326 lines
44 KiB
XML
326 lines
44 KiB
XML
<document id="376235C87BF993709763CECAC715F4F4" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-Zenodo-Dep="13742850" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan & Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFEF891D9D9AFDB1FE0CFC55" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis kethai Karimbumkara & Priyadarsanan, 2024, sp. nov." docType="treatment" docUuid="C3F62D8F-89D9-4034-A209-382401D8B861" docUuidSource="ZooBank" docVersion="4" lastPageNumber="28" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="26" updateTime="1726239157283" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="55D8FCCA55F33F976624426D20C154B4">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="2359DDBF9FC05D8F601985D3BCD49AD4">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:affiliation id="99F0879C3500F072224D5209E8C2B810">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:nameIdentifier id="39032423DE2A11BAC3DD8583774C802A" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
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<mods:namePart id="46C4D8C078DB1193C4929E3811BC8991">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="B98ACFB74F8B315B82FED6F636F948AE">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:nameIdentifier id="16A9FD58CD000668ABE81C3C018766DA" type="email">priyan@atree.org</mods:nameIdentifier>
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<mods:title id="624F3BDC065ACAD52480B44D43AC48C5">European Journal of Taxonomy</mods:title>
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<mods:part id="B731790AF88ECD76B4BB019F09B8447A">
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<mods:date id="F4D99C0143AA26118445155AEB9F7F88">2024</mods:date>
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<mods:number id="532A66D84DCFE13EE63D41450C3DFC66">2024-09-10</mods:number>
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<treatment id="557187D2FFEF891D9D9AFDB1FE0CFC55" ID-DOI="http://doi.org/10.5281/zenodo.13742909" ID-Zenodo-Dep="13742909" LSID="urn:lsid:zoobank.org:act:C3F62D8F-89D9-4034-A209-382401D8B861" httpUri="http://treatment.plazi.org/id/557187D2FFEF891D9D9AFDB1FE0CFC55" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
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<subSubSection id="95C2654FFFEF891F9D9AFDB1FC60FD91" box="[667,921,512,539]" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="DD6736C4FFEF891F9D9AFDB1FC60FD91" blockId="25.[373,1214,512,610]" box="[667,921,512,539]" pageId="25" pageNumber="26">
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<heading id="862F81A8FFEF891F9D9AFDB1FC60FD91" box="[667,921,512,539]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
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<taxonomicName id="1AD84D47FFEF891F9D9AFDB1FCCEFD91" authorityName="Karimbumkara & Priyadarsanan" authorityYear="2024" box="[667,823,512,539]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
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<emphasis id="EFACEAD6FFEF891F9D9AFDB1FCCEFD91" bold="true" box="[667,823,512,539]" italics="true" pageId="25" pageNumber="26">Onitis kethai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F49F57ADFFEF891F9C3FFDB0FC60FD91" box="[830,921,513,539]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="95C2654FFFEF891F9E74FD92FC0EFDEB" pageId="25" pageNumber="26" type="description">
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<paragraph id="DD6736C4FFEF891F9E74FD92FB47FDB7" blockId="25.[373,1214,512,610]" box="[373,1214,547,573]" pageId="25" pageNumber="26">
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<uri id="A9493AC6FFEF891F9E74FD92FB47FDB7" box="[373,1214,547,573]" pageId="25" pageNumber="26">
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urn:lsid:zoobank.org:act:
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<uuid id="A97E0C11FFEF891F9D96FD92FB47FDB7" box="[663,1214,547,573]" pageId="25" pageNumber="26">C3F62D8F-89D9-4034-A209-382401D8B861</uuid>
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</uri>
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</paragraph>
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<paragraph id="DD6736C4FFEF891F9D3DFDF6FC0EFDEB" blockId="25.[373,1214,512,610]" box="[572,1015,583,610]" pageId="25" pageNumber="26">
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<figureCitation id="45E32A41FFEF891F9D3DFDF6FD7EFDEB" box="[572,647,583,609]" captionStart="Fig" captionStartId="26.[189,232,1923,1949]" captionTargetBox="[299,1295,368,1851]" captionTargetId="figure-42@26.[191,1400,269,1864]" captionTargetPageId="26" captionText="Fig. 7. Onitis kethai sp. nov. A–E. Holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023B. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. E. Elytra. F. Paratype, ♀, habitus, dorsal view. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742869" httpUri="https://zenodo.org/record/13742869/files/figure.png" pageId="25" pageNumber="26">Figs 7</figureCitation>
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,
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<figureCitation id="45E32A41FFEF891F9D95FDF6FD27FDEB" box="[660,734,583,609]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="25" pageNumber="26">8A–D</figureCitation>
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,
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<figureCitation id="45E32A41FFEF891F9DEBFDF6FCF7FDEB" box="[746,782,583,609]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="25" pageNumber="26">9C</figureCitation>
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,
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<figureCitation id="45E32A41FFEF891F9C1AFDF6FCB7FDEB" box="[795,846,583,609]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="25" pageNumber="26">14C</figureCitation>
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,
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<figureCitation id="45E32A41FFEF891F9C5BFDF6FC77FDE8" box="[858,910,583,610]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="25" pageNumber="26">15C</figureCitation>
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,
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<figureCitation id="45E32A41FFEF891F9C9BFDF6FC34FDEB" box="[922,973,583,609]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="25" pageNumber="26">16C</figureCitation>
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,
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<figureCitation id="45E32A41FFEF891F9CDBFDF6FC0EFDEB" box="[986,1015,583,609]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="25" pageNumber="26">18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="95C2654FFFEF891F9FBCFD39FDFAFC60" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph id="DD6736C4FFEF891F9FBCFD39FE3FFD28" blockId="25.[189,454,648,674]" box="[189,454,648,674]" pageId="25" pageNumber="26">
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<heading id="862F81A8FFEF891F9FBCFD39FE3FFD28" bold="true" box="[189,454,648,674]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
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<emphasis id="EFACEAD6FFEF891F9FBCFD39FE3FFD28" bold="true" box="[189,454,648,674]" pageId="25" pageNumber="26">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="DD6736C4FFEF891F9FBCFD06FDFAFC60" blockId="25.[189,1399,695,1003]" pageId="25" pageNumber="26">
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<emphasis id="EFACEAD6FFEF891F9FBCFD06FEAAFD5B" box="[189,339,695,721]" italics="true" pageId="25" pageNumber="26">Onitiskethai</emphasis>
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<taxonomicNameLabel id="F49F57ADFFEF891F9E5BFD06FE4DFD5B" box="[346,436,695,721]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
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can be easily distinguished from
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<taxonomicName id="1AD84D47FFEF891F9C3DFD06FC35FD5B" authorityName="Fabricius" authorityYear="1801" box="[828,972,695,721]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
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<emphasis id="EFACEAD6FFEF891F9C3DFD06FC35FD5B" box="[828,972,695,721]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
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</taxonomicName>
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in having stronger and more closely placed punctures on the pronotum; genae smoothly rounded in
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<taxonomicName id="1AD84D47FFEF891F9CBFFD6BFBD4FD7E" authorityName="Karimbumkara & Priyadarsanan" authorityYear="2024" box="[958,1069,730,756]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
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<emphasis id="EFACEAD6FFEF891F9CBFFD6BFBD4FD7E" box="[958,1069,730,756]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
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</taxonomicName>
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, while strongly rounded in
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<taxonomicName id="1AD84D47FFEF891F9FBCFD4FFEA8FC9D" authorityName="Fabricius" authorityYear="1801" box="[189,337,765,792]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
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<emphasis id="EFACEAD6FFEF891F9FBCFD4FFEA8FC9D" box="[189,337,765,792]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
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</taxonomicName>
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, both inconspicuously punctured. The clypeus is rugulose in
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<taxonomicName id="1AD84D47FFEF891F9B3AFD4FFB48FC9D" authorityName="Karimbumkara & Priyadarsanan" authorityYear="2024" box="[1083,1201,765,792]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
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<emphasis id="EFACEAD6FFEF891F9B3AFD4FFB48FC9D" box="[1083,1201,765,792]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
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</taxonomicName>
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(
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<figureCitation id="45E32A41FFEF891F9BBCFD4CFAE3FC9D" box="[1213,1306,765,791]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="25" pageNumber="26">Fig. 9C</figureCitation>
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); while in
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<taxonomicName id="1AD84D47FFEF891F9FDCFC90FE96FCB0" authorityName="Fabricius" authorityYear="1801" box="[221,367,800,827]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
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<emphasis id="EFACEAD6FFEF891F9FDCFC90FE96FCB0" box="[221,367,800,827]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
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</taxonomicName>
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it is granular in male (
|
||
<figureCitation id="45E32A41FFEF891F9D85FC91FD1BFCB0" box="[644,738,800,826]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="25" pageNumber="26">Fig. 9D</figureCitation>
|
||
) and rugose in female.
|
||
<emphasis id="EFACEAD6FFEF891F9CFEFC91FB18FCB0" box="[1023,1249,800,826]" italics="true" pageId="25" pageNumber="26">Onitissinghalensis</emphasis>
|
||
differs from both these species in having strong rounded granules on clypeus and head while the genae are sparingly granular (
|
||
<figureCitation id="45E32A41FFEF891F9E33FCD6FE77FC0B" box="[306,398,871,897]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="25" pageNumber="26">Fig. 9F</figureCitation>
|
||
). In
|
||
<taxonomicName id="1AD84D47FFEF891F9EC9FCD6FDCEFC0B" authorityName="Karimbumkara & Priyadarsanan" authorityYear="2024" box="[456,567,871,897]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
|
||
<emphasis id="EFACEAD6FFEF891F9EC9FCD6FDCEFC0B" box="[456,567,871,897]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
|
||
</taxonomicName>
|
||
the space between the clypeal carina, frontal carina and the tubercle behind is strongly rugose, while in
|
||
<taxonomicName id="1AD84D47FFEF891F9D6DFC3BFD06FC2E" authorityName="Fabricius" authorityYear="1801" box="[620,767,906,932]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
|
||
<emphasis id="EFACEAD6FFEF891F9D6DFC3BFD06FC2E" box="[620,767,906,932]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1AD84D47FFEF891F9C3FFC3BFC0FFC2E" authorityName="Lansberge, AIMB" authorityYear="1875" box="[830,1014,906,932]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="singhalensis">
|
||
<emphasis id="EFACEAD6FFEF891F9C3FFC3BFC0FFC2E" box="[830,1014,906,932]" italics="true" pageId="25" pageNumber="26">O. singhalensis</emphasis>
|
||
</taxonomicName>
|
||
it is granular. Differences were also observed on comparing the aedeagus of
|
||
<taxonomicName id="1AD84D47FFEF891F9DEBFC1FFCA3FC4D" authorityName="Karimbumkara & Priyadarsanan" authorityYear="2024" box="[746,858,941,968]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
|
||
<emphasis id="EFACEAD6FFEF891F9DEBFC1FFCA3FC4D" box="[746,858,941,968]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="45E32A41FFEF891F9C6FFC1CFC0DFC4D" box="[878,1012,941,967]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="25" pageNumber="26">Fig. 8A–D</figureCitation>
|
||
),
|
||
<taxonomicName id="1AD84D47FFEF891F9B0FFC1FFB5AFC4D" authorityName="Fabricius" authorityYear="1801" box="[1038,1187,941,968]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
|
||
<emphasis id="EFACEAD6FFEF891F9B0FFC1FFB5AFC4D" box="[1038,1187,941,968]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="45E32A41FFEF891F9BB6FC1CFAC1FC4D" box="[1207,1336,941,967]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="25" pageNumber="26">Fig. 8E–H</figureCitation>
|
||
) and
|
||
<taxonomicName id="1AD84D47FFEF891F9FBCFC60FE8BFC60" authorityName="Lansberge, AIMB" authorityYear="1875" box="[189,370,976,1003]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="singhalensis">
|
||
<emphasis id="EFACEAD6FFEF891F9FBCFC60FE8BFC60" box="[189,370,976,1003]" italics="true" pageId="25" pageNumber="26">O. singhalensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="45E32A41FFEF891F9E83FC61FE0FFC60" box="[386,502,976,1002]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="25" pageNumber="26">Fig. 8I–L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95C2654FFFEF891F9FBCFBA3FC81FBF7" pageId="25" pageNumber="26" type="etymology">
|
||
<paragraph id="DD6736C4FFEF891F9FBCFBA3FEBDFBA6" blockId="25.[189,324,1042,1068]" box="[189,324,1042,1068]" pageId="25" pageNumber="26">
|
||
<heading id="862F81A8FFEF891F9FBCFBA3FEBDFBA6" bold="true" box="[189,324,1042,1068]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="EFACEAD6FFEF891F9FBCFBA3FEBDFBA6" bold="true" box="[189,324,1042,1068]" pageId="25" pageNumber="26">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFEF891F9FBCFBF1FC81FBF7" blockId="25.[189,1398,1088,1149]" pageId="25" pageNumber="26">Named in memory of our field assistant, the late Mr Ketha Gowda who assisted PDR in many of his field trips and helped in sampling dung beetles and other insects.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95C2654FFFEF891F9FBCFB14FD74F989" pageId="25" pageNumber="26" type="materials_examined">
|
||
<paragraph id="DD6736C4FFEF891F9FBCFB14FE15FB35" blockId="25.[189,492,1189,1215]" box="[189,492,1189,1215]" pageId="25" pageNumber="26">
|
||
<heading id="862F81A8FFEF891F9FBCFB14FE15FB35" bold="true" box="[189,492,1189,1215]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="EFACEAD6FFEF891F9FBCFB14FE15FB35" bold="true" box="[189,492,1189,1215]" pageId="25" pageNumber="26">Type material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="6DB03C99FFEF891F9FEBFB65FC7AFADD" collectingDate="1998-05-10" collectingMethod="open cow dung bait trap" collectionCode="NBAIR" collectorName="Priyadarsanan Dharma Rajan" country="India" county="Biligiri Rangaswamy" latitude="11.902133" location="Honnametti" longLatPrecision="1" longitude="77.19273" municipality="Tiger Reserve" pageId="25" pageNumber="26" specimenCode="NIM/NBAIR/COL/ONIT/H-271023B" specimenCount="2" specimenCount-male="1" stateProvince="Karnataka" typeStatus="holotype">
|
||
<paragraph id="DD6736C4FFEF891F9FEBFB65FEA5FB64" blockId="25.[189,1399,1236,1368]" box="[234,348,1236,1262]" pageId="25" pageNumber="26">
|
||
<emphasis id="EFACEAD6FFEF891F9FEBFB65FEA5FB64" bold="true" box="[234,348,1236,1262]" pageId="25" pageNumber="26">
|
||
<typeStatus id="02638866FFEF891F9FEBFB65FEA5FB64" box="[234,348,1236,1262]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFEF891F9FBCFB46FC7AFADD" blockId="25.[189,1399,1236,1368]" pageId="25" pageNumber="26">
|
||
<collectingCountry id="A5CF7654FFEF891F9FBCFB46FEF7FA9B" box="[189,270,1271,1297]" name="India" pageId="25" pageNumber="26">INDIA</collectingCountry>
|
||
•
|
||
<specimenCount id="CBDEFD4DFFEF891F9E28FB46FEBBFA9B" box="[297,322,1271,1297]" count="1" pageId="25" pageNumber="26" type="male">♂</specimenCount>
|
||
;
|
||
<collectingRegion id="1F1CF826FFEF891F9E4EFB46FE32FA9B" box="[335,459,1271,1297]" country="India" name="Karnataka" pageId="25" pageNumber="26">Karnataka</collectingRegion>
|
||
,
|
||
<collectingCounty id="34064E48FFEF891F9ED6FB46FD2BFA9B" box="[471,722,1271,1297]" pageId="25" pageNumber="26">Biligiri Rangaswamy</collectingCounty>
|
||
temple
|
||
<collectingMunicipality id="3D03ACBEFFEF891F9C30FB46FC20FA9B" box="[817,985,1271,1297]" pageId="25" pageNumber="26">Tiger Reserve</collectingMunicipality>
|
||
,
|
||
<location id="D807601FFFEF891F9CE7FB46FB88FA9B" LSID="urn:lsid:plazi:treatment:557187D2FFEF891D9D9AFDB1FE0CFC55:D807601FFFEF891F9CE7FB46FB88FA9B" box="[998,1137,1271,1297]" country="India" county="Biligiri Rangaswamy" latitude="11.902133" longLatPrecision="1" longitude="77.19273" municipality="Tiger Reserve" name="Honnametti" pageId="25" pageNumber="26" stateProvince="Karnataka">Honnametti</location>
|
||
shola;
|
||
<geoCoordinate id="B8EC5003FFEF891F9BC6FB44FA8BFA9B" box="[1223,1394,1269,1297]" degrees="11" direction="north" minutes="54" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="7.68" value="11.902133">11°54ʹ7.68″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B8EC5003FFEF891F9FBCFAA9FE8CFABE" box="[189,373,1304,1332]" degrees="77" direction="east" minutes="11" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="33.84" value="77.19273">77°11ʹ33.84″ E</geoCoordinate>
|
||
;
|
||
<collectingMethod id="04994ED3FFEF891F9E82FAABFD50FABE" box="[387,681,1306,1332]" pageId="25" pageNumber="26">open cow dung bait trap</collectingMethod>
|
||
;
|
||
<date id="A9661004FFEF891F9DB6FAABFCA0FABE" box="[695,857,1306,1332]" pageId="25" pageNumber="26" value="1998-05-10">
|
||
<collectingDate id="B922E9ECFFEF891F9DB6FAABFCA0FABE" box="[695,857,1306,1332]" pageId="25" pageNumber="26" value="1998-05-10">10 May 1998</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="702D5312FFEF891F9C66FAABFB3BFABE" box="[871,1218,1306,1332]" pageId="25" pageNumber="26">Priyadarsanan Dharma Rajan</collectorName>
|
||
leg.;
|
||
<specimenCount id="CBDEFD4DFFEF891F9A09FAABFA8EFABE" box="[1288,1399,1306,1332]" pageId="25" pageNumber="26" type="generic">specimen</specimenCount>
|
||
deposited at
|
||
<collectionCode id="BBC9AE01FFEF891F9E50FA8CFE56FADD" box="[337,431,1341,1367]" pageId="25" pageNumber="26">NBAIR</collectionCode>
|
||
,
|
||
<specimenCode id="8D7E9EBFFFEF891F9EBAFA8CFC86FADD" box="[443,895,1341,1367]" pageId="25" pageNumber="26">NIM/NBAIR/COL/ONIT/H-271023B</specimenCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<materialsCitation id="6DB03C99FFEF891F9FEBFACEFD70F989" collectingDate="1998-05-10" collectingMethod="open cow dung bait trap" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan" country="India" county="Biligiri Rangaswamy" latitude="11.902133" location="Tiger Reserve" longLatPrecision="1" longitude="77.19273" municipality="Tiger Reserve" pageId="25" pageNumber="26" specimenCode="Co/Sc1000293 to 1000305" specimenCount="26" specimenCount-female="5" specimenCount-male="8" stateProvince="Karnataka" typeStatus="paratype">
|
||
<paragraph id="DD6736C4FFEF891F9FEBFACEFDDBFA12" blockId="25.[189,1399,1406,1539]" box="[234,546,1406,1433]" pageId="25" pageNumber="26">
|
||
<emphasis id="EFACEAD6FFEF891F9FEBFACEFE9EFA13" bold="true" box="[234,359,1407,1433]" pageId="25" pageNumber="26">
|
||
<typeStatus id="02638866FFEF891F9FEBFACEFE9EFA13" box="[234,359,1407,1433]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="CBDEFD4DFFEF891F9E79FACEFDE3FA12" box="[376,538,1406,1433]" count="13" pageId="25" pageNumber="26" type="generic">13 specimens</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFEF891F9FBCFA13FD74F989" blockId="25.[189,1399,1406,1539]" pageId="25" pageNumber="26">
|
||
<collectingCountry id="A5CF7654FFEF891F9FBCFA13FEF7FA36" box="[189,270,1442,1468]" name="India" pageId="25" pageNumber="26">INDIA</collectingCountry>
|
||
•
|
||
<specimenCount id="CBDEFD4DFFEF891F9E33FA13FE82FA36" box="[306,379,1442,1468]" count="8" pageId="25" pageNumber="26" type="male">8 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="CBDEFD4DFFEF891F9E8CFA13FE2CFA36" box="[397,469,1442,1469]" count="5" pageId="25" pageNumber="26" type="female">5 ♀♀</specimenCount>
|
||
;
|
||
<collectingRegion id="1F1CF826FFEF891F9EE9FA13FD9DFA36" box="[488,612,1442,1468]" country="India" name="Karnataka" pageId="25" pageNumber="26">Karnataka</collectingRegion>
|
||
,
|
||
<collectingCounty id="34064E48FFEF891F9D74FA13FC8DFA36" box="[629,884,1442,1468]" pageId="25" pageNumber="26">Biligiri Rangaswamy</collectingCounty>
|
||
temple
|
||
<collectingMunicipality id="3D03ACBEFFEF891F9CDDFA13FB71FA36" box="[988,1160,1442,1468]" pageId="25" pageNumber="26">Tiger Reserve</collectingMunicipality>
|
||
, Honnametti shola;
|
||
<geoCoordinate id="B8EC5003FFEF891F9FBCFA72FE90FA55" box="[189,361,1475,1503]" degrees="11" direction="north" minutes="54" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="7.68" value="11.902133">11°54ʹ7.68″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B8EC5003FFEF891F9E77FA72FDD4FA55" box="[374,557,1475,1503]" degrees="77" direction="east" minutes="11" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="33.84" value="77.19273">77°11ʹ33.84″ E</geoCoordinate>
|
||
;
|
||
<collectingMethod id="04994ED3FFEF891F9D3AFA74FCA6FA55" box="[571,863,1477,1503]" pageId="25" pageNumber="26">open cow dung bait trap</collectingMethod>
|
||
;
|
||
<date id="A9661004FFEF891F9C6DFA74FBF6FA55" box="[876,1039,1477,1503]" pageId="25" pageNumber="26" value="1998-05-10">
|
||
<collectingDate id="B922E9ECFFEF891F9C6DFA74FBF6FA55" box="[876,1039,1477,1503]" pageId="25" pageNumber="26" value="1998-05-10">10 May 1998</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="702D5312FFEF891F9B1DFA74FA8FFA55" box="[1052,1398,1477,1503]" pageId="25" pageNumber="26">Priyadarsanan Dharma Rajan</collectorName>
|
||
leg.;
|
||
<collectionCode id="BBC9AE01FFEF891F9FF7FA59FEBAF988" box="[246,323,1512,1538]" pageId="25" pageNumber="26">AIMB</collectionCode>
|
||
/
|
||
<specimenCode id="8D7E9EBFFFEF891F9E4DFA59FD70F989" box="[332,649,1512,1539]" pageId="25" pageNumber="26">Co/Sc1000293 to 1000305</specimenCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="95C2654FFFEF891D9FBCF998FA8EFCC6" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="DD6736C4FFEF891F9FBCF998FEB7F9C9" blockId="25.[189,334,1577,1603]" box="[189,334,1577,1603]" pageId="25" pageNumber="26">
|
||
<heading id="862F81A8FFEF891F9FBCF998FEB7F9C9" bold="true" box="[189,334,1577,1603]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="EFACEAD6FFEF891F9FBCF998FEB7F9C9" bold="true" box="[189,334,1577,1603]" pageId="25" pageNumber="26">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFEF891F9FBCF9E9FEB6F91C" blockId="25.[189,1399,1624,1686]" pageId="25" pageNumber="26">MEASUREMENTS (in mm). TL=11.5–15; BW =6–7.5; PL=4–6; PW= 6–8; EL=6.5–8; HL=3–3.5; HW=3–4.5.</paragraph>
|
||
<paragraph id="DD6736C4FFEF891F9FEBF90CFE52F95D" blockId="25.[189,1399,1725,1998]" box="[234,427,1725,1751]" pageId="25" pageNumber="26">
|
||
<emphasis id="EFACEAD6FFEF891F9FEBF90CFED3F95D" bold="true" box="[234,298,1725,1751]" pageId="25" pageNumber="26">Male</emphasis>
|
||
(
|
||
<typeStatus id="02638866FFEF891F9E38F90CFE5AF95D" box="[313,419,1725,1751]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFEF891D9FBCF951FDC6FD09" blockId="25.[189,1399,1725,1998]" lastBlockId="27.[189,1399,265,643]" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
|
||
Body, elytra, head, legs, antenna and mouthparts reddish, elytra red with a tinge of metallic green; oval, slightly convex. Head (
|
||
<figureCitation id="45E32A41FFEF891F9D1BF8B2FD8EF897" box="[538,631,1795,1821]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="25" pageNumber="26">Fig. 9C</figureCitation>
|
||
) closely granular with some rugosity, hind part almost smooth; clypeus parabolic, rugulose, margin slightly reflexed; with a feeble emargination in the middle, separated from the vertex by an interrupted frontal carina with a short curved clypeal carina and a blunt tubercle behind; genae with scattered imperceptible punctures. Pronotum (
|
||
<figureCitation id="45E32A41FFEF891F9B95F8DCFAF8F80D" box="[1172,1281,1901,1927]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="25" pageNumber="26">Fig. 14C</figureCitation>
|
||
) strongly and closely punctured in the middle, less closely at the sides, more strongly and closely pitted anteriorly, with a very slight median line starting above the basal impression, extending almost up to the anterior margin as thin smooth area; anterior sides of pronotum slightly curved, strongly rounded in the middle, bisinuate behind; front angles sharp, base bluntly angulate in the middle; basal fovea deep near the base with imperceptible punctures in circular rows at the periphery. Elytra strongly striate, striae without punctures, elytral intervals finely and sparsely punctured with the 1
|
||
<superScript id="2AAD9B8CFFED891D9A62FEC3FA96FE0A" attach="none" box="[1379,1391,370,384]" fontSize="6" pageId="27" pageNumber="28">st</superScript>
|
||
, 3
|
||
<superScript id="2AAD9B8CFFED891D9FCDFE24FF23FE29" attach="left" box="[204,218,405,419]" fontSize="6" pageId="27" pageNumber="28">rd</superScript>
|
||
and the 5
|
||
<superScript id="2AAD9B8CFFED891D9E50FE24FEA7FE29" attach="left" box="[337,350,405,419]" fontSize="6" pageId="27" pageNumber="28">th</superScript>
|
||
intervals convex, sutural interval shining while the rest opaque. Pygidium not shining, imperceptibly punctured. Metasternum with deep broad longitudinal groove in the front, metasternal shield imperceptibly punctured, with scattered aciculate punctures at the sides anteriorly; sides of the metasternum granular and thinly covered with long hairs. Protibia slightly curved with four strong external teeth and a sharp process at the tip, profemur not toothed; mesofemur (
|
||
<figureCitation id="45E32A41FFED891D9B89FD93FB0CFDB6" box="[1160,1269,546,572]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="27" pageNumber="28">Fig. 16C</figureCitation>
|
||
) plain and curved near the tibial joint where it forms a sharp tooth-like process on both upper and lower ridges; hind trochanter slightly toothed.
|
||
</paragraph>
|
||
<caption id="89A7664CFFEC891C9FBCF832FD79F86E" ID-DOI="http://doi.org/10.5281/zenodo.13742869" ID-Zenodo-Dep="13742869" httpUri="https://zenodo.org/record/13742869/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1923,1949]" targetBox="[299,1295,368,1851]" targetPageId="26">
|
||
<paragraph id="DD6736C4FFEC891C9FBCF832FD79F86E" blockId="26.[189,1398,1923,2020]" pageId="26" pageNumber="27">
|
||
<emphasis id="EFACEAD6FFEC891C9FBCF832FEF6F814" bold="true" box="[189,271,1923,1950]" pageId="26" pageNumber="27">Fig. 7.</emphasis>
|
||
<taxonomicName id="1AD84D47FFEC891C9E1BF835FE4CF817" authorityName="Karimbumkara & Priyadarsanan" authorityYear="2024" box="[282,437,1923,1950]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
|
||
<emphasis id="EFACEAD6FFEC891C9E1BF835FE4CF817" box="[282,437,1923,1950]" italics="true" pageId="26" pageNumber="27">Onitis kethai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F49F57ADFFEC891C9EBEF835FDE7F814" box="[447,542,1924,1950]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="EFACEAD6FFEC891C9D29F832FDA6F817" bold="true" box="[552,607,1923,1949]" pageId="26" pageNumber="27">A–E</emphasis>
|
||
. Holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023B.
|
||
<emphasis id="EFACEAD6FFEC891C9BEDF832FAF8F817" bold="true" box="[1260,1281,1923,1949]" pageId="26" pageNumber="27">A</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="EFACEAD6FFEC891C9E57F816FE93F84B" bold="true" box="[342,362,1959,1985]" pageId="26" pageNumber="27">B</emphasis>
|
||
. Habitus, ventral view.
|
||
<emphasis id="EFACEAD6FFEC891C9D8BF817FD66F84A" bold="true" box="[650,671,1958,1984]" pageId="26" pageNumber="27">C</emphasis>
|
||
. Head, dorsal view.
|
||
<emphasis id="EFACEAD6FFEC891C9C99F816FC54F84B" bold="true" box="[920,941,1959,1985]" pageId="26" pageNumber="27">D</emphasis>
|
||
. Pygidium.
|
||
<emphasis id="EFACEAD6FFEC891C9B40F816FBACF84B" bold="true" box="[1089,1109,1959,1985]" pageId="26" pageNumber="27">E</emphasis>
|
||
. Elytra.
|
||
<emphasis id="EFACEAD6FFEC891C9BBEF816FB28F84B" bold="true" box="[1215,1233,1959,1985]" pageId="26" pageNumber="27">F</emphasis>
|
||
. Paratype, ♀, habitus, dorsal view. (Images: AIMB).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD6736C4FFED891D9FBCFD1AFAA3FD62" blockId="27.[189,1399,682,744]" pageId="27" pageNumber="28">
|
||
AEDEAGUS (in mm) (
|
||
<figureCitation id="45E32A41FFED891D9EACFD1BFDD6FD4F" box="[429,559,682,709]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. E–H. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. I–L. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. M–P. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742873" httpUri="https://zenodo.org/record/13742873/files/figure.png" pageId="27" pageNumber="28">Fig. 8A–D</figureCitation>
|
||
). LP= 2.5, Lp=2.2; BP=1.3, BpB=1, BpT=0.5. Phallobase longer than parameres. Parameres slightly sinuate near the base, curved inwards anteriorly and hooked at the tip.
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFED891D9FEBFCBEFEBCFCA3" blockId="27.[189,1399,783,845]" box="[234,325,783,809]" pageId="27" pageNumber="28">
|
||
<emphasis id="EFACEAD6FFED891D9FEBFCBEFEBCFCA3" bold="true" box="[234,325,783,809]" pageId="27" pageNumber="28">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFED891D9FBCFC83FA8EFCC6" blockId="27.[189,1399,783,845]" box="[189,1399,818,845]" pageId="27" pageNumber="28">Clypeus closely rugose; protibia with four strong teeth and an articulated spur; no protrusions on the legs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95C2654FFFED891D9FBCFCC2FE0CFC55" pageId="27" pageNumber="28" type="biology_ecology">
|
||
<paragraph id="DD6736C4FFED891D9FBCFCC2FEE4FC07" blockId="27.[189,285,883,909]" box="[189,285,883,909]" pageId="27" pageNumber="28">
|
||
<heading id="862F81A8FFED891D9FBCFCC2FEE4FC07" bold="true" box="[189,285,883,909]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="EFACEAD6FFED891D9FBCFCC2FEE4FC07" bold="true" box="[189,285,883,909]" pageId="27" pageNumber="28">Habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DD6736C4FFED891D9FBCFC13FE0CFC55" blockId="27.[189,1399,930,992]" pageId="27" pageNumber="28">
|
||
The collections were made in the cloud forests of Honnametti in Biligiri Rangaswamy Temple Tiger Reserve,
|
||
<collectingRegion id="1F1CF826FFED891D9E2BFC74FE5FFC55" box="[298,422,965,991]" country="India" name="Karnataka" pageId="27" pageNumber="28">Karnataka</collectingRegion>
|
||
,
|
||
<collectingCountry id="A5CF7654FFED891D9EB0FC74FE08FC55" box="[433,497,965,991]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |