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<document id="D14F410A99ACEDBDE7033945E3275EF1" ID-DOI="10.5281/zenodo.5355307" ID-ISSN="2345-7600" ID-Zenodo-Dep="5355307" ID-ZooBank="88473619-8A6C-4951-AD3C-89C14653CAC3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630215238892" checkinUser="valdenar" docAuthor="Ng, Peter K. L., Davie, Peter J. F. &amp; Li, Jheng-Jhang" docDate="2016" docId="84169C569355FFD22BC99BE3FDCC0526" docLanguage="en" docName="64rbz257-268.pdf" docOrigin="Raffles Bulletin of Zoology 64" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Parasesarma sigillatum" docType="treatment" docVersion="3" lastPageNumber="264" masterDocId="782FE42E9351FFD52B2C9F5DFFB20D28" masterDocTitle="On the identities of Parasesarma carolinense (Rathbun, 1907) and Parasesarma sigillatum (Tweedie, 1950), with description of a new species from Taiwan (Crustacea: Brachyura: Sesarmidae)" masterLastPageNumber="268" masterPageNumber="257" pageNumber="261" updateTime="1699189230292" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="95B1741CDBFD1FA08763DD0F41D12C59">On the identities of Parasesarma carolinense (Rathbun, 1907) and Parasesarma sigillatum (Tweedie, 1950), with description of a new species from Taiwan (Crustacea: Brachyura: Sesarmidae)</mods:title>
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<taxonomicName id="CBBF56C39355FFD12BC99BE3FD1B09FD" ID-CoL="4DBZQ" authority="(Tweedie, 1950)" baseAuthorityName="Tweedie" baseAuthorityYear="1950" box="[229,681,1213,1237]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="sigillatum">
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<emphasis id="3ECBF1529355FFD12BC99BE3FE5E09FD" bold="true" box="[229,492,1213,1237]" italics="true" pageId="4" pageNumber="261">Parasesarma sigillatum</emphasis>
(
<bibRefCitation id="682E50B19355FFD12AD79BE0FD1009FD" author="Tweedie MWF" box="[507,674,1213,1237]" pageId="4" pageNumber="261" pagination="105 - 148" refId="ref7623" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105 - 148, figs. 1 - 4, pls. 16, 17." type="journal article" year="1950">Tweedie, 1950</bibRefCitation>
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<paragraph id="0C002D409355FFD12A299B80FD3809DD" blockId="4.[229,681,1213,1269]" box="[261,650,1245,1269]" pageId="4" pageNumber="261">
(
<figureCitation id="948431C59355FFD12A219B80FE3D09DD" box="[269,399,1245,1269]" captionStart="Fig" captionStartId="1.[113,148,1887,1909]" captionTargetBox="[113,1451,308,1872]" captionTargetId="figure-15@1.[113,1451,308,1872]" captionTargetPageId="1" captionText="Fig. 1. Colour in life. AD, Parasesarma sigillatum, Cocos-Keeling Islands; EH, Parasesarma corallicum n. sp., Taiwan. A, male, not preserved; B, female (8.7 × 6.4 mm) (QM-W28343); C, D, male (9.4 × 6.9 mm) (QM-W28342); E, F, male, Hsiangchiaowan, Taiwan (10.3 × 7.6 mm) (NMMBCD 4698); G, Holotype, same data (9.5 × 7.0 mm) (NMMBCD 4697); H, Wanlitung, Taiwan (8.0 × 5.7 mm) (NMNS-7548-002)." figureDoi="http://doi.org/10.5281/zenodo.5355309" httpUri="https://zenodo.org/record/5355309/files/figure.png" pageId="4" pageNumber="261">Figs. 1AD</figureCitation>
;
<figureCitation id="948431C59355FFD12AB09B80FE1909DD" box="[412,427,1245,1269]" captionStart="Fig" captionStartId="5.[113,148,1948,1970]" captionTargetBox="[113,1451,248,1933]" captionTargetId="figure-92@5.[113,1451,248,1933]" captionTargetPageId="5" captionText="Fig. 3. Parasesarma sigillatum (Tweedie, 1950). AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="4" pageNumber="261">3</figureCitation>
;
<figureCitation id="948431C59355FFD12A969B80FE4C09DD" box="[442,510,1245,1269]" captionStart="Fig" captionStartId="7.[113,148,1491,1513]" captionTargetBox="[113,1451,189,1476]" captionTargetId="figure-93@7.[113,1451,189,1476]" captionTargetPageId="7" captionText="Fig. 5. Left ambulatory legs. AC, Parasesarma carolinense (Rathbun, 1914), holotype male (9.2 × 7.0 mm) (USNM 32861), Caroline Islands; DG, Parasesarma sigillatum (Tweedie, 1950), male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; HK, Parasesarma corallicum n. sp., holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. A, D, H, first ambulatory leg; B, E, I, second ambulatory leg; C, F, J, third ambulatory leg; G, K, fourth ambulatory leg. Legs of all species to scale." figureDoi="http://doi.org/10.5281/zenodo.5355317" httpUri="https://zenodo.org/record/5355317/files/figure.png" pageId="4" pageNumber="261">5DG</figureCitation>
;
<figureCitation id="948431C59355FFD129279B80FDE609DD" box="[523,596,1245,1269]" captionStart="Fig" captionStartId="8.[135,170,1920,1942]" captionTargetBox="[189,1437,184,1908]" captionTargetId="figure-8@8.[189,1437,184,1911]" captionTargetPageId="8" captionText="Fig. 6. AF, Parasesarma carolinense (Rathbun, 1914), holotype male (9.2 × 7.0 mm) (USNM 32861), Caroline Islands; GM, Parasesarma sigillatum (Tweedie, 1950), male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; NT, Parasesarma corallicum n. sp., holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. A, G, N, left third maxilliped; B, H, male pleon; O, male pleon (somites 1 and 3 broken); C, P, ventral view of left G1; D, Q, dorsal view of left G1; E, ventral view of right G1; F, distal chitinous part of right G1 (ventral view); I, ventral view of right G1; J, dorsal view of right G1; K, distal chitinous part of right G1 (ventral view); L, distal chitinous part of right G1 (dorsal view); M, right G2; R, distal chitinous part of left G1 (ventral view); S, distal chitinous part of left G1 (dorsal view); T, left G2. Setae for all structures denuded. Scale bars = 0.5 mm [AE, GJ, MQ, T]; 0.1 mm [F, K, L, R, S]." figureDoi="http://doi.org/10.5281/zenodo.5355319" httpUri="https://zenodo.org/record/5355319/files/figure.png" pageId="4" pageNumber="261">6GM</figureCitation>
;
<figureCitation id="948431C59355FFD1294D9B80FD3009DD" box="[609,642,1245,1269]" captionStart="Fig" captionStartId="9.[113,148,778,800]" captionTargetBox="[112,1450,192,765]" captionTargetId="figure-672@9.[100,1463,180,772]" captionTargetPageId="9" captionText="Fig. 7. Details of pectinate crests on chela. A, Parasesarma sigillatum (Tweedie, 1950), lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island; B, Parasesarma corallicum n. sp., holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. Arrow points to direction of dactylar finger. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5355321" httpUri="https://zenodo.org/record/5355321/files/figure.png" pageId="4" pageNumber="261">7A</figureCitation>
)
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<treatmentCitation id="8D1E0B519355FFD12BA49A41FE4C0819" author="Tweedie MWF" box="[136,510,1308,1330]" page="131" pageId="4" pageNumber="261" year="1950">
<taxonomicName id="CBBF56C39355FFD12BA49A41FE4C0819" ID-CoL="4WZ28" authority="Tweedie, 1950: 131" authorityName="Tweedie" authorityPageNumber="131" authorityYear="1950" box="[136,510,1308,1330]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="sigillata">
<emphasis id="3ECBF1529355FFD12BA49A41FE830819" box="[136,305,1308,1329]" italics="true" pageId="4" pageNumber="261">Sesarma sigillata</emphasis>
<bibRefCitation id="682E50B19355FFD12A159A41FE4C0819" author="Tweedie MWF" box="[313,510,1308,1330]" pageId="4" pageNumber="261" pagination="105 - 148" refId="ref7623" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105 - 148, figs. 1 - 4, pls. 16, 17." type="journal article" year="1950">Tweedie, 1950: 131</bibRefCitation>
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, fig. 3ae.
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<emphasis id="3ECBF1529355FFD12BA49A64FF690866" box="[136,219,1337,1358]" italics="true" pageId="4" pageNumber="261">Sesarma</emphasis>
(
<emphasis id="3ECBF1529355FFD12BC69A64FED80866" box="[234,362,1337,1358]" italics="true" pageId="4" pageNumber="261">Parasesarma</emphasis>
)
<emphasis id="3ECBF1529355FFD12A569A64FE680866" box="[378,474,1337,1358]" italics="true" pageId="4" pageNumber="261">sigillatum</emphasis>
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<treatmentCitation id="8D1E0B519355FFD12ADA9A64FD180866" author="Serene R" box="[502,682,1337,1359]" page="108" pageId="4" pageNumber="261" year="1968">
<bibRefCitation id="682E50B19355FFD12ADA9A64FD180866" author="Serene R" box="[502,682,1337,1359]" pageId="4" pageNumber="261" pagination="33 - 120" refId="ref7532" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, No. 1: 33 - 120." type="book chapter" year="1968">Serène, 1968: 108</bibRefCitation>
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.
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<treatmentCitationGroup id="2CAF0A6E9355FFD12BA49A0BFD0908A0" pageId="4" pageNumber="261">
<taxonomicName id="CBBF56C39355FFD12BA49A0BFE300842" ID-CoL="75QHG" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[136,386,1365,1387]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="carolinense">
<emphasis id="3ECBF1529355FFD12BA49A0BFE300842" box="[136,386,1365,1387]" italics="true" pageId="4" pageNumber="261">Parasesarma carolinense</emphasis>
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<treatmentCitation id="8D1E0B519355FFD12A8C9A0BFDDE0843" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[416,620,1365,1387]" page="222" pageId="4" pageNumber="261" year="2008">
<bibRefCitation id="682E50B19355FFD12A8C9A0BFDDE0843" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[416,620,1365,1387]" pageId="4" pageNumber="261" pagination="1 - 286" refId="ref7132" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 222</bibRefCitation>
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(list, part) (not
<taxonomicName id="CBBF56C39355FFD12B829A2EFD0208A0" ID-CoL="4WYVP" authority="Rathbun, 1907" authorityName="Rathbun" authorityYear="1907" box="[174,688,1394,1416]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="carolinensis" subGenus="Parasesarma">
<emphasis id="3ECBF1529355FFD12B829A2EFEB308A0" box="[174,257,1395,1416]" italics="true" pageId="4" pageNumber="261">Sesarma</emphasis>
(
<emphasis id="3ECBF1529355FFD12A3C9A2EFE2208A0" box="[272,400,1395,1416]" italics="true" pageId="4" pageNumber="261">Parasesarma</emphasis>
)
<emphasis id="3ECBF1529355FFD12A8C9A2FFDA708AF" box="[416,533,1394,1415]" italics="true" pageId="4" pageNumber="261">carolinensis</emphasis>
<bibRefCitation id="682E50B19355FFD129319A2FFD0208A0" author="Rathbun MJ" box="[541,688,1394,1416]" pageId="4" pageNumber="261" pagination="25 - 74" refId="ref7356" refString="Rathbun MJ (1907) Reports on the scientific results of the expedition to the tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding. IX. Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer &quot; Albatross, &quot; from October, 1904, to March, 1905, Lieut. - Commander L. M. Garrett, U. S. N., commanding. X: The Brachyura. Memoirs of the Museum of Comparative Zoology at Harvard College, 35 (2): 25 - 74." type="journal article" year="1907">Rathbun, 1907</bibRefCitation>
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).
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<emphasis id="3ECBF1529355FFD12BA49AEDFED108E0" bold="true" box="[136,355,1456,1480]" pageId="4" pageNumber="261">Material examined.</emphasis>
<materialsCitation id="BCD7271D9355FFD12A449AECFE370B21" collectionCode="NHM" collectorName="C. A. Gibson-Hill" location="Cocos-Keeling Islands" pageId="4" pageNumber="261" specimenCount="1" typeStatus="lectotype">
<typeStatus id="D30493E29355FFD12A449AECFE6008E1" box="[360,466,1457,1481]" pageId="4" pageNumber="261" type="lectotype">Lectotype</typeStatus>
(here designated): male (10.7 ×
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1950.4.17.4),
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<collectorName id="A14A48969355FFD12BA49AACFEF10B20" box="[136,323,1520,1545]" pageId="4" pageNumber="261">C.A. Gibson-Hill</collectorName>
, 1941
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<materialsCitation id="BCD7271D9355FFD12ABC9AADFCB10B00" collectionCode="NHM" collectorName="Others" location="Cocos-Keeling Islands" pageId="4" pageNumber="261" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="D30493E29355FFD12ABC9AADFD970B20" box="[400,549,1520,1544]" pageId="4" pageNumber="261" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCount id="1AB9E6C99355FFD129039AACFD3E0B20" box="[559,652,1520,1545]" pageId="4" pageNumber="261" type="female">1 female</specimenCount>
(10.4 ×
<quantity id="CB4780A59355FFD129C99AADFF010B01" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="261" unit="mm" value="7.5">7.5 mm</quantity>
) (
<collectionCode id="6AAEB5859355FFD12BE0994CFEBE0B01" box="[204,268,1553,1577]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="4" pageNumber="261" type="Herbarium">NHM</collectionCode>
1950.4.17.5), same data as lectotype.
<collectorName id="A14A48969355FFD12994994DFCB10B00" box="[696,771,1552,1576]" pageId="4" pageNumber="261">Others</collectorName>
</materialsCitation>
:
<specimenCount id="1AB9E6C99355FFD12BA4996CFF5B0B60" box="[136,233,1584,1609]" pageId="4" pageNumber="261" type="male">24 males</specimenCount>
(largest 11.1 ×
<quantity id="CB4780A59355FFD12ABB996CFE5C0B61" box="[407,494,1585,1609]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="4" pageNumber="261" unit="mm" value="7.8">7.8 mm</quantity>
),
<materialsCitation id="BCD7271D9355FFD1292E996CFD9E0BA0" collectionCode="ZRC" collectorName="C. A. Gibson-Hill" location="Keeling Islands" pageId="4" pageNumber="261" specimenCount="20" specimenCount-female="20" typeStatus="paralectotype">
<specimenCount id="1AB9E6C99355FFD1292E996CFDCA0B60" box="[514,632,1584,1609]" pageId="4" pageNumber="261" type="female">20 females</specimenCount>
(largest 11.0 ×
<quantity id="CB4780A59355FFD12BB3990CFF460B41" box="[159,244,1617,1641]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="4" pageNumber="261" unit="mm" value="7.8">7.8 mm</quantity>
) (1 ovigerous) (
<collectionCode id="6AAEB5859355FFD12A88990CFE680B41" box="[420,474,1617,1641]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="261">ZRC</collectionCode>
1965.8.2.260269), Cocos-
<location id="09607B9B9355FFD12BA4992DFE830BA0" LSID="urn:lsid:plazi:treatment:84169C569355FFD22BC99BE3FDCC0526:09607B9B9355FFD12BA4992DFE830BA0" box="[136,305,1648,1672]" name="Keeling Islands" pageId="4" pageNumber="261">Keeling Islands</location>
, coll.
<collectorName id="A14A48969355FFD12A5D992CFD9E0BA0" box="[369,556,1648,1673]" pageId="4" pageNumber="261">C.A. Gibson-Hill</collectorName>
</materialsCitation>
, 1941;
<materialsCitation id="BCD7271D9355FFD12956992CFE560BE1" collectingDate="2011-03-21" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; P. J. F. Davie" location="Cocos-Keeling Islands" pageId="4" pageNumber="261" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="1AB9E6C99355FFD12956992CFD650BA0" box="[634,727,1648,1673]" pageId="4" pageNumber="261" type="female">1 female</specimenCount>
(9.9 × 7.0 mm) (
<collectionCode id="6AAEB5859355FFD12A2199CCFEF40B81" box="[269,326,1681,1705]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="261">ZRC</collectionCode>
),
<location id="09607B9B9355FFD12A7799CDFDE00B80" LSID="urn:lsid:plazi:treatment:84169C569355FFD22BC99BE3FDCC0526:09607B9B9355FFD12A7799CDFDE00B80" box="[347,594,1680,1704]" name="Cocos-Keeling Islands" pageId="4" pageNumber="261">Cocos-Keeling Islands</location>
, coll.
<collectorName id="A14A48969355FFD129BB99CCFCB50B81" box="[663,775,1681,1705]" pageId="4" pageNumber="261">P.K.L. Ng</collectorName>
&amp;
<collectorName id="A14A48969355FFD12B8999ECFE9E0BE0" box="[165,300,1712,1736]" pageId="4" pageNumber="261">P.J.F. Davie</collectorName>
,
<date id="78010B809355FFD12A1599ECFE560BE1" box="[313,484,1712,1737]" pageId="4" pageNumber="261" value="2011-03-21">
<collectingDate id="6845F2689355FFD12A1599ECFE560BE1" box="[313,484,1712,1737]" pageId="4" pageNumber="261" value="2011-03-21">21 March 2011</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="1AB9E6C99355FFD12ADC99ECFD880BE0" box="[496,570,1712,1737]" pageId="4" pageNumber="261" type="male">1 male</specimenCount>
(9.1 ×
<quantity id="CB4780A59355FFD129A299EDFD570BE1" box="[654,741,1712,1737]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="261" unit="mm" value="6.7">6.7 mm</quantity>
),
<specimenCount id="1AB9E6C99355FFD129D599ECFF520BC0" pageId="4" pageNumber="261" type="juvenile">1 juvenile</specimenCount>
male,
<materialsCitation id="BCD7271D9355FFD12A06998DFE310A61" collectingDate="2011-03-24" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; P. J. F. Davie" county="Cocos-Keeling Islands" location="Meterological Station" municipality="West Island" pageId="4" pageNumber="261" specimenCount="6" specimenCount-female="6" typeStatus="paralectotype">
<specimenCount id="1AB9E6C99355FFD12A06998DFE200BC0" box="[298,402,1744,1768]" pageId="4" pageNumber="261" type="female">6 females</specimenCount>
(largest 8.6 ×
<quantity id="CB4780A59355FFD1291C998DFD360BC1" box="[560,644,1744,1769]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="4" pageNumber="261" unit="mm" value="6.3">6.3 mm</quantity>
) (
<collectionCode id="6AAEB5859355FFD129B7998CFD660BC1" box="[667,724,1745,1769]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="261">ZRC</collectionCode>
), stn CK 26, lagoon side of airport near
<location id="09607B9B9355FFD1293999ADFCB10A20" LSID="urn:lsid:plazi:treatment:84169C569355FFD22BC99BE3FDCC0526:09607B9B9355FFD1293999ADFCB10A20" box="[533,771,1776,1800]" county="Cocos-Keeling Islands" municipality="West Island" name="Meterological Station" pageId="4" pageNumber="261">Meterological Station</location>
,
<collectingMunicipality id="EC64B73A9355FFD12BA4984CFEBA0A00" box="[136,264,1808,1832]" pageId="4" pageNumber="261">West Island</collectingMunicipality>
,
<collectingCounty id="E56155CC9355FFD12A3D984DFDB10A00" box="[273,515,1808,1832]" pageId="4" pageNumber="261">Cocos-Keeling Islands</collectingCounty>
, coll.
<collectorName id="A14A48969355FFD1296C984CFD1E0A01" box="[576,684,1809,1833]" pageId="4" pageNumber="261">P.K.L. Ng</collectorName>
&amp;
<collectorName id="A14A48969355FFD129E7984CFF7B0A60" pageId="4" pageNumber="261">P.J.F. Davie</collectorName>
,
<date id="78010B809355FFD12BFB986CFE310A61" box="[215,387,1840,1865]" pageId="4" pageNumber="261" value="2011-03-24">
<collectingDate id="6845F2689355FFD12BFB986CFE310A61" box="[215,387,1840,1865]" pageId="4" pageNumber="261" value="2011-03-24">24 March 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="BCD7271D9355FFD12ABC986CFD110A81" collectingDate="2011-03-22" collectorName="P. J. F. Davie &amp; P. K. L. Ng" county="Cocos-Keeling Islands" location="Meterological Station" municipality="West Island" pageId="4" pageNumber="261" specimenCode="QM-W28342" specimenCount="2" specimenCount-male="2" typeStatus="paralectotype">
<specimenCount id="1AB9E6C99355FFD12ABC986CFE570A60" box="[400,485,1840,1865]" pageId="4" pageNumber="261" type="male">2 males</specimenCount>
(9.4 ×
<quantity id="CB4780A59355FFD12910986DFD210A60" box="[572,659,1840,1864]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="4" pageNumber="261" unit="mm" value="6.9">6.9 mm</quantity>
, 9.0 ×
<quantity id="CB4780A59355FFD129C8986DFF010A41" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="261" unit="mm" value="6.7">6.7 mm</quantity>
) (
<specimenCode id="5C19853B9355FFD12BE6980CFEED0A41" box="[202,351,1873,1897]" collectionCode="QM-W" pageId="4" pageNumber="261">QM-W28342</specimenCode>
), stn CK16, lagoon side of airport near
<location id="09607B9B9355FFD12BA4982DFEC10AA0" LSID="urn:lsid:plazi:treatment:84169C569355FFD22BC99BE3FDCC0526:09607B9B9355FFD12BA4982DFEC10AA0" box="[136,371,1904,1928]" county="Cocos-Keeling Islands" municipality="West Island" name="Meterological Station" pageId="4" pageNumber="261">Meterological Station</location>
,
<collectingMunicipality id="EC64B73A9355FFD12A53982CFDB00AA0" box="[383,514,1904,1928]" pageId="4" pageNumber="261">West Island</collectingMunicipality>
,
<collectingCounty id="E56155CC9355FFD12921982DFCB10AA0" box="[525,771,1904,1928]" pageId="4" pageNumber="261">Cocos-Keeling Islands</collectingCounty>
, coll.
<collectorName id="A14A48969355FFD12BEE98CCFEFB0A80" box="[194,329,1936,1960]" pageId="4" pageNumber="261">P.J.F. Davie</collectorName>
&amp;
<collectorName id="A14A48969355FFD12A5E98CCFE540A81" box="[370,486,1937,1961]" pageId="4" pageNumber="261">P.K.L. Ng</collectorName>
,
<date id="78010B809355FFD12AD998CCFD110A81" box="[501,675,1936,1961]" pageId="4" pageNumber="261" value="2011-03-22">
<collectingDate id="6845F2689355FFD12AD998CCFD110A81" box="[501,675,1936,1961]" pageId="4" pageNumber="261" value="2011-03-22">22 March 2011</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="1AB9E6C99355FFD1299E98CCFCBA0A80" box="[690,776,1936,1961]" pageId="4" pageNumber="261" type="male">3 males</specimenCount>
(largest 7.8 ×
<quantity id="CB4780A59355FFD12A0F98EDFEC80AE1" box="[291,378,1968,1993]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="4" pageNumber="261" unit="mm" value="5.9">5.9 mm</quantity>
),
<materialsCitation id="BCD7271D9355FFD12AA398EDFD810AC1" collectingDate="2011-03-22" collectorName="P. J. F. Davie &amp; P. K. L. Ng" county="Cocos-Keeling Islands" location="Meterological Station" municipality="West Island" pageId="4" pageNumber="261" specimenCode="QM-W28343" specimenCount="16" specimenCount-female="16" typeStatus="paralectotype">
<specimenCount id="1AB9E6C99355FFD12AA398EDFDB40AE0" box="[399,518,1968,1992]" pageId="4" pageNumber="261" type="female">16 females</specimenCount>
(largest 8.7 ×
<quantity id="CB4780A59355FFD1298698EDFD4D0AE1" box="[682,767,1968,1993]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="4" pageNumber="261" unit="mm" value="6.4">6.4 mm</quantity>
) (
<specimenCode id="5C19853B9355FFD12BA3988CFE960AC1" box="[143,292,2001,2025]" collectionCode="QM-W" pageId="4" pageNumber="261">QM-W28343</specimenCode>
), same data QM-W28342
</materialsCitation>
;
<materialsCitation id="BCD7271D9355FFD12910988CFA0C0DFD" collectingDate="2011-03-24" collectorName="P. J. F. Davie &amp; P. K. L. Ng" county="Cocos-Keeling Islands" location="Meterological Station" municipality="West Island" pageId="4" pageNumber="261" specimenCode="QM-W28344" specimenCount="2" specimenCount-male="2" typeStatus="paralectotype">
<specimenCount id="1AB9E6C99355FFD12910988CFD3F0AC0" box="[572,653,2000,2025]" pageId="4" pageNumber="261" type="male">2 males</specimenCount>
(largest 9.2 ×
<quantity id="CB4780A59355FFD12BB298ADFF430521" box="[158,241,2032,2057]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="4" pageNumber="261" unit="mm" value="6.9">6.9 mm</quantity>
) (
<specimenCode id="5C19853B9355FFD12A2A98ACFE2E0521" box="[262,412,2033,2057]" collectionCode="QM-W" pageId="4" pageNumber="261">QM-W28344</specimenCode>
), stn CK26, lagoon side of airport near
<location id="09607B9B9355FFD12BEE974DFE0B0500" LSID="urn:lsid:plazi:treatment:84169C569355FFD22BC99BE3FDCC0526:09607B9B9355FFD12BEE974DFE0B0500" box="[194,441,2064,2088]" county="Cocos-Keeling Islands" municipality="West Island" name="Meterological Station" pageId="4" pageNumber="261">Meterological Station</location>
,
<collectingMunicipality id="EC64B73A9355FFD12AE4974CFDE00500" box="[456,594,2064,2088]" pageId="4" pageNumber="261">West Island</collectingMunicipality>
,
<collectingCounty id="E56155CC9355FFD1294D974DFC200DFD" box="[609,914,189,2088]" pageId="4" pageNumber="261">Cocos-Keeling Islands</collectingCounty>
, coll.
<collectorName id="A14A48969355FFD128F79FE3FBD00DFD" box="[987,1122,189,213]" pageId="4" pageNumber="261">P.J.F. Davie</collectorName>
&amp;
<collectorName id="A14A48969355FFD12FA79FE3FAB20DFE" box="[1163,1280,190,214]" pageId="4" pageNumber="261">P.K.L. Ng</collectorName>
,
<date id="78010B809355FFD12E239FE0FA0C0DFD" box="[1295,1470,189,213]" pageId="4" pageNumber="261" value="2011-03-24">
<collectingDate id="6845F2689355FFD12E239FE0FA0C0DFD" box="[1295,1470,189,213]" pageId="4" pageNumber="261" value="2011-03-24">24 March 2011</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="1AB9E6C99355FFD1286F9F80FC280DDD" box="[835,922,221,245]" pageId="4" pageNumber="261" type="male">2 males</specimenCount>
(largest 10.5 ×
<quantity id="CB4780A59355FFD12F7E9F80FB1E0DDD" box="[1106,1196,221,245]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="4" pageNumber="261" unit="mm" value="7.9">7.9 mm</quantity>
),
<materialsCitation id="BCD7271D9355FFD12FEE9F80FAFD0C7D" collectingDate="2011-03-20" collectorName="P. J. F. Davie &amp; P. K. L. Ng" county="Cocos-Keeling Islands" latitude="-12.19315" location="Tanjong Puji" longLatPrecision="1" longitude="96.85915" municipality="West Island" pageId="4" pageNumber="261" specimenCode="QM-W28345" specimenCount="4" specimenCount-female="4" typeStatus="paralectotype">
<specimenCount id="1AB9E6C99355FFD12FEE9F80FA9C0DDD" box="[1218,1326,221,245]" pageId="4" pageNumber="261" type="female">4 females</specimenCount>
(largest 11.4 ×
<quantity id="CB4780A59355FFD128719FA0FC070C3D" box="[861,949,253,277]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="261" unit="mm" value="8.1">8.1 mm</quantity>
) (
<specimenCode id="5C19853B9355FFD128E29FA0FBD40C3D" box="[974,1126,253,277]" collectionCode="QM-W" pageId="4" pageNumber="261">QM-W28345</specimenCode>
), stn CK3,
<location id="09607B9B9355FFD12FC59FA0FACF0C3D" LSID="urn:lsid:plazi:treatment:84169C569355FFD22BC99BE3FDCC0526:09607B9B9355FFD12FC59FA0FACF0C3D" box="[1257,1405,253,277]" county="Cocos-Keeling Islands" latitude="-12.19315" longLatPrecision="1" longitude="96.85915" municipality="West Island" name="Tanjong Puji" pageId="4" pageNumber="261">Tanjong Puji</location>
,
<collectingMunicipality id="EC64B73A9355FFD12EA79FA3FC350C1D" pageId="4" pageNumber="261">West Island</collectingMunicipality>
,
<collectingCounty id="E56155CC9355FFD128BA9E40FB3C0C1D" box="[918,1166,285,309]" pageId="4" pageNumber="261">Cocos-Keeling Islands</collectingCounty>
,
<geoCoordinate id="698B4B879355FFD12FB09E40FA970C1D" box="[1180,1317,285,309]" degrees="12" direction="south" minutes="11.589" orientation="latitude" pageId="4" pageNumber="261" precision="1" value="-12.19315">12°11.589S</geoCoordinate>
,
<geoCoordinate id="698B4B879355FFD12E189E40FA0D0C1D" box="[1332,1471,285,309]" degrees="96" direction="east" minutes="51.549" orientation="longitude" pageId="4" pageNumber="261" precision="1" value="96.85915">96°51.549E</geoCoordinate>
, coll.
<collectorName id="A14A48969355FFD128579E63FC4D0C7D" box="[891,1023,317,341]" pageId="4" pageNumber="261">P.J.F. Davie</collectorName>
&amp;
<collectorName id="A14A48969355FFD12F099E63FB250C7E" box="[1061,1175,318,342]" pageId="4" pageNumber="261">P.K.L. Ng</collectorName>
,
<date id="78010B809355FFD12F889E60FAFD0C7D" box="[1188,1359,317,341]" pageId="4" pageNumber="261" value="2011-03-20">
<collectingDate id="6845F2689355FFD12F889E60FAFD0C7D" box="[1188,1359,317,341]" pageId="4" pageNumber="261" value="2011-03-20">20 March 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="44A57ECB9355FFD1286F9E20FB2208DD" pageId="4" pageNumber="261" type="diagnosis">
<paragraph id="0C002D409355FFD1286F9E20FA13091D" blockId="4.[834,1475,381,1077]" pageId="4" pageNumber="261">
<emphasis id="3ECBF1529355FFD1286F9E20FC090CBD" bold="true" box="[835,955,381,405]" pageId="4" pageNumber="261">Diagnosis.</emphasis>
Carapace 1.351.44 times broader than long; frontal margin bilobed, separated by broad, very shallow concavity (
<figureCitation id="948431C59355FFD128EF9EE0FBF30CFD" box="[963,1089,445,469]" captionStart="Fig" captionStartId="5.[113,148,1948,1970]" captionTargetBox="[113,1451,248,1933]" captionTargetId="figure-92@5.[113,1451,248,1933]" captionTargetPageId="5" captionText="Fig. 3. Parasesarma sigillatum (Tweedie, 1950). AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="4" pageNumber="261">Fig. 3AC</figureCitation>
); upper surface of cheliped palm with 2 transverse pectinate crests, primary (distalmost) crest with row of 1518 tall pectinate teeth, tip of each tooth gently curved, secondary crest with 9 or 10 relatively lower pectinate teeth, followed by patch of 1215 scattered low, more rounded pectinate tubercles on median part of palm (
<figureCitation id="948431C59355FFD128679D20FC1C0FBD" box="[843,942,637,661]" captionStart="Fig" captionStartId="5.[113,148,1948,1970]" captionTargetBox="[113,1451,248,1933]" captionTargetId="figure-92@5.[113,1451,248,1933]" captionTargetPageId="5" captionText="Fig. 3. Parasesarma sigillatum (Tweedie, 1950). AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="4" pageNumber="261">Figs. 3H</figureCitation>
;
<figureCitation id="948431C59355FFD128909D20FC520FBD" box="[956,992,637,661]" captionStart="Fig" captionStartId="9.[113,148,778,800]" captionTargetBox="[112,1450,192,765]" captionTargetId="figure-672@9.[100,1463,180,772]" captionTargetPageId="9" captionText="Fig. 7. Details of pectinate crests on chela. A, Parasesarma sigillatum (Tweedie, 1950), lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island; B, Parasesarma corallicum n. sp., holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. Arrow points to direction of dactylar finger. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5355321" httpUri="https://zenodo.org/record/5355321/files/figure.png" pageId="4" pageNumber="261">7A</figureCitation>
); inner surface of palm with low, obliquely transverse, submedian granulated ridge (
<figureCitation id="948431C59355FFD12E389DC0FAD70F9D" box="[1300,1381,669,693]" captionStart="Fig" captionStartId="5.[113,148,1948,1970]" captionTargetBox="[113,1451,248,1933]" captionTargetId="figure-92@5.[113,1451,248,1933]" captionTargetPageId="5" captionText="Fig. 3. Parasesarma sigillatum (Tweedie, 1950). AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="4" pageNumber="261">Fig. 3I</figureCitation>
); dorsal surface of dactylus with 1618 triangular tubercles, each dactylar tubercle with distinct transverse ridge (
<figureCitation id="948431C59355FFD12E4A9D80FC210E3D" captionStart="Fig" captionStartId="5.[113,148,1948,1970]" captionTargetBox="[113,1451,248,1933]" captionTargetId="figure-92@5.[113,1451,248,1933]" captionTargetPageId="5" captionText="Fig. 3. Parasesarma sigillatum (Tweedie, 1950). AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="4" pageNumber="261">Fig. 3E, F, H, I</figureCitation>
); inner distal margin of inner lamelliform process of merus lined with sharp triangular teeth (
<figureCitation id="948431C59355FFD12E099C40FA060E1D" box="[1317,1460,797,821]" captionStart="Fig" captionStartId="5.[113,148,1948,1970]" captionTargetBox="[113,1451,248,1933]" captionTargetId="figure-92@5.[113,1451,248,1933]" captionTargetPageId="5" captionText="Fig. 3. Parasesarma sigillatum (Tweedie, 1950). AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="4" pageNumber="261">Fig. 3E, F, I</figureCitation>
); ambulatory legs short, stout; merus of third leg 2.0 times as long as wide; upper margin of merus with subdistal spine; propodus of third leg 2.8 times as long as wide; dactylus of third and fourth legs 0.8 and 1.0 times length of propodus, respectively (
<figureCitation id="948431C59355FFD128FE9CE0FB800EFD" box="[978,1074,957,981]" captionStart="Fig" captionStartId="5.[113,148,1948,1970]" captionTargetBox="[113,1451,248,1933]" captionTargetId="figure-92@5.[113,1451,248,1933]" captionTargetPageId="5" captionText="Fig. 3. Parasesarma sigillatum (Tweedie, 1950). AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="4" pageNumber="261">Figs. 3A</figureCitation>
;
<figureCitation id="948431C59355FFD12F109CE0FB330EFD" box="[1084,1153,957,981]" captionStart="Fig" captionStartId="7.[113,148,1491,1513]" captionTargetBox="[113,1451,189,1476]" captionTargetId="figure-93@7.[113,1451,189,1476]" captionTargetPageId="7" captionText="Fig. 5. Left ambulatory legs. AC, Parasesarma carolinense (Rathbun, 1914), holotype male (9.2 × 7.0 mm) (USNM 32861), Caroline Islands; DG, Parasesarma sigillatum (Tweedie, 1950), male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; HK, Parasesarma corallicum n. sp., holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. A, D, H, first ambulatory leg; B, E, I, second ambulatory leg; C, F, J, third ambulatory leg; G, K, fourth ambulatory leg. Legs of all species to scale." figureDoi="http://doi.org/10.5281/zenodo.5355317" httpUri="https://zenodo.org/record/5355317/files/figure.png" pageId="4" pageNumber="261">5DG</figureCitation>
); male telson semicircular, ca. 1.2 times length of pleonal somite 6 (
<figureCitation id="948431C59355FFD12FE29C80FA910EDD" box="[1230,1315,989,1013]" captionStart="Fig" captionStartId="8.[135,170,1920,1942]" captionTargetBox="[189,1437,184,1908]" captionTargetId="figure-8@8.[189,1437,184,1911]" captionTargetPageId="8" captionText="Fig. 6. AF, Parasesarma carolinense (Rathbun, 1914), holotype male (9.2 × 7.0 mm) (USNM 32861), Caroline Islands; GM, Parasesarma sigillatum (Tweedie, 1950), male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; NT, Parasesarma corallicum n. sp., holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. A, G, N, left third maxilliped; B, H, male pleon; O, male pleon (somites 1 and 3 broken); C, P, ventral view of left G1; D, Q, dorsal view of left G1; E, ventral view of right G1; F, distal chitinous part of right G1 (ventral view); I, ventral view of right G1; J, dorsal view of right G1; K, distal chitinous part of right G1 (ventral view); L, distal chitinous part of right G1 (dorsal view); M, right G2; R, distal chitinous part of left G1 (ventral view); S, distal chitinous part of left G1 (dorsal view); T, left G2. Setae for all structures denuded. Scale bars = 0.5 mm [AE, GJ, MQ, T]; 0.1 mm [F, K, L, R, S]." figureDoi="http://doi.org/10.5281/zenodo.5355319" httpUri="https://zenodo.org/record/5355319/files/figure.png" pageId="4" pageNumber="261">Fig. 6H</figureCitation>
); G1 relatively slender, straight; apical process bent at 90° from the vertical, chitinous distal part long, aperture terminal (
<figureCitation id="948431C59355FFD12E059B40FA27091D" box="[1321,1429,1053,1077]" captionStart="Fig" captionStartId="8.[135,170,1920,1942]" captionTargetBox="[189,1437,184,1908]" captionTargetId="figure-8@8.[189,1437,184,1911]" captionTargetPageId="8" captionText="Fig. 6. AF, Parasesarma carolinense (Rathbun, 1914), holotype male (9.2 × 7.0 mm) (USNM 32861), Caroline Islands; GM, Parasesarma sigillatum (Tweedie, 1950), male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; NT, Parasesarma corallicum n. sp., holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. A, G, N, left third maxilliped; B, H, male pleon; O, male pleon (somites 1 and 3 broken); C, P, ventral view of left G1; D, Q, dorsal view of left G1; E, ventral view of right G1; F, distal chitinous part of right G1 (ventral view); I, ventral view of right G1; J, dorsal view of right G1; K, distal chitinous part of right G1 (ventral view); L, distal chitinous part of right G1 (dorsal view); M, right G2; R, distal chitinous part of left G1 (ventral view); S, distal chitinous part of left G1 (dorsal view); T, left G2. Setae for all structures denuded. Scale bars = 0.5 mm [AE, GJ, MQ, T]; 0.1 mm [F, K, L, R, S]." figureDoi="http://doi.org/10.5281/zenodo.5355319" httpUri="https://zenodo.org/record/5355319/files/figure.png" pageId="4" pageNumber="261">Fig. 6IL</figureCitation>
).
</paragraph>
<paragraph id="0C002D409355FFD1286F9B00FBBC09DD" blockId="4.[835,1474,1117,1269]" pageId="4" pageNumber="261">
<emphasis id="3ECBF1529355FFD1286F9B00FC51095D" bold="true" box="[835,995,1117,1141]" pageId="4" pageNumber="261">Colour in life.</emphasis>
Dorsal surfaces of carapace and ambulatory legs dirty white with large and small uneven patches of dark brown to grey (
<figureCitation id="948431C59355FFD128DC9BC0FBDB099D" box="[1008,1129,1181,1205]" captionStart="Fig" captionStartId="1.[113,148,1887,1909]" captionTargetBox="[113,1451,308,1872]" captionTargetId="figure-15@1.[113,1451,308,1872]" captionTargetPageId="1" captionText="Fig. 1. Colour in life. AD, Parasesarma sigillatum, Cocos-Keeling Islands; EH, Parasesarma corallicum n. sp., Taiwan. A, male, not preserved; B, female (8.7 × 6.4 mm) (QM-W28343); C, D, male (9.4 × 6.9 mm) (QM-W28342); E, F, male, Hsiangchiaowan, Taiwan (10.3 × 7.6 mm) (NMMBCD 4698); G, Holotype, same data (9.5 × 7.0 mm) (NMMBCD 4697); H, Wanlitung, Taiwan (8.0 × 5.7 mm) (NMNS-7548-002)." figureDoi="http://doi.org/10.5281/zenodo.5355309" httpUri="https://zenodo.org/record/5355309/files/figure.png" pageId="4" pageNumber="261">Fig. 1AC</figureCitation>
). Chelae light orange, although ventral half darker, with dorsal parts sometimes appearing paler (
<figureCitation id="948431C59355FFD128A59B80FBB009DD" box="[905,1026,1245,1269]" captionStart="Fig" captionStartId="1.[113,148,1887,1909]" captionTargetBox="[113,1451,308,1872]" captionTargetId="figure-15@1.[113,1451,308,1872]" captionTargetPageId="1" captionText="Fig. 1. Colour in life. AD, Parasesarma sigillatum, Cocos-Keeling Islands; EH, Parasesarma corallicum n. sp., Taiwan. A, male, not preserved; B, female (8.7 × 6.4 mm) (QM-W28343); C, D, male (9.4 × 6.9 mm) (QM-W28342); E, F, male, Hsiangchiaowan, Taiwan (10.3 × 7.6 mm) (NMMBCD 4698); G, Holotype, same data (9.5 × 7.0 mm) (NMMBCD 4697); H, Wanlitung, Taiwan (8.0 × 5.7 mm) (NMNS-7548-002)." figureDoi="http://doi.org/10.5281/zenodo.5355309" httpUri="https://zenodo.org/record/5355309/files/figure.png" pageId="4" pageNumber="261">Fig. 1AD</figureCitation>
).
</paragraph>
<paragraph id="0C002D409355FFD1286F9A40FB2208DD" blockId="4.[835,1475,1309,1525]" pageId="4" pageNumber="261">
<emphasis id="3ECBF1529355FFD1286F9A40FC11081D" bold="true" box="[835,931,1309,1333]" pageId="4" pageNumber="261">Habitat.</emphasis>
<taxonomicName id="CBBF56C39355FFD128819A43FB01081D" baseAuthorityName="Tweedie" baseAuthorityYear="1950" box="[941,1203,1309,1333]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="sigillatum">
<emphasis id="3ECBF1529355FFD128819A43FB01081D" box="[941,1203,1309,1333]" italics="true" pageId="4" pageNumber="261">Parasesarma sigillatum</emphasis>
</taxonomicName>
inhabits supralittoral or upper intertidal zones. It prefers a sandy substrate with some mud, and with grass growing nearby. The crabs were found hiding under flotsam, pieces of wood and small rocks. An allied species,
<taxonomicName id="CBBF56C39355FFD128C99AC3FBC1089D" authorityName="De Man" authorityYear="1895" box="[997,1139,1438,1461]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="3ECBF1529355FFD128C99AC3FBC1089D" box="[997,1139,1438,1461]" italics="true" pageId="4" pageNumber="261">Parasesarma</emphasis>
</taxonomicName>
cf.
<emphasis id="3ECBF1529355FFD12F8D9AC0FB69089D" box="[1185,1243,1437,1461]" italics="true" pageId="4" pageNumber="261">lenzii</emphasis>
(De
<bibRefCitation id="682E50B19355FFD12E399AC3FA3F089D" author="Man JG De" box="[1301,1421,1437,1461]" pageId="4" pageNumber="261" pagination="485 - 609" refId="ref7027" refString="Man JG De (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8: 485 - 609, pls. 12 - 14." type="journal article" year="1895">Man, 1895</bibRefCitation>
) can sometimes be found with it, although this species prefers muddier and wetter substrates.
</paragraph>
</subSubSection>
<subSubSection id="44A57ECB9355FFD2286F9940FDCC0526" lastPageId="7" lastPageNumber="264" pageId="4" pageNumber="261" type="discussion">
<paragraph id="0C002D409355FFD2286F9940FF4E0A26" blockId="4.[835,1475,1565,2101]" lastBlockId="7.[113,753,1654,1806]" lastPageId="7" lastPageNumber="264" pageId="4" pageNumber="261">
<emphasis id="3ECBF1529355FFD1286F9940FC010B1D" bold="true" box="[835,947,1565,1589]" pageId="4" pageNumber="261">Remarks.</emphasis>
<bibRefCitation id="682E50B19355FFD128919940FB170B1D" author="Tweedie MWF" box="[957,1189,1565,1589]" pageId="4" pageNumber="261" pagination="105 - 148" refId="ref7623" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105 - 148, figs. 1 - 4, pls. 16, 17." type="journal article" year="1950">Tweedie (1950: 131)</bibRefCitation>
stated that
<specimenCount id="1AB9E6C99355FFD12E009943FA230B1D" box="[1324,1425,1565,1589]" pageId="4" pageNumber="261" type="male">one male</specimenCount>
and
<specimenCount id="1AB9E6C99355FFD1286F9963FC700B7D" box="[835,962,1597,1621]" pageId="4" pageNumber="261" type="female">one female</specimenCount>
were “cotypes”, but noted that he also had 43 additional specimens. This means that only the
<specimenCount id="1AB9E6C99355FFD12EB59903FC010BBD" pageId="4" pageNumber="261" type="generic">two specimens</specimenCount>
in the NHM (labelled as cotypes) are
<typeStatus id="D30493E29355FFD12E739923FA0D0BBE" box="[1375,1471,1662,1686]" pageId="4" pageNumber="261" type="syntype">syntypes</typeStatus>
, with the remaining material in ZRC not types. The NHM male is here designated the
<typeStatus id="D30493E29355FFD12F5699E0FB6E0BFD" box="[1146,1244,1725,1749]" pageId="4" pageNumber="261" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="CBBF56C39355FFD12E2999E3FC560BDD" authority="Tweedie, 1950" authorityName="Tweedie" authorityYear="1950" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="sigillata">
<emphasis id="3ECBF1529355FFD12E2999E3FA700BFD" box="[1285,1474,1725,1749]" italics="true" pageId="4" pageNumber="261">Sesarma sigillata</emphasis>
<bibRefCitation id="682E50B19355FFD1286F9980FC560BDD" author="Tweedie MWF" box="[835,996,1757,1781]" pageId="4" pageNumber="261" pagination="105 - 148" refId="ref7623" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105 - 148, figs. 1 - 4, pls. 16, 17." type="journal article" year="1950">Tweedie, 1950</bibRefCitation>
</taxonomicName>
. There are, however,
<specimenCount id="1AB9E6C99355FFD12FE99980FAE60BDD" box="[1221,1364,1757,1781]" pageId="4" pageNumber="261" type="generic">44 specimens</specimenCount>
in a bottle in the ZRC (
<specimenCount id="1AB9E6C99355FFD128E199A0FB800A3D" box="[973,1074,1789,1813]" pageId="4" pageNumber="261" type="male">24 males</specimenCount>
,
<specimenCount id="1AB9E6C99355FFD12F1299A0FB0A0A3D" box="[1086,1208,1789,1813]" pageId="4" pageNumber="261" type="female">20 females</specimenCount>
), not 43, suggesting that Tweedie miscounted. The two NHM specimens are in very poor condition: all the chelipeds and ambulatory legs are detached; and their carapaces are very delicate, with that of the female already medially broken. However, they agree with the ZRC specimens in all aspects, including the structure of the G1. Both G1s of the
<typeStatus id="D30493E29355FFD12F7A98E0FB0A0AFD" box="[1110,1208,1981,2005]" pageId="4" pageNumber="261" type="lectotype">lectotype</typeStatus>
male are intact and still on the specimen, and the G1 figured by
<bibRefCitation id="682E50B19355FFD12E3F9880FA0C0ADD" author="Tweedie MWF" box="[1299,1470,2013,2037]" pageId="4" pageNumber="261" pagination="105 - 148" refId="ref7623" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105 - 148, figs. 1 - 4, pls. 16, 17." type="journal article" year="1950">Tweedie (1950</bibRefCitation>
: fig. 3d) must have therefore come from one of the males in the ZRC. It was probably from the male in ZRC which has its own catalogue number (ZRC 1965.8.2.260) and a note in Tweedies handwriting that it was photographed; this specimen no longer has an pleon or gonopods suggesting they may have been removed for figuring and not returned to the bottle.
</paragraph>
<caption id="58C07DC89354FFD02B5D98C1FCFD0520" ID-DOI="http://doi.org/10.5281/zenodo.5355313" ID-Zenodo-Dep="5355313" httpUri="https://zenodo.org/record/5355313/files/figure.png" pageId="5" pageNumber="262" startId="5.[113,148,1948,1970]" targetBox="[113,1451,248,1933]" targetPageId="5">
<paragraph id="0C002D409354FFD02B5D98C1FCFD0520" blockId="5.[113,1452,1948,2056]" pageId="5" pageNumber="262">
Fig. 3.
<taxonomicName id="CBBF56C39354FFD02B9098C1FDE50A9A" authority="(Tweedie, 1950)" baseAuthorityName="Tweedie" baseAuthorityYear="1950" box="[188,599,1948,1970]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="sigillatum">
<emphasis id="3ECBF1529354FFD02B9098C1FE1A0A99" box="[188,424,1948,1969]" italics="true" pageId="5" pageNumber="262">Parasesarma sigillatum</emphasis>
(
<bibRefCitation id="682E50B19354FFD02A9498C1FDE20A9A" author="Tweedie MWF" box="[440,592,1948,1970]" pageId="5" pageNumber="262" pagination="105 - 148" refId="ref7623" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105 - 148, figs. 1 - 4, pls. 16, 17." type="journal article" year="1950">Tweedie, 1950</bibRefCitation>
)
</taxonomicName>
. AH, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; I, lectotype male (10.7 × 7.6 mm) (NHM 1950.4.17.4), Cocos-Keeling Island. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and pleon; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela.
</paragraph>
</caption>
<caption id="58C07DC89357FFD32BA498FBFB450ADE" ID-DOI="http://doi.org/10.5281/zenodo.5355315" ID-Zenodo-Dep="5355315" httpUri="https://zenodo.org/record/5355315/files/figure.png" pageId="6" pageNumber="263" startId="6.[136,171,1958,1980]" targetBox="[136,1474,258,1943]" targetPageId="6">
<paragraph id="0C002D409357FFD32BA498FBFB450ADE" blockId="6.[136,1475,1958,2038]" pageId="6" pageNumber="263">
Fig. 4.
<taxonomicName id="CBBF56C39357FFD32BFD98FAFE730A93" authority="Ng &amp; Davie &amp; Li, 2016" authorityName="Ng &amp; Davie &amp; Li" authorityYear="2016" box="[209,449,1958,1980]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="263" phylum="Arthropoda" rank="species" species="corallicum" status="sp. nov.">
<emphasis id="3ECBF1529357FFD32BFD98FAFE730A93" box="[209,449,1958,1980]" italics="true" pageId="6" pageNumber="263">Parasesarma corallicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25F84C299357FFD32AE598FAFDB20A94" box="[457,512,1959,1980]" pageId="6" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
, holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, anterior thoracic sternum and sternopleonal cavity; E, dorsal view of left cheliped; F, dorsal view of right cheliped; G, outer view of right chela; H, dactylar tubercles of right chela; I, inner view of left chela.
</paragraph>
</caption>
<caption id="58C07DC89356FFD22B5D9A8EFC460B17" ID-DOI="http://doi.org/10.5281/zenodo.5355317" ID-Zenodo-Dep="5355317" httpUri="https://zenodo.org/record/5355317/files/figure.png" pageId="7" pageNumber="264" startId="7.[113,148,1491,1513]" targetBox="[113,1451,189,1476]" targetPageId="7">
<paragraph id="0C002D409356FFD22B5D9A8EFC460B17" blockId="7.[113,1452,1491,1599]" pageId="7" pageNumber="264">
Fig. 5. Left ambulatory legs. AC,
<taxonomicName id="CBBF56C39356FFD22AFC9A89FCC008C1" authority="(Rathbun, 1914)" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[464,882,1491,1513]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" species="carolinense">
<emphasis id="3ECBF1529356FFD22AFC9A89FD7A08C0" box="[464,712,1491,1513]" italics="true" pageId="7" pageNumber="264">Parasesarma carolinense</emphasis>
(Rathbun, 1914)
</taxonomicName>
, holotype male (9.2 × 7.0 mm) (USNM 32861), Caroline Islands; DG,
<taxonomicName id="CBBF56C39356FFD22BD59AADFD330B2E" authority="(Tweedie, 1950)" baseAuthorityName="Tweedie" baseAuthorityYear="1950" box="[249,641,1520,1542]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" species="sigillatum">
<emphasis id="3ECBF1529356FFD22BD59AADFE6F0B2D" box="[249,477,1520,1541]" italics="true" pageId="7" pageNumber="264">Parasesarma sigillatum</emphasis>
(
<bibRefCitation id="682E50B19356FFD22AC49AADFDC80B2E" author="Tweedie MWF" box="[488,634,1520,1542]" pageId="7" pageNumber="264" pagination="105 - 148" refId="ref7623" refString="Tweedie MWF (1950) The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105 - 148, figs. 1 - 4, pls. 16, 17." type="journal article" year="1950">Tweedie, 1950</bibRefCitation>
)
</taxonomicName>
, male (9.5 × 6.8 mm) (ZRC 1965.8.2.269), Cocos-Keeling Island; HK,
<taxonomicName id="CBBF56C39356FFD22E009AADFF6B0B0A" authority="Ng &amp; Davie &amp; Li, 2016" authorityName="Ng &amp; Davie &amp; Li" authorityYear="2016" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" species="corallicum" status="sp. nov.">
<emphasis id="3ECBF1529356FFD22E009AADFF6B0B0A" italics="true" pageId="7" pageNumber="264">Parasesarma corallicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25F84C299356FFD22BCD9950FEAB0B0A" box="[225,281,1549,1570]" pageId="7" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
, holotype male (9.5 × 7.0 mm) (NMMBCD 4697), Taiwan. A, D, H, first ambulatory leg; B, E, I, second ambulatory leg; C, F, J, third ambulatory leg; G, K, fourth ambulatory leg. Legs of all species to scale.
</paragraph>
</caption>
<paragraph id="0C002D409356FFD22B5F986BFDEF0A86" blockId="7.[113,755,1846,1966]" pageId="7" pageNumber="264">
There is a label with the NHM material that states that this species is conspecific with “
<taxonomicName id="CBBF56C39356FFD22ABD980AFDDB0A46" authorityName="Rathbun" authorityYear="1907" box="[401,617,1878,1902]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" species="carolinense">
<emphasis id="3ECBF1529356FFD22ABD980AFDDB0A46" box="[401,617,1878,1902]" italics="true" pageId="7" pageNumber="264">Sesarma carolinense</emphasis>
</taxonomicName>
”, suggesting that Serène may have examined the material before he synonymized
<taxonomicName id="CBBF56C39356FFD22A2498CAFE130A86" authorityName="Rathbun" authorityYear="1907" box="[264,417,1942,1966]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" species="carolinense">
<emphasis id="3ECBF1529356FFD22A2498CAFE130A86" box="[264,417,1942,1966]" italics="true" pageId="7" pageNumber="264">S. carolinense</emphasis>
</taxonomicName>
with
<taxonomicName id="CBBF56C39356FFD22AC898CAFDE40A86" authorityName="Tweedie" authorityYear="1950" box="[484,598,1942,1966]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" species="sigillata">
<emphasis id="3ECBF1529356FFD22AC898CAFDE40A86" box="[484,598,1942,1966]" italics="true" pageId="7" pageNumber="264">S. sigillata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0C002D409356FFD22B5D988BFDCC0526" blockId="7.[113,752,2006,2062]" pageId="7" pageNumber="264">
The differences between
<taxonomicName id="CBBF56C39356FFD22A8C988AFDFB0AC6" authorityName="Rathbun" authorityYear="1907" box="[416,585,2006,2030]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" sensu="stricto" species="carolinense">
<emphasis id="3ECBF1529356FFD22A8C988AFDFB0AC6" box="[416,585,2006,2030]" italics="true" pageId="7" pageNumber="264">P. carolinense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25F84C299356FFD2297A988AFD2A0AC6" box="[598,664,2007,2030]" pageId="7" pageNumber="264" sensu="stricto">s. str.</taxonomicNameLabel>
and
<taxonomicName id="CBBF56C39356FFD229F4988AFF570526" authority="Ng &amp; Davie &amp; Li, 2016" authorityName="Ng &amp; Davie &amp; Li" authorityYear="2016" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="264" phylum="Arthropoda" rank="species" species="corallicum" status="sp. nov.">
<emphasis id="3ECBF1529356FFD229F4988AFF570526" italics="true" pageId="7" pageNumber="264">P. corallicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25F84C299356FFD22BC298AAFE9B0526" box="[238,297,2039,2062]" pageId="7" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
are outlined in the Discussion.
</paragraph>
</subSubSection>
</treatment>
</document>