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<document id="59DE2C96B7EFDBBE05B3CE11CB4604C9" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-Zenodo-Dep="13742850" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFE789159D7BFEB8FB02FE12" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis crassus Sharp 1875" docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="18" updateTime="1726490331196" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AD0DF08B8CB7CA18FA3E51D3963215FC">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="393FBC4F206B6A47DEED75228998FBFC">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:affiliation id="DBFE0131D6B87C3558ABCB4A6274AADD">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="22B2A93D84F22FB6CCD0773ADD3AA71D" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
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<mods:namePart id="DA11C91C0C367CC006087614C7A38E5B">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="99FC6ABB42FECEFD1B15E965FB4533D0">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:title id="F1DEC1138C0A5F6E5E2130994D115F2C">European Journal of Taxonomy</mods:title>
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<mods:date id="131BAC5A416CEFFAC935250487E5D43F">2024</mods:date>
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<treatment id="557187D2FFE789159D7BFEB8FB02FE12" LSID="urn:lsid:plazi:treatment:557187D2FFE789159D7BFEB8FB02FE12" httpUri="http://treatment.plazi.org/id/557187D2FFE789159D7BFEB8FB02FE12" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="95C2654FFFE789179D7BFEB8FC43FEAE" box="[634,954,265,292]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="DD6736C4FFE789179D7BFEB8FC43FEAE" blockId="17.[634,954,265,334]" box="[634,954,265,292]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179D7BFEB8FC43FEAE" box="[634,954,265,292]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="1AD84D47FFE789179D7BFEB8FC43FEAE" ID-CoL="49KFD" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[634,954,265,292]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179D7BFEB8FCDCFEA9" bold="true" box="[634,805,265,291]" italics="true" pageId="17" pageNumber="18">Onitis crassus</emphasis>
Sharp, 1875
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</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179DD4FE82FCA7FEC7" box="[725,862,307,334]" pageId="17" pageNumber="18" type="description">
<paragraph id="DD6736C4FFE789179DD4FE82FCA7FEC7" blockId="17.[634,954,265,334]" box="[725,862,307,334]" pageId="17" pageNumber="18">
<figureCitation id="45E32A41FFE789179DD4FE82FCCCFEC4" box="[725,821,307,334]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="17" pageNumber="18">Figs 5D</figureCitation>
,
<figureCitation id="45E32A41FFE789179C40FE82FCA7FEC7" box="[833,862,307,333]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="17" pageNumber="18">18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCFECDFDF0FD32" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="DD6736C4FFE789179FBCFECDFCD7FE1C" blockId="17.[189,814,379,406]" box="[189,814,379,406]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FBCFECDFE9DFE1C" box="[189,356,380,406]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Sharp, 1875: 48 (original description).
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFE74FAFDFDE1" blockId="17.[189,1399,452,697]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="FDC811EAFFE789179FBCFE74FAFDFDE1" pageId="17" pageNumber="18">
<taxonomicName id="1AD84D47FFE789179FBCFE74FE9DFE54" ID-CoL="49KFD" authorityName="Sharp" authorityYear="1875" box="[189,356,453,479]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179FBCFE74FE9DFE54" box="[189,356,453,479]" italics="true" pageId="17" pageNumber="18">Onitis crassus</emphasis>
</taxonomicName>
–
<treatmentCitation id="5C7910D5FFE789179E7EFE75FDB0FE54" author="Arrow G. J." box="[383,585,452,478]" page="400" pageId="17" pageNumber="18" year="1931">
<bibRefCitation id="B9494B35FFE789179E7EFE75FDB0FE54" author="Arrow G. J." box="[383,585,452,478]" pageId="17" pageNumber="18" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 400</bibRefCitation>
</treatmentCitation>
(keys and description). —
<treatmentCitation id="5C7910D5FFE789179C89FE74FBA2FE54" author="Janssens A." box="[904,1115,452,479]" page="73" pageId="17" pageNumber="18" year="1937">
<bibRefCitation id="B9494B35FFE789179C89FE74FBA2FE54" author="Janssens A." box="[904,1115,452,479]" pageId="17" pageNumber="18" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 73</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFE789179A09FE75FEADFD88" author="Balthasar V." page="45" pageId="17" pageNumber="18" year="1963">
<bibRefCitation id="B9494B35FFE789179A09FE75FEADFD88" author="Balthasar V." pageId="17" pageNumber="18" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar 1963: 45</bibRefCitation>
</treatmentCitation>
(monograph). —
<treatmentCitation id="5C7910D5FFE789179D2EFE56FCEDFD8B" author="Chandra K." box="[559,788,487,513]" page="360" pageId="17" pageNumber="18" year="2000">
<bibRefCitation id="B9494B35FFE789179D2EFE56FCEDFD8B" author="Chandra K." box="[559,788,487,513]" pageId="17" pageNumber="18" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
</treatmentCitation>
(distribution); 2008: 176 (distribution). —
<treatmentCitation id="5C7910D5FFE789179A1EFE56FE28FDAF" author="Bezdek A. &amp; Krell F. - T." page="159" pageId="17" pageNumber="18" year="2006">
<bibRefCitation id="B9494B35FFE789179A1EFE56FE28FDAF" author="Bezdek A. &amp; Krell F. - T." pageId="17" pageNumber="18" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE789179DB4FDBBFBD6FDAF" author="Chandra K. &amp; Ahirwar S. C." box="[693,1071,522,549]" page="287" pageId="17" pageNumber="18" year="2007">
<bibRefCitation id="B9494B35FFE789179DB4FDBBFBD6FDAF" author="Chandra K. &amp; Ahirwar S. C." box="[693,1071,522,549]" pageId="17" pageNumber="18" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007: 287</bibRefCitation>
</treatmentCitation>
(distribution). — Chandra
<emphasis id="EFACEAD6FFE789179FEBFD9EFED0FDC2" box="[234,297,558,584]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2012b: 29 (distribution). —
<treatmentCitation id="5C7910D5FFE789179D8EFD9FFC38FDC2" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[655,961,558,584]" page="299" pageId="17" pageNumber="18" year="2014">
<bibRefCitation id="B9494B35FFE789179D8EFD9FFC38FDC2" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[655,961,558,584]" pageId="17" pageNumber="18" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFE789179DFCFD9EFCC5FDC2" box="[765,828,558,584]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2014: 299
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE789179BA1FD9FFA8EFDC2" author="Ali S. M. S. &amp; Naeem M. &amp; Baig F. &amp; Shazad A. &amp; Zia A." box="[1184,1399,558,585]" page="4" pageId="17" pageNumber="18" year="2015">
<bibRefCitation id="B9494B35FFE789179BA1FD9FFA8EFDC2" author="Ali S. M. S. &amp; Naeem M. &amp; Baig F. &amp; Shazad A. &amp; Zia A." box="[1184,1399,558,585]" pageId="17" pageNumber="18" pagination="1 - 6" refId="ref26882" refString="Ali S. M. S., Naeem M., Baig F., Shazad A. &amp; Zia A. 2015. New records, distributional notes and species diversity of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Pothohar Plateau of Punjab. Pakistan. Journal of Entomology and Zoology Studies 3 (3): 1 - 6. Available from: https: // www. entomoljournal. com / archives / 2015 / vol 3 issue 3 / PartA / 3 - 3 - 23. pdf [accessed 27 Nov. 2019]" type="journal article" year="2015">
Ali
<emphasis id="EFACEAD6FFE789179BD1FD9EFAF6FDC2" box="[1232,1295,558,584]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2015: 4
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE789179EB9FDE0FD36FDE1" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[440,719,593,620]" page="1037" pageId="17" pageNumber="18" year="2015">
<bibRefCitation id="B9494B35FFE789179EB9FDE0FD36FDE1" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[440,719,593,620]" pageId="17" pageNumber="18" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE789179D06FDE3FDBBFDE1" box="[519,578,593,619]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="5C7910D5FFE789179C84FDE0FBA3FDE1" author="Bezdek A." box="[901,1114,593,619]" page="179" pageId="17" pageNumber="18" year="2016">
<bibRefCitation id="B9494B35FFE789179C84FDE0FBA3FDE1" author="Bezdek A." box="[901,1114,593,619]" pageId="17" pageNumber="18" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFDCAFDF0FD32" blockId="17.[189,1399,452,697]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FBCFDCAFE9CFD1F" box="[189,357,635,661]" italics="true" pageId="17" pageNumber="18">Onitisvischnu</emphasis>
–
<bibRefCitation id="B9494B35FFE789179E83FDCAFD28FD1F" author="Preudhomme de Borre C. F. P. A." box="[386,721,635,662]" pageId="17" pageNumber="18" refId="ref30731" refString="Preudhomme de Borre C. F. P. A. 1881. Bulletin ou Comptes-Rendus des Seances de la Societe entomologique de Belgique XXXIX." type="book" year="1881">Preudhomme de Borre 1881</bibRefCitation>
: xxxix (synonymised by
<bibRefCitation id="B9494B35FFE789179CF5FDCAFB7FFD1F" author="Arrow G. J." box="[1012,1158,635,661]" pageId="17" pageNumber="18" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931</bibRefCitation>
). —
<bibRefCitation id="B9494B35FFE789179BBFFDCAFEDCFD33" author="Schoolmeesters P." pageId="17" pageNumber="18" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCFD56FE9AFC4E" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="DD6736C4FFE789179FBCFD56FE3FFC8B" blockId="17.[189,454,743,769]" box="[189,454,743,769]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179FBCFD56FE3FFC8B" bold="true" box="[189,454,743,769]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="EFACEAD6FFE789179FBCFD56FE3FFC8B" bold="true" box="[189,454,743,769]" pageId="17" pageNumber="18">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFCACFE9AFC4E" blockId="17.[189,1399,796,964]" pageId="17" pageNumber="18">
<taxonomicName id="1AD84D47FFE789179FBCFCACFE91FCBC" authorityName="Sharp" authorityYear="1875" box="[189,360,797,823]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179FBCFCACFE91FCBC" box="[189,360,797,823]" italics="true" pageId="17" pageNumber="18">Onitis crassus</emphasis>
</taxonomicName>
is closer to
<taxonomicName id="1AD84D47FFE789179D06FCACFD5AFCBC" box="[519,675,797,823]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFE789179D06FCACFD5AFCBC" box="[519,675,797,823]" italics="true" pageId="17" pageNumber="18">O. castaneus</emphasis>
</taxonomicName>
in having hairy pygidium and uninterrupted clypeo-frontal carina but differs from the latter in having short scanty setae on the pygidium; frontal tubercle being placed behind the carina while in
<taxonomicName id="1AD84D47FFE789179D5AFCD2FD0FFCF7" box="[603,758,867,893]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFE789179D5AFCD2FD0FFCF7" box="[603,758,867,893]" italics="true" pageId="17" pageNumber="18">O. castaneus</emphasis>
</taxonomicName>
it is placed on the carina and the anterior margin of clypeus being gently excised in the middle while in the latter it is reflexed and straight. See diagnosis of
<taxonomicName id="1AD84D47FFE789179FDCFC1BFEA5FC49" authorityName="Lansberge" authorityYear="1875" box="[221,348,937,964]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFE789179FDCFC1BFEA5FC49" box="[221,348,937,964]" italics="true" pageId="17" pageNumber="18">O. brahma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCFC43FCECFA5F" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="DD6736C4FFE789179FBCFC43FE15FB81" blockId="17.[189,492,1009,1036]" box="[189,492,1009,1036]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179FBCFC43FE15FB81" bold="true" box="[189,492,1009,1036]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="EFACEAD6FFE789179FBCFC43FE15FB81" bold="true" box="[189,492,1009,1036]" pageId="17" pageNumber="18">Type material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFE789179FEBFB96FBD5FB05" collectionCode="MNHN" collectorName="A. Janssens" country="India" location="INDIA" pageId="17" pageNumber="18" specimenCode="EC4813" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<paragraph id="DD6736C4FFE789179FEBFB96FEA5FBCB" blockId="17.[234,348,1063,1089]" box="[234,348,1063,1089]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBFB96FEA5FBCB" bold="true" box="[234,348,1063,1089]" pageId="17" pageNumber="18">
<typeStatus id="02638866FFE789179FEBFB96FEA5FBCB" box="[234,348,1063,1089]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFBE3FBD5FB05" blockId="17.[189,1398,1105,1167]" pageId="17" pageNumber="18">
<collectingCountry id="A5CF7654FFE789179FBCFBE3FEF7FBE6" box="[189,270,1106,1132]" name="India" pageId="17" pageNumber="18">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFE789179E29FBE3FEC6FBE6" box="[296,319,1106,1132]" count="1" pageId="17" pageNumber="18" type="female">♀</specimenCount>
; “
<emphasis id="EFACEAD6FFE789179E5AFBE3FDFBFBE6" box="[347,514,1106,1132]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Type D.S.;
<collectorName id="702D5312FFE789179D91FBE3FC74FBE1" box="[656,909,1105,1132]" pageId="17" pageNumber="18">Ex. Musaeo D. Sharp</collectorName>
1890,
<collectorName id="702D5312FFE789179CDEFBE3FB90FBE6" box="[991,1129,1106,1132]" pageId="17" pageNumber="18">G. J. Arrow</collectorName>
vid. 1928,
<collectorName id="702D5312FFE789179BECFBE3FA8FFBE6" box="[1261,1398,1106,1132]" pageId="17" pageNumber="18">A. Janssens</collectorName>
vid., 1936:
<emphasis id="EFACEAD6FFE789179E42FBC4FE13FB05" box="[323,490,1141,1167]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Sharp; Holotype (Red Label)”;
<collectionCode id="BBC9AE01FFE789179C62FBC4FC44FB05" box="[867,957,1141,1167]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFE789179CC5FBC4FBD1FB05" box="[964,1064,1141,1167]" collectionCode="EC" pageId="17" pageNumber="18">EC4813</specimenCode>
.
</paragraph>
</materialsCitation>
<paragraph id="DD6736C4FFE789179FEBFB0CFCF5FB5D" blockId="17.[234,780,1213,1239]" box="[234,780,1213,1239]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBFB0CFD9BFB5D" bold="true" box="[234,610,1213,1239]" pageId="17" pageNumber="18">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFE789179D70FB0CFCFDFB5D" box="[625,772,1213,1239]" count="4" pageId="17" pageNumber="18" type="generic">4 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFB59FCECFA5F" blockId="17.[189,1399,1255,1493]" pageId="17" pageNumber="18">
<materialsCitation id="6DB03C99FFE789179FBCFB59FC44FAAF" collectionCode="BMNH" country="India" determinerName="G. J. A. Onitis Vischnu" location="Borre" pageId="17" pageNumber="18" specimenCode="(E) 1237163" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFE789179FBCFB59FEF7FA88" box="[189,270,1256,1282]" name="India" pageId="17" pageNumber="18">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFE789179E2DFB59FEBCFA88" box="[300,325,1256,1282]" count="1" pageId="17" pageNumber="18" type="male">♂</specimenCount>
; “Determined from description
<determinerName id="28BBDA00FFE789179DC2FB59FC3CFA8B" box="[707,965,1255,1282]" pageId="17" pageNumber="18">
G.J.A.
<taxonomicName id="1AD84D47FFE789179C19FB59FC3CFA8B" authority=", de Borre" authorityName="de Borre" box="[792,965,1255,1282]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">
<emphasis id="EFACEAD6FFE789179C19FB59FC3CFA8B" box="[792,965,1255,1282]" italics="true" pageId="17" pageNumber="18">Onitis Vischnu</emphasis>
</taxonomicName>
</determinerName>
, de
<location id="D807601FFFE789179CFDFB59FBC6FA88" LSID="urn:lsid:plazi:treatment:557187D2FFE789159D7BFEB8FB02FE12:D807601FFFE789179CFDFB59FBC6FA88" box="[1020,1087,1256,1282]" country="India" name="Borre" pageId="17" pageNumber="18">Borre</location>
;
<taxonomicName id="1AD84D47FFE789179B51FB59FECEFAAF" authority="de Borre, Compared" authorityName="de Borre, Compared" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">
<emphasis id="EFACEAD6FFE789179B51FB59FB04FA8B" box="[1104,1277,1255,1282]" italics="true" pageId="17" pageNumber="18">Onitis Vischnu</emphasis>
de Borre, Compared
</taxonomicName>
with type G.J.A., 6745, 7030”;
<collectionCode id="BBC9AE01FFE789179DC2FABAFCE2FAAF" box="[707,795,1291,1317]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="17" pageNumber="18">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFE789179C27FABAFC44FAAF" box="[806,957,1291,1317]" pageId="17" pageNumber="18">(E) 1237163</specimenCode>
</materialsCitation>
•
<materialsCitation id="6DB03C99FFE789179CDDFABAFAC2FAC2" collectingMethod="Cattle dung on sand" collectionCode="BMNH" collectorName="Dehra Dun" country="India" location="River Suswa" pageId="17" pageNumber="18" specimenCode="(E) 1237164" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFE789179CDDFABAFC0AFAAF" box="[988,1011,1291,1317]" count="1" pageId="17" pageNumber="18" type="female">♀</specimenCount>
; “
<collectorName id="702D5312FFE789179B0EFABAFB6EFAAF" box="[1039,1175,1291,1317]" pageId="17" pageNumber="18">Dehra Dun</collectorName>
Coll. B.M. 1931– 2.,
<location id="D807601FFFE789179FE5FA9FFE84FAC2" LSID="urn:lsid:plazi:treatment:557187D2FFE789159D7BFEB8FB02FE12:D807601FFFE789179FE5FA9FFE84FAC2" box="[228,381,1326,1352]" country="India" name="River Suswa" pageId="17" pageNumber="18">River Suswa</location>
,
<location id="D807601FFFE789179E8DFA9FFDFAFAC2" LSID="urn:lsid:plazi:treatment:557187D2FFE789159D7BFEB8FB02FE12:D807601FFFE789179E8DFA9FFDFAFAC2" box="[396,515,1326,1352]" country="India" name="Dehradun" pageId="17" pageNumber="18">Dehradun</location>
,
<location id="D807601FFFE789179D11FA9FFD30FAC2" LSID="urn:lsid:plazi:treatment:557187D2FFE789159D7BFEB8FB02FE12:D807601FFFE789179D11FA9FFD30FAC2" box="[528,713,1326,1352]" country="India" name="U. P. C. Beeson" pageId="17" pageNumber="18">U.P. C. Beeson</location>
. XI.’30;
<collectingMethod id="04994ED3FFE789179C34FA9FFBD2FAC2" box="[821,1067,1326,1352]" pageId="17" pageNumber="18">Cattle dung on sand</collectingMethod>
”;
<collectionCode id="BBC9AE01FFE789179B45FA9FFB65FAC2" box="[1092,1180,1326,1352]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="17" pageNumber="18">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFE789179BA4FA9FFAC2FAC2" box="[1189,1339,1326,1352]" pageId="17" pageNumber="18">(E) 1237164</specimenCode>
</materialsCitation>
•
<materialsCitation id="6DB03C99FFE789179A59FA9FFA8FFA3B" collectorName="Coll. J. Thomson" country="India" location="India" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFE789179A59FA9FFA88FAC2" box="[1368,1393,1326,1352]" count="1" pageId="17" pageNumber="18" type="male">♂</specimenCount>
; “A, Janssens vid. 1936:
<taxonomicName id="1AD84D47FFE789179EDAFAE0FD1BFAE1" authority="Sharp" authorityName="Sharp" authorityYear="1875" box="[475,738,1361,1387]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179EDAFAE0FD6FFAE1" box="[475,662,1361,1387]" italics="true" pageId="17" pageNumber="18">ONITIS crassus</emphasis>
Sharp
</taxonomicName>
;
<emphasis id="EFACEAD6FFE789179DF2FAE3FC62FAE6" box="[755,923,1362,1388]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Shp. Revis. Arrow 1926;
<emphasis id="EFACEAD6FFE789179BCEFAE3FA8EFAE6" box="[1231,1399,1362,1388]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Shp. Compared with type G.J.A.; cf. Ann. Soc. Ent. Belg. Xxv, 1881, C.R. p. 39; Ex-Typis de
<taxonomicName id="1AD84D47FFE789179A17FAC5FEF9FA38" authority="Borre" authorityName="Borre" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">Vischnu Borre</taxonomicName>
; det: Predhomme Borre,
<taxonomicName id="1AD84D47FFE789179D29FA29FCE4FA38" authority="Borre" authorityName="Borre" box="[552,797,1431,1458]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">
<emphasis id="EFACEAD6FFE789179D29FA29FD2AFA3B" box="[552,723,1431,1458]" italics="true" pageId="17" pageNumber="18">Onitis Vischnu</emphasis>
Borre
</taxonomicName>
;
<emphasis id="EFACEAD6FFE789179C2DFA29FC67FA3B" box="[812,926,1431,1458]" italics="true" pageId="17" pageNumber="18">O. Vishnu</emphasis>
, Reiche,
<collectingCountry id="A5CF7654FFE789179B0CFA26FBB5FA3B" box="[1037,1100,1431,1457]" name="India" pageId="17" pageNumber="18">India</collectingCountry>
, 3050,
<collectorName id="702D5312FFE789179BA1FA26FA94FA3B" box="[1184,1389,1431,1458]" pageId="17" pageNumber="18">Coll. J. Thomson</collectorName>
”
</materialsCitation>
•
<materialsCitation id="6DB03C99FFE789179FCFFA0AFCE8FA5F" box="[206,785,1467,1493]" collectionCode="IRSNB" collectorName="O. Vishnu &amp; Coll. J. Thomson" country="India" location="India" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFE789179FCFFA0AFF1CFA5F" box="[206,229,1467,1493]" count="1" pageId="17" pageNumber="18" type="female">♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="BBC9AE01FFE789179DB6FA0AFCE8FA5F" box="[695,785,1467,1493]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="17" pageNumber="18">IRSNB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCF9B2FA8EF863" pageId="17" pageNumber="18" type="description">
<paragraph id="DD6736C4FFE789179FBCF9B2FEB7F997" blockId="17.[189,334,1539,1565]" box="[189,334,1539,1565]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179FBCF9B2FEB7F997" bold="true" box="[189,334,1539,1565]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="EFACEAD6FFE789179FBCF9B2FEB7F997" bold="true" box="[189,334,1539,1565]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCF988FCCBF9DE" blockId="17.[189,818,1593,1620]" box="[189,818,1593,1620]" pageId="17" pageNumber="18">MEASUREMENTS (in mm). TL=15.5–21; BW=9–10.5.</paragraph>
<paragraph id="DD6736C4FFE789179FEBF930FED3F911" blockId="17.[234,298,1665,1691]" box="[234,298,1665,1691]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBF930FED3F911" bold="true" box="[234,298,1665,1691]" pageId="17" pageNumber="18">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCF91AFBBFF8FF" blockId="17.[189,1399,1707,1910]" pageId="17" pageNumber="18">Black; clypeus rugulose, anterior margin strongly elevated; gently excised in the middle; frontal carina entire, curved, with a tubercle behind; pronotum strongly and closely punctured, rugose anteriorly; elytra finely striate, intervals angularly elevated along the middle, finely, fairly closely punctured; pygidium opaque, rugulose with short setae; metasternal shield smooth, shining, feebly, sparsely punctured, sides of metasternum rugosely punctured, with long hairs; front legs elongate with a blunt process on its trochanter; hind femur with a hooked process at the middle of its lower edge.</paragraph>
<paragraph id="DD6736C4FFE789179FEBF815FEBCF834" blockId="17.[234,325,1956,1982]" box="[234,325,1956,1982]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBF815FEBCF834" bold="true" box="[234,325,1956,1982]" pageId="17" pageNumber="18">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCF87FFA8EF863" blockId="17.[189,1399,1998,2025]" box="[189,1399,1998,2025]" pageId="17" pageNumber="18">Clypeal margin scarcely excised in the middle; protibia broad with four blunt teeth and an articulated spur.</paragraph>
</subSubSection>
<caption id="89A7664CFFE489149FBFF8D4FDB7F863" ID-DOI="http://doi.org/10.5281/zenodo.13742867" ID-Zenodo-Dep="13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="18" pageNumber="19" startId="18.[190,234,1893,1919]" targetBox="[295,1246,340,1832]" targetPageId="18">
<paragraph id="DD6736C4FFE489149FBFF8D4FDB7F863" blockId="18.[190,1407,1893,2025]" pageId="18" pageNumber="19">
<emphasis id="EFACEAD6FFE489149FBFF8D4FEEBF8F5" bold="true" box="[190,274,1893,1919]" pageId="18" pageNumber="19">Fig. 6.</emphasis>
Aedeagus, lateral, dorsal, ventral and apical views.
<emphasis id="EFACEAD6FFE489149CACF8D4FC1EF8F5" bold="true" box="[941,999,1893,1920]" pageId="18" pageNumber="19">A–D</emphasis>
.
<taxonomicName id="1AD84D47FFE489149CFAF8D7FA80F8F5" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[1019,1401,1893,1920]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFE489149CFAF8D7FB30F8F5" box="[1019,1225,1894,1920]" italics="true" pageId="18" pageNumber="19">Onitis excavatus</emphasis>
<bibRefCitation id="B9494B35FFE489149BD2F8D7FA80F8F5" author="Arrow G. J." box="[1235,1401,1893,1920]" pageId="18" pageNumber="19" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000037.
<emphasis id="EFACEAD6FFE489149ECEF838FDF3F829" bold="true" box="[463,522,1929,1955]" pageId="18" pageNumber="19">E–H</emphasis>
.
<taxonomicName id="1AD84D47FFE489149D19F838FCA7F828" authority="(Wulfen, 1786)" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[536,862,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFE489149D19F838FD64F828" box="[536,669,1928,1955]" italics="true" pageId="18" pageNumber="19">O. falcatus</emphasis>
(
<bibRefCitation id="B9494B35FFE489149DADF839FCACF828" author="Wulfen X." box="[684,853,1928,1954]" pageId="18" pageNumber="19" refId="ref31404" refString="Wulfen X. 1786. Descriptiones quorumdam Capensium insectorum. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1786">Wulfen, 1786</bibRefCitation>
)
</taxonomicName>
, AIMB/Co/Sc1000070.
<emphasis id="EFACEAD6FFE489149B7DF838FB52F829" bold="true" box="[1148,1195,1929,1955]" pageId="18" pageNumber="19">I–L</emphasis>
.
<taxonomicName id="1AD84D47FFE489149BB8F838FA81F828" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1209,1400,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFE489149BB8F838FA81F828" box="[1209,1400,1928,1955]" italics="true" pageId="18" pageNumber="19">O. bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFE489149FBFF81DFED8F84C" box="[190,289,1964,1990]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, NIM/NBAIR/COL/ONIT/H-271023A.
<emphasis id="EFACEAD6FFE489149C08F81DFCBEF84C" bold="true" box="[777,839,1964,1990]" pageId="18" pageNumber="19">M–P</emphasis>
.
<taxonomicName id="1AD84D47FFE489149C5AF81DFAE8F84C" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[859,1297,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFE489149C5AF81DFBB2F84C" box="[859,1099,1964,1990]" italics="true" pageId="18" pageNumber="19">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFE489149B59F81DFAE8F84C" author="Janssens A." box="[1112,1297,1964,1990]" pageId="18" pageNumber="19" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
, AIMB/ Co/Sc1000492. (Images: AIMB).
</paragraph>
</caption>
<subSubSection id="95C2654FFFE589159FBCFEB9FB02FE12" pageId="19" pageNumber="20" type="distribution">
<paragraph id="DD6736C4FFE589159FBCFEB9FEAFFEA8" blockId="19.[189,342,264,290]" box="[189,342,264,290]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="EFACEAD6FFE589159FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFE86FB02FE12" blockId="19.[189,1398,311,408]" pageId="19" pageNumber="20">
<collectingCountry id="A5CF7654FFE589159FBCFE86FF07FEDB" box="[189,254,311,337]" name="India" pageId="19" pageNumber="20">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFE589159E15FE86FE1FFEDB" box="[276,486,311,337]" country="India" name="Madhya Pradesh" pageId="19" pageNumber="20">Madhya Pradesh</collectingRegion>
, Punjab,
<collectingRegion id="1F1CF826FFE589159D69FE86FD05FEDB" box="[616,764,311,337]" country="India" name="Uttarakhand" pageId="19" pageNumber="20">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFE589159C13FE89FC3BFEDB" box="[786,962,311,338]" country="India" name="Uttar Pradesh" pageId="19" pageNumber="20">Uttar Pradesh</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFE589159BB2FE86FB04FEDB" box="[1203,1277,311,337]" name="Nepal" pageId="19" pageNumber="20">Nepal</collectingCountry>
;
<collectingCountry id="A5CF7654FFE589159A12FE86FA8FFEDB" box="[1299,1398,311,337]" name="Pakistan" pageId="19" pageNumber="20">Pakistan</collectingCountry>
(
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;
<bibRefCitation id="B9494B35FFE589159E7DFEEBFD82FEFE" author="Bezdek A. &amp; Krell F. - T." box="[380,635,346,372]" pageId="19" pageNumber="20" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
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2012b;
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).
</paragraph>
</subSubSection>
</treatment>
</document>
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