumber="62">Checklist: Part 4A, Hesperioidea-Papilionoidea</emphasis>
,
<emphasis id="F083EABDFFB29A16FA09B5DA6D96B5B9" bold="true" box="[1415,1436,641,662]" italics="true" pageId="3" pageNumber="62">in</emphasis>
Heppner (ed.).
<emphasis id="F083EABDFFB29A16FEEEB5FB6A53B599" bold="true" box="[352,601,672,694]" pageId="3" pageNumber="62">
<taxonomicName id="05F74D2CFFB29A16FEEEB5FB6A3EB599" authority="Neotrop. Lep." authorityName="Neotrop. Lep." box="[352,564,672,694]" class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
5A
</emphasis>
.—
<bibRefCitation id="A6664B5EFFB29A16FDF9B5FB6B09B599" author="Mielke" box="[631,771,672,694]" pageId="3" pageNumber="62" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 5: Hesperiinae 2. In: Megaleas - Zenis. Sociedade Brasileira de Zoologia, Curitiba, pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
.
<emphasis id="F083EABDFFB29A16FC82B5FB6C20B599" bold="true" box="[780,1066,672,694]" pageId="3" pageNumber="62">Cat. Amer. Hesperioidea 5</emphasis>
, p. 1363.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB29A16FF19B5B96C8BB442" pageId="3" pageNumber="62" type="diagnosis">
<paragraph id="C24836AFFFB29A16FF19B5B96C8BB442" blockId="3.[151,1436,738,985]" pageId="3" pageNumber="62">
<emphasis id="F083EABDFFB29A16FF19B5B9696BB5D4" bold="true" box="[151,353,738,763]" pageId="3" pageNumber="62">Diagnosis. Male:</emphasis>
Forewing length
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. Nudum 12. DFW brand large, subapical spot yellowish, in R5-M1. VFW costal area, apical spot and marginal band homogeneous yellowish. VHW with faint yellowish discal band and streak in CuA2-2A. Median apophyses of tegumen developed, large, simple, with a proximal constriction; lateral processes pointed. Harpe thin, curved. Ventro-posterior end of aedeagus truncated.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB29A16FF49B4236B40B4F6" pageId="3" pageNumber="62" type="materials_examined">
<paragraph id="C24836AFFFB29A16FF49B4236A3EB4BE" blockId="3.[151,1436,738,985]" box="[199,564,888,913]" pageId="3" pageNumber="62">
<emphasis id="F083EABDFFB29A16FF49B4236900B4BE" bold="true" box="[199,266,888,913]" pageId="3" pageNumber="62">
<typeStatus id="1D4C880DFFB29A16FF49B423690CB4BE" box="[199,262,888,913]" pageId="3" pageNumber="62">Type</typeStatus>
:
</emphasis>
<typeStatus id="1D4C880DFFB29A16FE9CB4236970B4BE" box="[274,378,888,913]" pageId="3" pageNumber="62" type="holotype">Holotype</typeStatus>
male in
<collectionCode id="A4E6AE6AFFB29A16FE55B4226A25B4BE" box="[475,559,889,913]" pageId="3" pageNumber="62">MGCL</collectionCode>
.
</paragraph>
<paragraph id="C24836AFFFB29A16FF49B4C76B40B4F6" blockId="3.[151,1436,738,985]" pageId="3" pageNumber="62">
<emphasis id="F083EABDFFB29A16FF49B4C76A11B49A" bold="true" box="[199,539,924,949]" pageId="3" pageNumber="62">
Studied material:
<collectingCountry id="BAE0763FFFB29A16FE2EB4C76A1CB49A" box="[416,534,924,949]" name="Colombia" pageId="3" pageNumber="62">Colombia</collectingCountry>
,
</emphasis>
Antioquia, Cerro del Padre Amaya,
<quantity id="050F9B4AFFB29A16FC33B4C76C04B49A" box="[957,1038,924,949]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="62" unit="m" value="2500.0">2500m</quantity>
,
<date id="B649106FFFB29A16FB94B4C66CA0B49A" box="[1050,1194,925,949]" pageId="3" pageNumber="62" value="1989-12-18">18-XII-1989</date>
,
<specimenCount id="D4F1FD26FFB29A16FB38B4C66D0CB49A" box="[1206,1286,924,949]" pageId="3" pageNumber="62" type="male">1 male</specimenCount>
, OM 50.981; Antioquia, Envigado,
<quantity id="050F9B4AFFB29A16FE00B49A69D5B4F6" box="[398,479,961,985]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="62" unit="m" value="2200.0">2200m</quantity>
,
<date id="B649106FFFB29A16FE64B49A6A56B4F6" box="[490,604,961,985]" pageId="3" pageNumber="62" value="1989-01-18">18-I-1989</date>
,
<specimenCount id="D4F1FD26FFB29A16FDE6B49A6ABCB4F6" box="[616,694,960,985]" pageId="3" pageNumber="62" type="male">1 male</specimenCount>
, OM 51.142.
</paragraph>
</subSubSection>
</treatment>
</document>