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<document id="E829E59D0FEFF5C4A97CD1C97C9D4D5F" ID-CLB-Dataset="31573" ID-DOI="10.11646/zootaxa.4347.1.13" ID-GBIF-Dataset="165e3b68-777e-46c9-a3b1-2b3fe2d3f102" ID-ISSN="1175-5326" ID-Zenodo-Dep="1044820" ID-ZooBank="A758BE2A-7541-422D-83C3-C66D30497CCE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510293117519" checkinUser="plazi" docAuthor="Nekhaev, Ivan O." docDate="2017" docId="910E878B0172FFE4E2C9FF58E2E9A02E" docLanguage="en" docName="zootaxa.4347.1.13.pdf" docOrigin="Zootaxa 4347 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Menestho schikovi Nekhaev 2017, sp. n." docType="treatment" docVersion="7" masterDocId="6D37FFF30171FFE0E25EFFCFE37DA139" masterDocTitle="A new species of Menestho Møller, 1842 from the Arctic with remarks on Menestho albula (Fabricius, 1780) (Gastropoda: Heterobranchia: Pyramidellidae)" updateTime="1698481053428" updateUser="ExternalLinkService">
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<mods:title id="D7954A9D618C3FC0C9086AEC3BB0DBBF">A new species of Menestho Møller, 1842 from the Arctic with remarks on Menestho albula (Fabricius, 1780) (Gastropoda: Heterobranchia: Pyramidellidae)</mods:title>
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<mods:name id="4ACE036EDDBA205A874043BF3D66C351" type="personal">
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<mods:namePart id="A53E3CDE5367F1239DA1FC51BD9E4A71">Nekhaev, Ivan O.</mods:namePart>
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<mods:title id="72C5605A63BA2F4D340D7E7C6932B55F">Zootaxa</mods:title>
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<mods:date id="F6877E62040DE1268DF9EBF6D09FBE8A">2017</mods:date>
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<mods:number id="3085BDA16558A17CFABAB3227B5372EE">2017-11-10</mods:number>
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<treatment id="910E878B0172FFE4E2C9FF58E2E9A02E" ID-DOI="http://doi.org/10.5281/zenodo.6021783" ID-GBIF-Taxon="137255915" ID-Zenodo-Dep="6021783" LSID="urn:lsid:plazi:treatment:910E878B0172FFE4E2C9FF58E2E9A02E" httpUri="http://treatment.plazi.org/id/910E878B0172FFE4E2C9FF58E2E9A02E" lastPageId="4" pageId="3">
<subSubSection id="51BD65160172FFE3E2C9FF58E14BA18B" box="[151,566,151,178]" pageId="3" type="nomenclature">
<paragraph id="1918369D0172FFE3E2C9FF58E14BA18B" blockId="3.[151,1436,151,2007]" box="[151,566,151,178]" pageId="3">
<heading id="425081F10172FFE3E2C9FF58E14BA18B" bold="true" box="[151,566,151,178]" fontSize="11" level="1" pageId="3" reason="1">
<emphasis id="2BD3EA8F0172FFE3E2C9FF58E14BA18B" bold="true" box="[151,566,151,178]" pageId="3">
<taxonomicName id="DEA74D1E0172FFE3E2C9FF58E290A18B" ID-CoL="6QZT3" authority="Nekhaev" authorityName="Nekhaev" authorityYear="2017" box="[151,493,151,178]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="schikovi" status="sp. nov.">
<emphasis id="2BD3EA8F0172FFE3E2C9FF58E205A188" bold="true" box="[151,376,151,177]" italics="true" pageId="3">Menestho schikovi</emphasis>
Nekhaev
</taxonomicName>
<taxonomicNameLabel id="30E057F40172FFE3E3AAFF57E14BA18B" box="[500,566,152,178]" pageId="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE3E2C9FF75E09BA1F6" box="[151,998,186,207]" pageId="3" type="reference_group">
<paragraph id="1918369D0172FFE3E2C9FF75E09BA1F6" blockId="3.[151,1436,151,2007]" box="[151,998,186,207]" pageId="3">
<treatmentCitationGroup id="39B711B30172FFE3E2C9FF75E09BA1F6" box="[151,998,186,207]" pageId="3">
<taxonomicName id="DEA74D1E0172FFE3E2C9FF75E104A1F6" ID-CoL="3ZQF3" authority=": Kantor &amp; Sysoev 2006: 243" authorityName=": Kantor &amp; Sysoev" authorityPageNumber="243" authorityYear="2006" box="[151,633,186,207]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="truncatula">
<emphasis id="2BD3EA8F0172FFE3E2C9FF75E218A1F6" box="[151,357,186,207]" italics="true" pageId="3">Menestho truncatula:</emphasis>
Kantor &amp;
<treatmentCitation id="9806108C0172FFE3E393FF75E104A1F6" author="Sysoev" box="[461,633,186,207]" page="243" pageId="3" year="2006">
<bibRefCitation id="7D364B6C0172FFE3E393FF75E136A1F6" author="Sysoev" box="[461,587,186,207]" pageId="3" refString="Kantor, Yu. I. &amp; Sysoev, A. V. (2006) Marine and brackish water Gastropoda of Russia and adjacent countries: an illustrated catalogue. KMK Scientific Press Ltd, Moscow, 371 pp., 140 pls." type="book" year="2006">Sysoev 2006</bibRefCitation>
: 243
</treatmentCitation>
</taxonomicName>
fig. 122С (partly), non
<bibRefCitation id="7D364B6C0172FFE3E103FF75E09EA1F6" author="Odhner" box="[861,995,186,207]" pageId="3" refString="Odhner, N. H. (1915) Die Molluskenfauna des Eisfjordes. In: Geer, G. (Ed.), Schwedischen Expedition nach Spitsbergen 1908. Teil II. Almqvist &amp; Wiksells Boktrvckeri, Stockholm, pp. 3 - 274." type="book chapter" year="1915">Odhner, 1915</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE3E2C9FF17E108A370" pageId="3" type="materials_examined">
<paragraph id="1918369D0172FFE3E2C9FF17E65BA028" blockId="3.[151,1436,151,2007]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E2C9FF17E243A1D6" bold="true" box="[151,318,216,239]" pageId="3">
<typeStatus id="C61C883F0172FFE3E2C9FF17E3B2A1D6" box="[151,207,216,239]" pageId="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="C61C883F0172FFE3E318FF17E2DEA1D6" box="[326,419,216,239]" pageId="3" type="holotype">holotype</typeStatus>
(empty shell) (
<figureCitation id="819C2A180172FFE3E015FF17E1D8A1D6" box="[587,677,216,239]" captionStart="FIGURE 1" captionStartId="1.[151,244,1908,1930]" captionTargetBox="[151,1436,191,1887]" captionTargetId="figure@1.[151,1436,191,1887]" captionTargetPageId="1" captionText="FIGURE 1. Arctic species of Menestho, photographs of shells. A, C, E — Menestho schikovi sp. n.: A — holotype, C — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m, E — Laptev Sea, 76 ° 36 ' N, 118 ° 30 ' E, 51 m; B — Menestho albula, Barents Sea, Murman Coast, Dalne- Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m; D — Menestho truncatula, Svalbard, Spitsbergen, Grønfjorden, 77 ° 57 ' N, 14 ° 16 ' E. Scale bar = 3 mm." httpUri="https://zenodo.org/record/1044822/files/figure.png" pageId="3">Figs 1A</figureCitation>
,
<figureCitation id="819C2A180172FFE3E0ECFF17E188A1D6" box="[690,757,216,239]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">2 A, I</figureCitation>
),
<date id="6D19105D0172FFE3E152FF17E005A1D6" box="[780,888,216,239]" pageId="3" value="2015-03-12">3.12.2015</date>
, van Veen grab,
<collectionCode id="7FB6AE580172FFE3E66EFF17E721A1D6" box="[1072,1116,216,239]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3">ZIN</collectionCode>
62135/1,
<specimenCount id="0FA1FD140172FFE3E695FF17E63AA1D6" box="[1227,1351,216,239]" pageId="3" type="generic" typeStatus="paratypes">6 paratypes</specimenCount>
(empty shells) from the same sample
<collectionCode id="7FB6AE580172FFE3E38AFF35E17DA028" box="[468,512,250,273]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3">ZIN</collectionCode>
62136/2 and
<specimenCount id="0FA1FD140172FFE3E0CCFF35E070A028" box="[658,781,250,273]" pageId="3" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
(empty shells) from the same sample
<collectionCode id="7FB6AE580172FFE3E6FDFF35E668A028" box="[1187,1301,250,273]" pageId="3">LMBI IN-</collectionCode>
3.
</paragraph>
<paragraph id="1918369D0172FFE3E299FED2E3CEA06E" blockId="3.[151,1436,151,2007]" pageId="3">
<materialsCitation id="A9CF3CC00172FFE3E299FED2E3CEA06E" ID-GBIF-Occurrence="1699887054" collectorName="Billefjorden, R &amp; V Dalnie Zelentsy" country="Svalbard and Jan Mayen" elevation="188" latitude="78.66" location="Spitsbergen Island" longLatPrecision="94" longitude="16.673334" pageId="3" specimenCount="188" typeStatus="holotype">
<emphasis id="2BD3EA8F0172FFE3E299FED2E21DA00D" bold="true" box="[199,352,285,308]" pageId="3">
<typeStatus id="C61C883F0172FFE3E299FED2E382A00D" box="[199,255,285,308]" pageId="3">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="61B0760D0172FFE3E339FED2E2B5A00D" box="[359,456,285,308]" name="Svalbard and Jan Mayen" pageId="3">Svalbard</collectingCountry>
,
<location id="1C7860460172FFE3E38BFED2E1E2A00D" LSID="urn:lsid:plazi:treatment:910E878B0172FFE4E2C9FF58E2E9A02E:1C7860460172FFE3E38BFED2E1E2A00D" box="[469,671,285,308]" country="Svalbard and Jan Mayen" latitude="78.66" longLatPrecision="94" longitude="16.673334" name="Spitsbergen Island" pageId="3">Spitsbergen Island</location>
,
<collectorName id="B452534B0172FFE3E0F5FED2E030A00D" box="[683,845,285,308]" pageId="3">Billefjorden, R</collectorName>
/
<collectorName id="B452534B0172FFE3E10BFED2E760A00D" box="[853,1053,285,308]" pageId="3">V Dalnie Zelentsy</collectorName>
, sta. 44,
<geoCoordinate id="7C93505A0172FFE3E6DEFED2E793A00D" box="[1152,1262,285,308]" direction="north" orientation="latitude" pageId="3" precision="92" value="78.66">78°39.6'N</geoCoordinate>
,
<geoCoordinate id="7C93505A0172FFE3E6A5FED2E61AA00D" box="[1275,1383,285,308]" direction="east" orientation="longitude" pageId="3" precision="92" value="16.673334">16°40.4'E</geoCoordinate>
,
<specimenCount id="0FA1FD140172FFE3E72DFED2E3CEA06E" pageId="3" type="generic">
<elevation id="928AD1AE0172FFE3E72DFED2E3CDA06E" metricMagnitude="2" metricUnit="m" metricValue="1.88" pageId="3" unit="m" value="188.0">
<quantity id="DE5F9B780172FFE3E72DFED2E3CDA06E" metricMagnitude="2" metricUnit="m" metricValue="1.88" pageId="3" unit="m" value="188.0">188 m</quantity>
</elevation>
.
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="1918369D0172FFE3E299FEADE108A370" blockId="3.[151,1436,151,2007]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FEADE286A040" bold="true" box="[199,507,354,377]" pageId="3">Other examined material:</emphasis>
3 spms, 2 shells, Kara Sea, 76°41'N, 88°14'E,
<quantity id="DE5F9B780172FFE3E67DFEADE71CA040" box="[1059,1121,354,377]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="3" unit="m" value="45.0">45 m</quantity>
, R/V G. Sedov, leg. Vagin, 0 6.09.1934, ZIN 22732 (studied with SEM); 4 spms, Kara Sea, 75°40'N, 80°26'E,
<quantity id="DE5F9B780172FFE3E66FFE4AE710A0A5" box="[1073,1133,389,412]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="3" unit="m" value="42.0">42 m</quantity>
, R/V G. Sedov, leg. Vagin, 0 9.09.1934, ZIN 35691; 3 spms, Kara Sea, 76°17'N, 75°08'E,
<quantity id="DE5F9B780172FFE3E172FE67E019A086" box="[812,868,424,447]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="3" unit="m" value="75.0">75 m</quantity>
, R/V G. Sedov, leg. Vagin,
<date id="6D19105D0172FFE3E6D8FE67E67DA086" box="[1158,1280,424,447]" pageId="3" value="1934-09-21">21.09.1934</date>
, ZIN 35693; 1 spm, Severnaya Zemlya, 80°30'N, 90°03E,
<quantity id="DE5F9B780172FFE3E0D8FE05E1ADA0D8" box="[646,720,458,481]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="3" unit="m" value="174.0">174 m</quantity>
, R/V Fyodor Litke,
<date id="6D19105D0172FFE3E1E6FE05E74FA0D8" box="[952,1074,458,481]" pageId="3" value="1948-09-29">29.09.1948</date>
, ZIN 35701; 1 spm,Laptev Sea, 77°25'N, 115°48'E,
<quantity id="DE5F9B780172FFE3E32BFE22E2CFA33D" box="[373,434,493,516]" metricMagnitude="1" metricUnit="m" metricValue="6.2" pageId="3" unit="m" value="62.0">62 m</quantity>
, R/V Sadko, leg. G.P. Gorbunov,
<date id="6D19105D0172FFE3E16AFE22E0D2A33D" box="[820,943,493,516]" pageId="3" value="1937-08-16">16.08.1937</date>
, ZIN 35696; 2 spms, Laptev Sea, 76°36'N, 118°30'E,
<quantity id="DE5F9B780172FFE3E358FDDFE242A31E" box="[262,319,528,551]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="3" unit="m" value="51.0">51 m</quantity>
, R/V Sadko, leg. G.P. Gorbunov, 0
<date id="6D19105D0172FFE3E0EDFDDFE05DA31E" box="[691,800,528,551]" pageId="3" value="1937-02-10">2.10.1937</date>
, ZIN 35697; 1 spm, 71°24'78''N, 174°22'03''E,
<quantity id="DE5F9B780172FFE3E743FDDFE61EA31E" box="[1309,1379,528,551]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="3" unit="m" value="585.0">585 m</quantity>
, R/V Professor Chromov,
<date id="6D19105D0172FFE3E32CFDFDE291A370" box="[370,492,562,585]" pageId="3" value="2004-08-22">22.08.2004</date>
, ZIN 60879.
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE3E299FD9AE6EEA355" box="[199,1427,597,620]" pageId="3" type="diagnosis">
<paragraph id="1918369D0172FFE3E299FD9AE6EEA355" blockId="3.[151,1436,151,2007]" box="[199,1427,597,620]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FD9AE240A355" bold="true" box="[199,317,597,620]" pageId="3">Diagnosis:</emphasis>
<taxonomicName id="DEA74D1E0172FFE3E31AFD9AE2D6A355" authorityName="Moller" authorityYear="1842" box="[324,427,597,620]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="genus">
<emphasis id="2BD3EA8F0172FFE3E31AFD9AE2D6A355" box="[324,427,597,620]" italics="true" pageId="3">Menestho</emphasis>
</taxonomicName>
with conical shell, convex, evenly rounded whorls, open umbilicus and straight growth-lines.
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE3E299FDB7E26BA2FE" pageId="3" type="description">
<paragraph id="1918369D0172FFE3E299FDB7E04CA205" blockId="3.[151,1436,151,2007]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FDB7E229A3B6" bold="true" box="[199,340,632,655]" pageId="3">Description:</emphasis>
Shell small, conical, white or yellowish with convex, evenly rounded whorls divided by deep suture (
<figureCitation id="819C2A180172FFE3E2C1FD55E23EA388" box="[159,323,666,689]" captionStart="FIGURE 1" captionStartId="1.[151,244,1908,1930]" captionTargetBox="[151,1436,191,1887]" captionTargetId="figure@1.[151,1436,191,1887]" captionTargetPageId="1" captionText="FIGURE 1. Arctic species of Menestho, photographs of shells. A, C, E — Menestho schikovi sp. n.: A — holotype, C — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m, E — Laptev Sea, 76 ° 36 ' N, 118 ° 30 ' E, 51 m; B — Menestho albula, Barents Sea, Murman Coast, Dalne- Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m; D — Menestho truncatula, Svalbard, Spitsbergen, Grønfjorden, 77 ° 57 ' N, 14 ° 16 ' E. Scale bar = 3 mm." httpUri="https://zenodo.org/record/1044822/files/figure.png" pageId="3">Figs. 1 A, C, E</figureCitation>
,
<figureCitation id="819C2A180172FFE3E310FD55E20EA388" box="[334,371,666,689]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">2A</figureCitation>
). Shell surface with fine, frequent (811 on body whorl), spiral ribs, almost semicircular in section, divided by very narrow interspaces (
<figureCitation id="819C2A180172FFE3E06BFD72E1F8A3ED" box="[565,645,701,724]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">
Fig.
<date id="6D19105D0172FFE3E032FD72E1F8A3ED" box="[620,645,701,724]" pageId="3">2I</date>
</figureCitation>
). Axial sculpture, except for straight growth-lines (
<figureCitation id="819C2A180172FFE3E697FD72E641A3ED" box="[1225,1340,701,724]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2A, I</figureCitation>
) absent. Embryonic shell intorted, of
<typeStatus id="C61C883F0172FFE3E38DFD2FE17CA3CE" box="[467,513,736,759]" pageId="3">type</typeStatus>
B, partially eroded on studied specimens, smooth or with very low pattern of spiral sculpture (
<figureCitation id="819C2A180172FFE3E354FCCDE219A220" box="[266,356,770,793]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2G</figureCitation>
). Aperture oval with acute angle in upper part, no teeth or folds on columella. Umbilicus clearly visible, narrow. Operculum transparent, without peg (
<figureCitation id="819C2A180172FFE3E08CFCEAE058A205" box="[722,805,805,828]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="1918369D0172FFE3E299FC87E166A2B8" blockId="3.[151,1436,151,2007]" pageId="3">
Measurements of
<typeStatus id="C61C883F0172FFE3E3DAFC87E29BA266" box="[388,486,840,863]" pageId="3" type="holotype">holotype</typeStatus>
: shell height =
<quantity id="DE5F9B780172FFE3E0DBFC87E1A4A266" box="[645,729,840,863]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="3" unit="mm" value="3.6">3.6 mm</quantity>
, aperture height =
<quantity id="DE5F9B780172FFE3E1C0FC87E08CA266" box="[926,1009,840,863]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" unit="mm" value="1.2">1.2 mm</quantity>
, last whorl height =
<quantity id="DE5F9B780172FFE3E697FC87E661A266" box="[1225,1308,840,863]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" unit="mm" value="2.3">2.3 mm</quantity>
, shell width = 2.0 mm, aperture width = 1.0 mm.
</paragraph>
<paragraph id="1918369D0172FFE3E299FC42E26BA2FE" blockId="3.[151,1436,151,2007]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FC42E23BA29D" bold="true" box="[199,326,909,932]" pageId="3">Variability.</emphasis>
Studied shells differ in width/height proportions (
<figureCitation id="819C2A180172FFE3E133FC42E777A29D" box="[877,1034,909,932]" captionStart="FIGURE 1" captionStartId="1.[151,244,1908,1930]" captionTargetBox="[151,1436,191,1887]" captionTargetId="figure@1.[151,1436,191,1887]" captionTargetPageId="1" captionText="FIGURE 1. Arctic species of Menestho, photographs of shells. A, C, E — Menestho schikovi sp. n.: A — holotype, C — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m, E — Laptev Sea, 76 ° 36 ' N, 118 ° 30 ' E, 51 m; B — Menestho albula, Barents Sea, Murman Coast, Dalne- Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m; D — Menestho truncatula, Svalbard, Spitsbergen, Grønfjorden, 77 ° 57 ' N, 14 ° 16 ' E. Scale bar = 3 mm." httpUri="https://zenodo.org/record/1044822/files/figure.png" pageId="3">Fig 1 A, C, E</figureCitation>
), no other significant variability had been found.
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE3E299FC1DE181A5AE" pageId="3" type="distribution">
<paragraph id="1918369D0172FFE3E299FC1DE065A516" blockId="3.[151,1436,151,2007]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FC1DE224A2D0" bold="true" box="[199,345,978,1001]" pageId="3">Distribution.</emphasis>
Known from the
<collectingCountry id="61B0760D0172FFE3E043FC1DE103A2D0" box="[541,638,978,1001]" name="Svalbard and Jan Mayen" pageId="3">Svalbard</collectingCountry>
, the Kara and the Laptev seas. Single young specimen (ZIN 60879) with shell height =
<quantity id="DE5F9B780172FFE3E369FC3AE2F6A535" box="[311,395,1013,1036]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" unit="mm" value="2.5">2.5 mm</quantity>
corresponded to
<taxonomicName id="DEA74D1E0172FFE3E015FC3AE06EA535" authorityName="Nekhaev" authorityYear="2017" box="[587,787,1013,1036]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="schikovi">
<emphasis id="2BD3EA8F0172FFE3E015FC3AE06EA535" box="[587,787,1013,1036]" italics="true" pageId="3">Menestho schikovi</emphasis>
</taxonomicName>
was found from the Chukchi Sea. Depth range
<quantity id="DE5F9B780172FFE3E779FC3AE6E5A535" box="[1319,1432,1013,1036]" metricMagnitude="2" metricUnit="m" metricValue="3.135" metricValueMax="5.85" metricValueMin="0.42" pageId="3" unit="m" value="313.5" valueMax="585.0" valueMin="42.0">
42
<specimenCount id="0FA1FD140172FFE3E70EFC3AE6E5A535" box="[1360,1432,1013,1036]" pageId="3" type="generic">585 m</specimenCount>
</quantity>
. Probably, the species lives in all seas of the Eurasian Arctic.
</paragraph>
<paragraph id="1918369D0172FFE3E299FBF5E181A5AE" blockId="3.[151,1436,151,2007]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FBF5E258A568" bold="true" box="[199,293,1082,1105]" pageId="3">Habitat.</emphasis>
The species had been collected from silt substrates. The observed near-bottom temperature and salinity at the
<typeStatus id="C61C883F0172FFE3E2E1FB92E390A54D" box="[191,237,1117,1140]" pageId="3">type</typeStatus>
locality were -1.4°C and 34.8 psu respectively.
<taxonomicName id="DEA74D1E0172FFE3E0A7FB92E0C2A54D" authorityName="Nekhaev" authorityYear="2017" box="[761,959,1117,1140]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="schikovi">
<emphasis id="2BD3EA8F0172FFE3E0A7FB92E0C2A54D" box="[761,959,1117,1140]" italics="true" pageId="3">Menestho schikovi</emphasis>
</taxonomicName>
probably have a parasitic mode of life, like other pyramidellid species. However its host is unknown.
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE3E299FB6DE620A580" box="[199,1373,1186,1210]" pageId="3" type="description">
<paragraph id="1918369D0172FFE3E299FB6DE620A580" blockId="3.[151,1436,151,2007]" box="[199,1373,1186,1210]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FB6DE2D4A583" bold="true" box="[199,425,1186,1210]" pageId="3">Derivation of name.</emphasis>
The species is named after Dr. Evgenij Schikov, enthusiastic malacologist and ecologist.
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE3E299FB0AE1CEA45E" pageId="3" type="discussion">
<paragraph id="1918369D0172FFE3E299FB0AE1CEA45E" blockId="3.[151,1436,151,2007]" pageId="3">
<emphasis id="2BD3EA8F0172FFE3E299FB0AE248A5E5" bold="true" box="[199,309,1221,1244]" pageId="3">Remarks.</emphasis>
Shell of
<taxonomicName id="DEA74D1E0172FFE3E3CBFB0AE10EA5E5" authorityName=": Kantor &amp; Sysoev" authorityYear="2006" box="[405,627,1221,1244]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="truncatula">
<emphasis id="2BD3EA8F0172FFE3E3CBFB0AE10EA5E5" box="[405,627,1221,1244]" italics="true" pageId="3">Menestho truncatula</emphasis>
</taxonomicName>
(
<figureCitation id="819C2A180172FFE3E0DFFB0AE1A2A5E5" box="[641,735,1221,1244]" captionStart="FIGURE 1" captionStartId="1.[151,244,1908,1930]" captionTargetBox="[151,1436,191,1887]" captionTargetId="figure@1.[151,1436,191,1887]" captionTargetPageId="1" captionText="FIGURE 1. Arctic species of Menestho, photographs of shells. A, C, E — Menestho schikovi sp. n.: A — holotype, C — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m, E — Laptev Sea, 76 ° 36 ' N, 118 ° 30 ' E, 51 m; B — Menestho albula, Barents Sea, Murman Coast, Dalne- Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m; D — Menestho truncatula, Svalbard, Spitsbergen, Grønfjorden, 77 ° 57 ' N, 14 ° 16 ' E. Scale bar = 3 mm." httpUri="https://zenodo.org/record/1044822/files/figure.png" pageId="3">Figs. 1D</figureCitation>
,
<figureCitation id="819C2A180172FFE3E0B5FB0AE076A5E5" box="[747,779,1221,1244]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">2B</figureCitation>
) (see also
<bibRefCitation id="7D364B6C0172FFE3E122FB0AE767A5E5" author="Odhner" box="[892,1050,1221,1244]" pageId="3" refString="Odhner, N. H. (1915) Die Molluskenfauna des Eisfjordes. In: Geer, G. (Ed.), Schwedischen Expedition nach Spitsbergen 1908. Teil II. Almqvist &amp; Wiksells Boktrvckeri, Stockholm, pp. 3 - 274." type="book chapter" year="1915">Odhner (1915)</bibRefCitation>
for the figures of
<typeStatus id="C61C883F0172FFE3E684FB0AE644A5E5" box="[1242,1337,1221,1244]" pageId="3" type="syntype">syntypes</typeStatus>
) is wider than
<taxonomicName id="DEA74D1E0172FFE3E295FB27E238A5C6" authorityName="Nekhaev" authorityYear="2017" box="[203,325,1256,1279]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="schikovi">
<emphasis id="2BD3EA8F0172FFE3E295FB27E238A5C6" box="[203,325,1256,1279]" italics="true" pageId="3">M. schikovi</emphasis>
</taxonomicName>
, upper part of its last whorl with flattened shoulder, not rounded. The aperture height in
<taxonomicName id="DEA74D1E0172FFE3E6AFFB27E6F9A5C6" authorityName=": Kantor &amp; Sysoev" authorityYear="2006" box="[1265,1412,1256,1279]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="truncatula">
<emphasis id="2BD3EA8F0172FFE3E6AFFB27E6F9A5C6" box="[1265,1412,1256,1279]" italics="true" pageId="3">M. truncatula</emphasis>
</taxonomicName>
is about a half of shell height. In addition,
<taxonomicName id="DEA74D1E0172FFE3E00EFAC5E199A418" authorityName=": Kantor &amp; Sysoev" authorityYear="2006" box="[592,740,1290,1313]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="truncatula">
<emphasis id="2BD3EA8F0172FFE3E00EFAC5E199A418" box="[592,740,1290,1313]" italics="true" pageId="3">M. truncatula</emphasis>
</taxonomicName>
have a reticulate pattern on the protoconch (
<figureCitation id="819C2A180172FFE3E693FAC5E662A418" box="[1229,1311,1290,1313]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2F</figureCitation>
).
<taxonomicName id="DEA74D1E0172FFE3E76AFAC5E3A6A47D" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">
<emphasis id="2BD3EA8F0172FFE3E76AFAC5E3A6A47D" italics="true" pageId="3">Menestho albula</emphasis>
</taxonomicName>
differs from
<taxonomicName id="DEA74D1E0172FFE3E334FAE2E29BA47D" authorityName="Nekhaev" authorityYear="2017" box="[362,486,1325,1348]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="schikovi">
<emphasis id="2BD3EA8F0172FFE3E334FAE2E29BA47D" box="[362,486,1325,1348]" italics="true" pageId="3">M. schikovi</emphasis>
</taxonomicName>
by prosocline growth-lines and absence of an umbilicus. Usually
<taxonomicName id="DEA74D1E0172FFE3E6F1FAE2E61FA47D" box="[1199,1378,1325,1348]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">
<emphasis id="2BD3EA8F0172FFE3E6F1FAE2E61FA47D" box="[1199,1378,1325,1348]" italics="true" pageId="3">Menestho albula</emphasis>
</taxonomicName>
have coarser spiral sculpture and more flattened whorls.
</paragraph>
</subSubSection>
<subSubSection id="51BD65160172FFE4E299FABDE2E9A02E" lastPageId="4" pageId="3" type="reference_group">
<paragraph id="1918369D0172FFE3E299FABDE6F5A656" blockId="3.[151,1436,151,2007]" pageId="3">
<taxonomicName id="DEA74D1E0172FFE3E299FABDE2EDA4B0" ID-CoL="6QZT3" authorityName="Nekhaev" authorityYear="2017" box="[199,400,1394,1417]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="schikovi">
<emphasis id="2BD3EA8F0172FFE3E299FABDE2EDA4B0" box="[199,400,1394,1417]" italics="true" pageId="3">Menestho schikovi</emphasis>
</taxonomicName>
very resembles
<emphasis id="2BD3EA8F0172FFE3E017FABDE07BA4B0" box="[585,774,1394,1417]" italics="true" pageId="3">
<taxonomicName id="DEA74D1E0172FFE3E017FABDE07FA4B0" ID-CoL="6QZR5" box="[585,770,1394,1417]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">Menestho albula</taxonomicName>
,
</emphasis>
which was described from West
<collectingCountry id="61B0760D0172FFE3E622FABDE792A4B0" box="[1148,1263,1394,1417]" name="Greenland" pageId="3">Greenland</collectingCountry>
. However,
<typeStatus id="C61C883F0172FFE3E730FABDE6E1A4B0" box="[1390,1436,1394,1417]" pageId="3">type</typeStatus>
material of
<taxonomicName id="DEA74D1E0172FFE3E349FA5AE2D3A495" box="[279,430,1429,1452]" class="Gastropoda" family="Turbinidae" genus="Turbo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="3" phylum="Mollusca" rank="species" species="albulus">
<emphasis id="2BD3EA8F0172FFE3E349FA5AE2D3A495" box="[279,430,1429,1452]" italics="true" pageId="3">Turbo albulus</emphasis>
</taxonomicName>
had not been found neither during previous studies nor during my own examination of the collections of the Natural History Museum of
<collectingCountry id="61B0760D0172FFE3E0D6FA77E193A4F6" box="[648,750,1464,1487]" name="Denmark" pageId="3">Denmark</collectingCountry>
, where the collections by Otto Fabricius are stored (
<bibRefCitation id="7D364B6C0172FFE3E77EFA77E388A4C8" author="Nekhaev" pageId="3" refString="Nekhaev, I. O., Schiotte, T. &amp; Vinarski, M. V. (2015) Type materials of European freshwater molluscs described by Otto Friedrich Muller. Archiv fur Molluskenkunde, 144 (1), 51 - 64. https: // doi. org / 10.1127 / arch. moll / 1869 - 0963 / 144 / 051 - 064" type="journal article" year="2015">Nekhaev et al. 2015</bibRefCitation>
). Hence, current knowledge on taxonomic status of
<taxonomicName id="DEA74D1E0172FFE3E165FA15E08CA4C8" ID-CoL="6QZR5" box="[827,1009,1498,1521]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">
<emphasis id="2BD3EA8F0172FFE3E165FA15E08CA4C8" box="[827,1009,1498,1521]" italics="true" pageId="3">Menestho albula</emphasis>
</taxonomicName>
is mainly based on topotypic samples collected by H.P.C. Møller (
<bibRefCitation id="7D364B6C0172FFE3E397FA32E12CA72D" author="Waren" box="[457,593,1533,1556]" pageId="3" refString="Waren, A. (1991) New and little known Mollusca from Iceland and Scandinavia. Sarsia, 76, 53 - 124. https: // doi. org / 10.1080 / 00364827.1991.10413466" type="journal article" year="1991">Warén 1991</bibRefCitation>
). A single shell from ZIN corresponds with the original description of
<taxonomicName id="DEA74D1E0172FFE3E702FA32E398A70E" class="Gastropoda" family="Turbinidae" genus="Turbo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="3" phylum="Mollusca" rank="species" species="albulus">
<emphasis id="2BD3EA8F0172FFE3E702FA32E398A70E" italics="true" pageId="3">Turbo albulus</emphasis>
</taxonomicName>
in shell shape, size and sculpture as well as with specimens from
<collectingCountry id="61B0760D0172FFE3E1C0F9EFE091A70E" box="[926,1004,1568,1591]" name="Iceland" pageId="3">Iceland</collectingCountry>
studied by
<bibRefCitation id="7D364B6C0172FFE3E639F9EFE784A70E" author="Waren" box="[1127,1273,1568,1591]" pageId="3" refString="Waren, A. (1991) New and little known Mollusca from Iceland and Scandinavia. Sarsia, 76, 53 - 124. https: // doi. org / 10.1080 / 00364827.1991.10413466" type="journal article" year="1991">Warén (1991)</bibRefCitation>
and specimens from the Murman Coast studied in this work. All mollusks have small (
<quantity id="DE5F9B780172FFE3E1D1F98DE09CA760" box="[911,993,1602,1625]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" unit="mm" value="3.5">3.5 mm</quantity>
in height and lesser) conical, white, shell with flattened whorls (
<figureCitation id="819C2A180172FFE3E3D4F9AAE2A3A745" box="[394,478,1637,1660]" captionStart="FIGURE 1" captionStartId="1.[151,244,1908,1930]" captionTargetBox="[151,1436,191,1887]" captionTargetId="figure@1.[151,1436,191,1887]" captionTargetPageId="1" captionText="FIGURE 1. Arctic species of Menestho, photographs of shells. A, C, E — Menestho schikovi sp. n.: A — holotype, C — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m, E — Laptev Sea, 76 ° 36 ' N, 118 ° 30 ' E, 51 m; B — Menestho albula, Barents Sea, Murman Coast, Dalne- Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m; D — Menestho truncatula, Svalbard, Spitsbergen, Grønfjorden, 77 ° 57 ' N, 14 ° 16 ' E. Scale bar = 3 mm." httpUri="https://zenodo.org/record/1044822/files/figure.png" pageId="3">Fig. 1B</figureCitation>
,
<figureCitation id="819C2A180172FFE3E3B4F9AAE153A745" box="[490,558,1637,1660]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">2CD</figureCitation>
). The reported size of
<quantity id="DE5F9B780172FFE3E17FF9AAE022A745" box="[801,863,1637,1660]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" unit="mm" value="9.0">9 mm</quantity>
for
<taxonomicName id="DEA74D1E0172FFE3E1D0F9AAE73CA745" ID-CoL="6QZR5" box="[910,1089,1637,1660]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">
<emphasis id="2BD3EA8F0172FFE3E1D0F9AAE73CA745" box="[910,1089,1637,1660]" italics="true" pageId="3">Menestho albula</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation id="7D364B6C0172FFE3E6DEF9AAE64DA745" author="Rosenberg" box="[1152,1328,1637,1660]" pageId="3" refString="Rosenberg, G. (2009) Malacolog Ver. 4.1. 1. Available from http: // www. malacolog. org / (accessed 16 April 2016)" type="journal article" year="2009">Rosenberg 2009</bibRefCitation>
) seems to be a result of confusion with another species. Spiral ribs of
<taxonomicName id="DEA74D1E0172FFE3E114F947E77CA7A6" ID-CoL="6QZR5" box="[842,1025,1672,1695]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">
<emphasis id="2BD3EA8F0172FFE3E114F947E77CA7A6" box="[842,1025,1672,1695]" italics="true" pageId="3">Menestho albula</emphasis>
</taxonomicName>
are usually of the same width with interspaces (
<figureCitation id="819C2A180172FFE3E343F965E2E9A7F8" box="[285,404,1706,1729]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2 JK</figureCitation>
). Some of the studied specimens from the Murman Coast have a «bend» on the suture (
<figureCitation id="819C2A180172FFE3E716F965E6EEA7F8" box="[1352,1427,1706,1729]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2J</figureCitation>
) whereas others have not. The bend also has not been found in the topotypic specimen (
<figureCitation id="819C2A180172FFE3E605F902E7CFA7DD" box="[1115,1202,1741,1764]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2K</figureCitation>
) and on a shell from
<collectingCountry id="61B0760D0172FFE3E2C9F93FE398A63E" box="[151,229,1776,1799]" name="Iceland" pageId="3">Iceland</collectingCountry>
depicted by
<bibRefCitation id="7D364B6C0172FFE3E32EF93FE10CA63E" author="Waren" box="[368,625,1776,1799]" pageId="3" refString="Waren, A. (1991) New and little known Mollusca from Iceland and Scandinavia. Sarsia, 76, 53 - 124. https: // doi. org / 10.1080 / 00364827.1991.10413466" type="journal article" year="1991">Warén (1991: Fig. 27F)</bibRefCitation>
, and hence, with limited knowledge available to date, I suggest that it is an intraspecific variation. All specimens had clearly prosocline growth-lines (
<figureCitation id="819C2A180172FFE3E1ECF8DDE71BA610" box="[946,1126,1810,1833]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2 CD, JK</figureCitation>
). The umbilicus of
<taxonomicName id="DEA74D1E0172FFE3E76AF8DDE3A6A675" ID-CoL="6QZR5" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">
<emphasis id="2BD3EA8F0172FFE3E76AF8DDE3A6A675" italics="true" pageId="3">Menestho albula</emphasis>
</taxonomicName>
is closed, its embryonic shell is smooth (
<figureCitation id="819C2A180172FFE3E0C1F8FAE18AA675" box="[671,759,1845,1868]" captionStart="FIGURE 2" captionStartId="2.[151,244,1822,1844]" captionTargetBox="[185,1409,179,1809]" captionTargetId="figure@2.[185,1409,179,1809]" captionTargetPageId="2" captionText="FIGURE 2. Teleoconchs (A-D), operculum (E), embryonic shells (F-H) and details of sculpture (I-K) of the Arctic species of Menestho, SEM micrographs. A, E, G, I — Menestho schikovi sp. n.: A, I — holotype, E, G — Kara Sea, 76 ° 41 ' N, 88 ° 14 ' E, 45 m (G - same specimen as on Fig. 1 C); B, F — Menestho truncatula, Kara Sea, 79 ° 17.2 ' N, 76 ° 03 ' E, 54 m; C-D, H, J-K — Menestho albula, C, J — Barents Sea, Murman Coast, Dalne-Zelenetskaya Bay, 69 ° 07.85 ' N, 36 ° 04.81 ' E, 46 m (same specimen as on Fig. 1 B), H — Barents Sea, Murman Coast, Kola Inlet, 69 ° 18.90 ' N, 33 ° 29.08 ' E, 43 m, D, K — between Labrador and Greenland; Scale bars: A-D at same scale bar = 3 mm, E = 500 µm, F-H at same scale bar = 200 µm, scale bars for I, J, K = 200 µm. Arrows indicate direction of growthlines." httpUri="https://zenodo.org/record/1044824/files/figure.png" pageId="3">Fig. 2H</figureCitation>
).
<taxonomicName id="DEA74D1E0172FFE3E155F8FAE0C2A675" ID-CoL="6QZR5" box="[779,959,1845,1868]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="albula">
<emphasis id="2BD3EA8F0172FFE3E155F8FAE0C2A675" box="[779,959,1845,1868]" italics="true" pageId="3">Menestho albula</emphasis>
</taxonomicName>
have an Atlantic distribution: the species is known in the waters around
<collectingCountry id="61B0760D0172FFE3E39BF897E14AA656" box="[453,567,1880,1903]" name="Greenland" pageId="3">Greenland</collectingCountry>
, North of
<collectingCountry id="61B0760D0172FFE3E0FDF897E189A656" box="[675,756,1880,1903]" name="Iceland" pageId="3">Iceland</collectingCountry>
,
<collectingCountry id="61B0760D0172FFE3E0A1F897E023A656" box="[767,862,1880,1903]" name="Svalbard and Jan Mayen" pageId="3">Svalbard</collectingCountry>
and SW Barents Sea (
<bibRefCitation id="7D364B6C0172FFE3E612F897E7AFA656" author="Waren" box="[1100,1234,1880,1903]" pageId="3" refString="Waren, A. (1991) New and little known Mollusca from Iceland and Scandinavia. Sarsia, 76, 53 - 124. https: // doi. org / 10.1080 / 00364827.1991.10413466" type="journal article" year="1991">Warén 1991</bibRefCitation>
;
<bibRefCitation id="7D364B6C0172FFE3E683F897E601A656" author="Nekhaev" box="[1245,1404,1880,1903]" pageId="3" refString="Nekhaev, I. O. (2014) Marine shell-bearing Gastropoda of Murman (Barents Sea): an annotated check-list. Ruthenica, Russian Malacological Journal, 24, 75 - 121." type="journal article" year="2014">Nekhaev 2014</bibRefCitation>
).
</paragraph>
<paragraph id="1918369D0172FFE4E299F8B5E2E9A02E" blockId="3.[151,1436,151,2007]" lastBlockId="4.[151,1436,152,279]" lastPageId="4" pageId="3">
Examination of the museum collection revealed that
<taxonomicName id="DEA74D1E0172FFE3E158F8B5E0B0A6A8" ID-CoL="6QZT3" authorityName="Nekhaev" authorityYear="2017" box="[774,973,1914,1937]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="schikovi">
<emphasis id="2BD3EA8F0172FFE3E158F8B5E0B0A6A8" box="[774,973,1914,1937]" italics="true" pageId="3">Menestho schikovi</emphasis>
</taxonomicName>
had been usually confused with
<emphasis id="2BD3EA8F0172FFE3E76AF8B5E270A68D" italics="true" pageId="3">
<taxonomicName id="DEA74D1E0172FFE3E76AF8B5E274A68D" ID-CoL="3ZQF3" authorityName=": Kantor &amp; Sysoev" authorityYear="2006" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="truncatula">Menestho truncatula</taxonomicName>
.
</emphasis>
A specimen of the former species was also mistakenly depicted by Kantor &amp;
<bibRefCitation id="7D364B6C0172FFE3E630F852E676A68D" box="[1134,1291,1949,1972]" pageId="3" refString="Kantor, Yu. I. &amp; Sysoev, A. V. (2006) Marine and brackish water Gastropoda of Russia and adjacent countries: an illustrated catalogue. KMK Scientific Press Ltd, Moscow, 371 pp., 140 pls." type="book">Sysoev (2006)</bibRefCitation>
as
<taxonomicName id="DEA74D1E0172FFE3E76AF852E27BA6EE" ID-CoL="3ZQF3" authorityName=": Kantor &amp; Sysoev" authorityYear="2006" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="3" phylum="Mollusca" rank="species" species="truncatula">
<emphasis id="2BD3EA8F0172FFE3E76AF852E27BA6EE" italics="true" pageId="3">Menestho truncatula</emphasis>
</taxonomicName>
. Examination of museum collections showed that both species occur sympatrically in the Arctic. However,
<taxonomicName id="DEA74D1E0175FFE4E2C9FF57E222A196" ID-CoL="6QZT3" authorityName="Nekhaev" authorityYear="2017" box="[151,351,152,175]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" phylum="Mollusca" rank="species" species="schikovi">
<emphasis id="2BD3EA8F0175FFE4E2C9FF57E222A196" box="[151,351,152,175]" italics="true" pageId="4">Menestho schikovi</emphasis>
</taxonomicName>
had not been found in the Barents and White seas yet.
<taxonomicName id="DEA74D1E0175FFE4E194FF57E7D7A196" ID-CoL="3ZQF3" authorityName=": Kantor &amp; Sysoev" authorityYear="2006" box="[970,1194,152,175]" class="Gastropoda" family="Pyramidellidae" genus="Menestho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" phylum="Mollusca" rank="species" species="truncatula">
<emphasis id="2BD3EA8F0175FFE4E194FF57E7D7A196" box="[970,1194,152,175]" italics="true" pageId="4">Menestho truncatula</emphasis>
</taxonomicName>
seems to be the most common representative of the family in the Arctic and has been found with certainty from
<collectingCountry id="61B0760D0175FFE4E6D4FF75E796A1E8" box="[1162,1259,186,209]" name="Svalbard and Jan Mayen" pageId="4">Svalbard</collectingCountry>
, all parts of the Barents Sea, the Kara, White, Laptev and East-Siberian Seas (
<bibRefCitation id="7D364B6C0175FFE4E109FF12E089A1CD" author="Golikov" box="[855,1012,221,244]" pageId="4" refString="Golikov, A. N. (1987) Classis Gastropoda (part.). In: Starobogatov, Ya. I., Naumov, A. D. (Eds.), Molluscs of the White Sea. Nauka, Leningrad, pp. 41 - 148 [in Russian]" type="book chapter" year="1987">Golikov 1987</bibRefCitation>
;
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</paragraph>
</subSubSection>
</treatment>
</document>