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<document id="92756CB941D536285BAD39565AAA6D40" ID-CLB-Dataset="29502" ID-DOI="10.11646/zootaxa.4484.1.1" ID-GBIF-Dataset="74b777c9-eae0-4770-8c86-dcbb10fb06b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437848" ID-ZooBank="D66524CF-9C6D-4DF4-8CA2-B2C9708CF5FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538125412450" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P. &amp; Melnik, Viacheslav P." docDate="2018" docId="521587E45600553109EEFA53884FFB88" docLanguage="en" docName="zootaxa.4484.1.1.pdf" docOrigin="Zootaxa 4484 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Calyssopora clarionensis Grischenko &amp; Gordon &amp; Melnik 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="AE2CFF9C563255070979FF888A4AFF87" masterDocTitle="Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining" masterLastPageNumber="91" masterPageNumber="1" pageNumber="51" updateTime="1698557051871" updateUser="ExternalLinkService">
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<mods:title id="CFDB5EBB63FF27AE000AD5173286A8F6">Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining</mods:title>
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<mods:namePart id="5F4D9E438C5CBDE2A815FB1B3B823341">Grischenko, Andrei V.</mods:namePart>
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<mods:namePart id="B5AC04E8F3E30BA431714148FE9C59B3">Gordon, Dennis P.</mods:namePart>
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<mods:namePart id="94BA1C7D574A4E50F1562AE6316A6A64">Melnik, Viacheslav P.</mods:namePart>
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<treatment id="521587E45600553109EEFA53884FFB88" ID-DOI="http://doi.org/10.5281/zenodo.5989823" ID-GBIF-Taxon="148403739" ID-Zenodo-Dep="5989823" LSID="urn:lsid:plazi:treatment:521587E45600553109EEFA53884FFB88" httpUri="http://treatment.plazi.org/id/521587E45600553109EEFA53884FFB88" lastPageId="54" lastPageNumber="55" pageId="50" pageNumber="51">
<subSubSection id="92A665795600553509EEFA538B6DF990" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="DA0336F25600553509EEFA538845FA72" blockId="50.[151,527,1499,1559]" box="[151,527,1499,1525]" pageId="50" pageNumber="51">
<heading id="814B819E5600553509EEFA538845FA72" bold="true" box="[151,527,1499,1525]" fontSize="11" level="1" pageId="50" pageNumber="51" reason="1">
<emphasis id="E8C8EAE05600553509EEFA538845FA72" bold="true" box="[151,527,1499,1525]" pageId="50" pageNumber="51">
<taxonomicName id="1DBC4D715600553509EEFA538B8FFA72" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[151,453,1499,1525]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="50" pageNumber="51" phylum="Bryozoa" rank="species" species="clarionensis" status="sp. nov.">
<emphasis id="E8C8EAE05600553509EEFA538B8FFA72" bold="true" box="[151,453,1499,1525]" italics="true" pageId="50" pageNumber="51">Calyssopora clarionensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B5600553508B5FA538845FA72" box="[460,527,1499,1525]" pageId="50" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DA0336F25600553509EEFA768B6DF990" blockId="50.[151,527,1499,1559]" box="[151,295,1534,1559]" pageId="50" pageNumber="51">
(
<figureCitation id="42872A775600553509E6FA768B55F990" box="[159,287,1534,1559]" captionStart-0="FIGURE 33" captionStart-1="FIGURE 34" captionStart-2="FIGURE 35" captionStartId-0="51.[151,250,1901,1923]" captionStartId-1="52.[151,250,1815,1837]" captionStartId-2="53.[151,250,1910,1932]" captionTargetBox-0="[162,1427,193,1885]" captionTargetBox-1="[247,1349,193,1803]" captionTargetBox-2="[163,1428,193,1896]" captionTargetId-0="figure@51.[162,1427,193,1885]" captionTargetId-1="figure@52.[247,1349,193,1803]" captionTargetId-2="figure@53.[163,1428,193,1896]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionText-0="FIGURE 33. Calyssopora clarionensis n. gen., n. sp. General view of holotype and paratype colonies. A, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome; B, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome; C, paratype 2, ZIRAS 3/50721, with ooeciopore visible; D, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm." captionText-1="FIGURE 34. Calyssopora clarionensis n. gen., n. sp. A, D, G, J, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome; B, E, H, K, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome; C, F, I, L, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome; M, specimen YMG413, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view; N, specimen GLD411, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces; O, P, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: AF, 200 µm; GI, 100 µm; JP, 50 µm." captionText-2="FIGURE 35. Calyssopora clarionensis n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. AD, specimen YMG1801, Stn 24, three-zooid colony; EH, specimen GLD408, Stn 144, fourfive-zooid stage; IL, specimen YMG407, Stn 134, seven-zooid stage with central cavity presumably representing incipient incubation chamber; MP, specimen GLD411, Stn 209, eight-zooid stage with paired central cavities. Scale bars: 200 µm." httpUri-0="https://zenodo.org/record/1437915/files/figure.png" httpUri-1="https://zenodo.org/record/1437917/files/figure.png" httpUri-2="https://zenodo.org/record/1437919/files/figure.png" pageId="50" pageNumber="51">Figs 3335</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="92A665795600553509EEF9CE8E35F8DC" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="DA0336F25600553509EEF9CE8E35F8DC" blockId="50.[151,1437,1606,2027]" pageId="50" pageNumber="51">
<emphasis id="E8C8EAE05600553509EEF9CE8BCBF9D8" bold="true" box="[151,385,1606,1631]" pageId="50" pageNumber="51">Material examined.</emphasis>
<emphasis id="E8C8EAE05600553508F0F9CE8BA5F9D8" box="[393,495,1606,1631]" italics="true" pageId="50" pageNumber="51">Holotype</emphasis>
: ZIRAS 1/50719, colony attached to nodule particle, YMG R.V.
<emphasis id="E8C8EAE0560055350DAAF9CE8F1BF9D8" box="[1235,1361,1606,1631]" italics="true" pageId="50" pageNumber="51">Gelendzhik</emphasis>
cruise GLD411, Stn 220,
<date id="AE0210325600553508FBF9E38862F904" box="[386,552,1642,1667]" pageId="50" pageNumber="51" value="2012-04-30">30 April 2012</date>
, 12.88593° N, 133.18168° W,
<quantity id="1D449B17560055350AF6F9E289A0F905" box="[911,1002,1642,1667]" metricMagnitude="3" metricUnit="m" metricValue="4.916" pageId="50" pageNumber="51" unit="m" value="4916.0">4916 m</quantity>
.
<emphasis id="E8C8EAE0560055350A8EF9E38E17F905" box="[1015,1117,1643,1666]" italics="true" pageId="50" pageNumber="51">Paratype</emphasis>
1: ZIRAS 2/50720, colony attached to nodule particle, YMG R.V.
<emphasis id="E8C8EAE0560055350B2DF9068968F920" box="[596,802,1678,1703]" italics="true" pageId="50" pageNumber="51">Yuzhmorgeologiya</emphasis>
cruise YMG413, Stn 291,
<date id="AE021032560055350D1AF9078EB3F920" box="[1123,1273,1678,1703]" pageId="50" pageNumber="51" value="2014-07-27">27 July 2014</date>
, 13.00083° N, 132.49723° W,
<quantity id="1D449B17560055350833F93A8BEEF94D" box="[330,420,1714,1739]" metricMagnitude="3" metricUnit="m" metricValue="4.865" pageId="50" pageNumber="51" unit="m" value="4865.0">4865 m</quantity>
.
<emphasis id="E8C8EAE05600553508C8F93B8852F94D" box="[433,536,1715,1738]" italics="true" pageId="50" pageNumber="51">Paratype</emphasis>
2: ZIRAS 3/50721, colony attached to nodule particle, YMG R.V.
<emphasis id="E8C8EAE0560055350C67F93A8FD6F94C" box="[1310,1436,1714,1739]" italics="true" pageId="50" pageNumber="51">Gelendzhik</emphasis>
cruise GLD409, Stn 186,
<date id="AE0210325600553508B6F95F88CBF968" box="[463,641,1751,1775]" pageId="50" pageNumber="51" value="2011-01-01">1 January 2011</date>
, 13.45140° N, 133.53807° W,
<quantity id="1D449B17560055350A9EF95F8E0BF969" box="[999,1089,1751,1775]" metricMagnitude="3" metricUnit="m" metricValue="4.872" pageId="50" pageNumber="51" unit="m" value="4872.0">4872 m</quantity>
.
<emphasis id="E8C8EAE0560055350D36F95F8EFCF969" box="[1103,1206,1751,1774]" italics="true" pageId="50" pageNumber="51">Paratype</emphasis>
3: ZIRAS 4/50722, detached colony, YMG R.V.
<emphasis id="E8C8EAE056005535089FF972882EF894" box="[486,612,1786,1811]" italics="true" pageId="50" pageNumber="51">Gelendzhik</emphasis>
cruise GLD412, Stn 246,
<date id="AE021032560055350ADEF9738E0AF894" box="[935,1088,1786,1811]" pageId="50" pageNumber="51" value="2013-04-03">3 April 2013</date>
, 13.00578° N, 132.95280° W,
<quantity id="1D449B175600553509EEF8968ABAF8B1" box="[151,240,1822,1847]" metricMagnitude="3" metricUnit="m" metricValue="4.935" pageId="50" pageNumber="51" unit="m" value="4935.0">4935 m</quantity>
.
<emphasis id="E8C8EAE0560055350985F8968B90F8B0" box="[252,474,1822,1847]" italics="true" pageId="50" pageNumber="51">Additional material</emphasis>
: YMG1801, Stn 24; YMG407, Stn 134; GLD408, Stns 144, 150; GLD411, Stn 209; YMG413, Stns 287, 290; YMG414, Stns 327, 339. Total specimens examined 13.
</paragraph>
</subSubSection>
<subSubSection id="92A665795600553509BEF8EE8FD7F8F9" box="[199,1437,1894,1919]" pageId="50" pageNumber="51" type="etymology">
<paragraph id="DA0336F25600553509BEF8EE8FD7F8F9" blockId="50.[151,1437,1606,2027]" box="[199,1437,1894,1919]" pageId="50" pageNumber="51">
<emphasis id="E8C8EAE05600553509BEF8EE8B04F8F8" bold="true" box="[199,334,1894,1919]" pageId="50" pageNumber="51">Etymology.</emphasis>
Pertaining to the Clarion Fracture, which defines the northern margin of the potential mining area.
</paragraph>
</subSubSection>
<subSubSection id="92A665795600553109BEF8028E0EFC07" lastPageId="54" lastPageNumber="55" pageId="50" pageNumber="51" type="description">
<paragraph id="DA0336F25600553509BEF8028F04F86D" blockId="50.[151,1437,1606,2027]" pageId="50" pageNumber="51">
<emphasis id="E8C8EAE05600553509BEF8028B13F824" bold="true" box="[199,345,1930,1955]" pageId="50" pageNumber="51">Description.</emphasis>
Colony erect, compact (
<figureCitation id="42872A77560055350B12F80288F7F824" box="[619,701,1930,1955]" captionStart="FIGURE 33" captionStartId="51.[151,250,1901,1923]" captionTargetBox="[162,1427,193,1885]" captionTargetId="figure@51.[162,1427,193,1885]" captionTargetPageId="51" captionText="FIGURE 33. Calyssopora clarionensis n. gen., n. sp. General view of holotype and paratype colonies. A, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome; B, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome; C, paratype 2, ZIRAS 3/50721, with ooeciopore visible; D, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437915/files/figure.png" pageId="50" pageNumber="51">Fig. 33</figureCitation>
) relative to other species, with apron-like base, vasiform; column topped by capitulum comprising peripheral autozooidal peristomes surrounding central concavity. Skeletal surface irregularly textured with minute dimples and granules except for base of column. Pores tiny, simple, sparse.
</paragraph>
<caption id="8EC3667A5601553409EEF8E58B94F858" httpUri="https://zenodo.org/record/1437915/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,250,1901,1923]" targetBox="[162,1427,193,1885]" targetPageId="51">
<paragraph id="DA0336F25601553409EEF8E58B94F858" blockId="51.[151,1436,1901,2015]" pageId="51" pageNumber="52">
<emphasis id="E8C8EAE05601553409EEF8E58B6AF804" bold="true" box="[151,288,1901,1923]" pageId="51" pageNumber="52">FIGURE 33.</emphasis>
<taxonomicName id="1DBC4D7156015534085FF8E58862F804" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[294,552,1901,1923]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="51" pageNumber="52" phylum="Bryozoa" rank="species" species="clarionensis" status="gen. nov.">
<emphasis id="E8C8EAE056015534085FF8E58862F804" box="[294,552,1901,1923]" italics="true" pageId="51" pageNumber="52">Calyssopora clarionensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B560155340B57F8E588F1F804" box="[558,699,1901,1923]" pageId="51" pageNumber="52" rank="species">
<emphasis id="E8C8EAE0560155340B57F8E5883CF804" bold="true" box="[558,630,1901,1923]" pageId="51" pageNumber="52">n. gen.</emphasis>
,
<emphasis id="E8C8EAE0560155340BFBF8E588F1F804" bold="true" box="[642,699,1901,1923]" pageId="51" pageNumber="52">n. sp.</emphasis>
</taxonomicNameLabel>
General view of holotype and paratype colonies.
<emphasis id="E8C8EAE0560155340DCDF8E58E8CF804" bold="true" box="[1204,1222,1901,1923]" pageId="51" pageNumber="52">A</emphasis>
, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome;
<emphasis id="E8C8EAE0560155340B51F8038873F826" bold="true" box="[552,569,1931,1953]" pageId="51" pageNumber="52">B</emphasis>
, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome;
<emphasis id="E8C8EAE0560155340DC7F8038E9AF826" bold="true" box="[1214,1232,1931,1953]" pageId="51" pageNumber="52">C</emphasis>
, paratype 2, ZIRAS 3/50721, with ooeciopore visible;
<emphasis id="E8C8EAE0560155340885F8228844F847" bold="true" box="[508,526,1962,1984]" pageId="51" pageNumber="52">D</emphasis>
, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm.
</paragraph>
</caption>
<caption id="8EC3667A5606553309EEF89F8F0AF862" httpUri="https://zenodo.org/record/1437917/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1815,1837]" targetBox="[247,1349,193,1803]" targetPageId="52">
<paragraph id="DA0336F25606553309EEF89F8F0AF862" blockId="52.[151,1436,1815,2021]" pageId="52" pageNumber="53">
<emphasis id="E8C8EAE05606553309EEF89F8B6BF8AA" bold="true" box="[151,289,1815,1837]" pageId="52" pageNumber="53">FIGURE 34.</emphasis>
<taxonomicName id="1DBC4D71560655330851F89F8866F8AA" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[296,556,1815,1837]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="52" pageNumber="53" phylum="Bryozoa" rank="species" species="clarionensis" status="gen. nov.">
<emphasis id="E8C8EAE0560655330851F89F8866F8AA" box="[296,556,1815,1837]" italics="true" pageId="52" pageNumber="53">Calyssopora clarionensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B560655330B4DF89F888FF8AA" box="[564,709,1815,1837]" pageId="52" pageNumber="53" rank="species">
<emphasis id="E8C8EAE0560655330B4DF89F8837F8AA" bold="true" box="[564,637,1815,1837]" pageId="52" pageNumber="53">n. gen.</emphasis>
,
<emphasis id="E8C8EAE0560655330BF2F89F888FF8AA" bold="true" box="[651,709,1815,1837]" pageId="52" pageNumber="53">n. sp.</emphasis>
</taxonomicNameLabel>
<emphasis id="E8C8EAE0560655330BB4F89F8895F8AA" bold="true" box="[717,735,1815,1837]" pageId="52" pageNumber="53">A</emphasis>
,
<emphasis id="E8C8EAE0560655330B94F89F88B5F8AA" bold="true" box="[749,767,1815,1837]" pageId="52" pageNumber="53">D</emphasis>
,
<emphasis id="E8C8EAE0560655330A74F89F8954F8AA" bold="true" box="[781,798,1815,1837]" pageId="52" pageNumber="53">G</emphasis>
,
<emphasis id="E8C8EAE0560655330A55F89F8972F8AA" bold="true" box="[812,824,1815,1837]" pageId="52" pageNumber="53">J</emphasis>
, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome;
<emphasis id="E8C8EAE0560655330A1AF8BD893EF8CC" bold="true" box="[867,884,1845,1867]" pageId="52" pageNumber="53">B</emphasis>
,
<emphasis id="E8C8EAE0560655330AFDF8BD89DFF8CC" bold="true" box="[900,917,1845,1867]" pageId="52" pageNumber="53">E</emphasis>
,
<emphasis id="E8C8EAE0560655330ADCF8BD89F3F8CC" bold="true" box="[933,953,1845,1867]" pageId="52" pageNumber="53">H</emphasis>
,
<emphasis id="E8C8EAE0560655330AB0F8BD8997F8CC" bold="true" box="[969,989,1845,1867]" pageId="52" pageNumber="53">K</emphasis>
, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome;
<emphasis id="E8C8EAE0560655330D6BF8DC8E6EF8ED" bold="true" box="[1042,1060,1876,1898]" pageId="52" pageNumber="53">C</emphasis>
,
<emphasis id="E8C8EAE0560655330D48F8DC8E74F8ED" bold="true" box="[1073,1086,1876,1898]" pageId="52" pageNumber="53">F</emphasis>
,
<emphasis id="E8C8EAE0560655330D33F8DC8E1EF8ED" bold="true" box="[1098,1108,1876,1898]" pageId="52" pageNumber="53">I</emphasis>
,
<emphasis id="E8C8EAE0560655330D19F8DC8E3BF8ED" bold="true" box="[1120,1137,1876,1898]" pageId="52" pageNumber="53">L</emphasis>
, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome;
<emphasis id="E8C8EAE0560655330B97F8FB894CF80E" bold="true" box="[750,774,1907,1929]" pageId="52" pageNumber="53">M</emphasis>
, specimen YMG413, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view;
<emphasis id="E8C8EAE0560655330B39F81A8818F82F" bold="true" box="[576,594,1938,1960]" pageId="52" pageNumber="53">N</emphasis>
, specimen GLD411, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces;
<emphasis id="E8C8EAE0560655330A7EF8388951F841" bold="true" box="[775,795,1968,1990]" pageId="52" pageNumber="53">O</emphasis>
,
<emphasis id="E8C8EAE0560655330A5EF838897EF841" bold="true" box="[807,820,1968,1990]" pageId="52" pageNumber="53">P</emphasis>
, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: AF, 200 µm; GI, 100 µm; JP, 50 µm.
</paragraph>
</caption>
<caption id="8EC3667A5607553209EEF8FE8EE9F875" httpUri="https://zenodo.org/record/1437919/files/figure.png" pageId="53" pageNumber="54" startId="53.[151,250,1910,1932]" targetBox="[163,1428,193,1896]" targetPageId="53">
<paragraph id="DA0336F25607553209EEF8FE8EE9F875" blockId="53.[151,1437,1910,2034]" pageId="53" pageNumber="54">
<emphasis id="E8C8EAE05607553209EEF8FE8B6BF80B" bold="true" box="[151,289,1910,1932]" pageId="53" pageNumber="54">FIGURE 35.</emphasis>
<taxonomicName id="1DBC4D71560755320850F8FE8867F80B" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[297,557,1910,1932]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="species" species="clarionensis" status="gen. nov.">
<emphasis id="E8C8EAE0560755320850F8FE8867F80B" box="[297,557,1910,1932]" italics="true" pageId="53" pageNumber="54">Calyssopora clarionensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B560755320B4DF8FE888CF80B" box="[564,710,1910,1932]" pageId="53" pageNumber="54" rank="species">
<emphasis id="E8C8EAE0560755320B4DF8FE8837F80B" bold="true" box="[564,637,1910,1932]" pageId="53" pageNumber="54">n. gen.</emphasis>
,
<emphasis id="E8C8EAE0560755320BF2F8FE888CF80B" bold="true" box="[651,710,1910,1932]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
Progressive stages of development of ancestrular and young colonies.
<emphasis id="E8C8EAE05607553209EEF81F8A82F82A" bold="true" box="[151,200,1943,1965]" pageId="53" pageNumber="54">AD</emphasis>
, specimen YMG1801, Stn 24, three-zooid colony;
<emphasis id="E8C8EAE0560755320B80F81F8960F82A" bold="true" box="[761,810,1943,1965]" pageId="53" pageNumber="54">EH</emphasis>
, specimen GLD408, Stn 144, fourfive-zooid stage;
<emphasis id="E8C8EAE0560755320C17F81F8FDFF82A" bold="true" box="[1390,1429,1943,1965]" pageId="53" pageNumber="54">IL</emphasis>
, specimen YMG407, Stn 134, seven-zooid stage with central cavity presumably representing incipient incubation chamber;
<emphasis id="E8C8EAE05607553209EEF8548A82F875" bold="true" box="[151,200,2012,2034]" pageId="53" pageNumber="54">MP</emphasis>
, specimen GLD411, Stn 209, eight-zooid stage with paired central cavities. Scale bars: 200 µm.
</paragraph>
</caption>
<paragraph id="DA0336F25604553109BEFF1F887EFEE3" blockId="54.[151,1437,151,1040]" pageId="54" pageNumber="55">
Autozooids in well-developed infertile colonies in single whorl of 67 (
<figureCitation id="42872A77560455310A8FFF1F8E39FF37" box="[1014,1139,151,176]" captionStart="FIGURE 35" captionStartId="53.[151,250,1910,1932]" captionTargetBox="[163,1428,193,1896]" captionTargetId="figure@53.[163,1428,193,1896]" captionTargetPageId="53" captionText="FIGURE 35. Calyssopora clarionensis n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. AD, specimen YMG1801, Stn 24, three-zooid colony; EH, specimen GLD408, Stn 144, fourfive-zooid stage; IL, specimen YMG407, Stn 134, seven-zooid stage with central cavity presumably representing incipient incubation chamber; MP, specimen GLD411, Stn 209, eight-zooid stage with paired central cavities. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437919/files/figure.png" pageId="54" pageNumber="55">
Fig.
<date id="AE021032560455310D55FF1F8E18FF37" box="[1068,1106,151,176]" pageId="54" pageNumber="55" value="1935-01">35I</date>
P
</figureCitation>
), with incipient zooid bud developing frontally at lower level between each pair; apertures bilaterally symmetrical, with raised, granular, arcuate lateral rims and concave frontal and abfrontal rims (
<figureCitation id="42872A77560455310A3DFF5789FBFF7F" box="[836,945,223,248]" captionStart="FIGURE 33" captionStartId="51.[151,250,1901,1923]" captionTargetBox="[162,1427,193,1885]" captionTargetId="figure@51.[162,1427,193,1885]" captionTargetPageId="51" captionText="FIGURE 33. Calyssopora clarionensis n. gen., n. sp. General view of holotype and paratype colonies. A, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome; B, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome; C, paratype 2, ZIRAS 3/50721, with ooeciopore visible; D, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437915/files/figure.png" pageId="54" pageNumber="55">Figs 33A</figureCitation>
,
<figureCitation id="42872A77560455310AC6FF688E7CFF7F" box="[959,1078,224,248]" captionStart="FIGURE 34" captionStartId="52.[151,250,1815,1837]" captionTargetBox="[247,1349,193,1803]" captionTargetId="figure@52.[247,1349,193,1803]" captionTargetPageId="52" captionText="FIGURE 34. Calyssopora clarionensis n. gen., n. sp. A, D, G, J, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome; B, E, H, K, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome; C, F, I, L, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome; M, specimen YMG413, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view; N, specimen GLD411, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces; O, P, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: AF, 200 µm; GI, 100 µm; JP, 50 µm." httpUri="https://zenodo.org/record/1437917/files/figure.png" pageId="54" pageNumber="55">34AF, M</figureCitation>
), hence somewhat mitriform in profile. Wall interiors densely covered by prickle-like granules (
<figureCitation id="42872A77560455310A18FE8C89ACFE9B" box="[865,998,260,285]" captionStart="FIGURE 34" captionStartId="52.[151,250,1815,1837]" captionTargetBox="[247,1349,193,1803]" captionTargetId="figure@52.[247,1349,193,1803]" captionTargetPageId="52" captionText="FIGURE 34. Calyssopora clarionensis n. gen., n. sp. A, D, G, J, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome; B, E, H, K, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome; C, F, I, L, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome; M, specimen YMG413, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view; N, specimen GLD411, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces; O, P, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: AF, 200 µm; GI, 100 µm; JP, 50 µm." httpUri="https://zenodo.org/record/1437917/files/figure.png" pageId="54" pageNumber="55">Fig. 34N, P</figureCitation>
). Alveoli rare between peristome bases abfrontally. Autozooids in mature brooding colonies with
<emphasis id="E8C8EAE0560455310A67FEA18960FEC7" box="[798,810,297,320]" italics="true" pageId="54" pageNumber="55">c</emphasis>
. 8 peristomes in primary whorl and
<emphasis id="E8C8EAE0560455310DB9FEA18E86FEC7" box="[1216,1228,297,320]" italics="true" pageId="54" pageNumber="55">c</emphasis>
. 7 slightly smaller peristomes at lower level (
<figureCitation id="42872A775604553108C6FEC4886DFEE3" box="[447,551,332,357]" captionStart="FIGURE 34" captionStartId="52.[151,250,1815,1837]" captionTargetBox="[247,1349,193,1803]" captionTargetId="figure@52.[247,1349,193,1803]" captionTargetPageId="52" captionText="FIGURE 34. Calyssopora clarionensis n. gen., n. sp. A, D, G, J, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome; B, E, H, K, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome; C, F, I, L, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome; M, specimen YMG413, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view; N, specimen GLD411, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces; O, P, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: AF, 200 µm; GI, 100 µm; JP, 50 µm." httpUri="https://zenodo.org/record/1437917/files/figure.png" pageId="54" pageNumber="55">Fig. 34A</figureCitation>
).
</paragraph>
<paragraph id="DA0336F25604553109BEFEE788BCFE73" blockId="54.[151,1437,151,1040]" pageId="54" pageNumber="55">
Gonozooid single, developing from female zooid in central concavity of capitulum, ventricose in vicinity of ooeciostome only, occupying part or all of concavity, ooeciostome strongly curved and hood-like (
<figureCitation id="42872A77560455310D94FE1C8F13FE2B" box="[1261,1369,404,429]" captionStart="FIGURE 33" captionStartId="51.[151,250,1901,1923]" captionTargetBox="[162,1427,193,1885]" captionTargetId="figure@51.[162,1427,193,1885]" captionTargetPageId="51" captionText="FIGURE 33. Calyssopora clarionensis n. gen., n. sp. General view of holotype and paratype colonies. A, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome; B, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome; C, paratype 2, ZIRAS 3/50721, with ooeciopore visible; D, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437915/files/figure.png" pageId="54" pageNumber="55">Figs 33A</figureCitation>
,
<figureCitation id="42872A77560455310C1CFE1C8B4CFE57" captionStart="FIGURE 34" captionStartId="52.[151,250,1815,1837]" captionTargetBox="[247,1349,193,1803]" captionTargetId="figure@52.[247,1349,193,1803]" captionTargetPageId="52" captionText="FIGURE 34. Calyssopora clarionensis n. gen., n. sp. A, D, G, J, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome; B, E, H, K, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome; C, F, I, L, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome; M, specimen YMG413, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view; N, specimen GLD411, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces; O, P, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: AF, 200 µm; GI, 100 µm; JP, 50 µm." httpUri="https://zenodo.org/record/1437917/files/figure.png" pageId="54" pageNumber="55">34A, B, G, H, J</figureCitation>
), its surface granular-tubercular, ooeciopore concealed from frontal view. Sparse, relatively large alveoli adjacent to gonozooid within concavity (
<figureCitation id="42872A77560455310B19FE5488A2FE73" box="[608,744,476,501]" captionStart="FIGURE 34" captionStartId="52.[151,250,1815,1837]" captionTargetBox="[247,1349,193,1803]" captionTargetId="figure@52.[247,1349,193,1803]" captionTargetPageId="52" captionText="FIGURE 34. Calyssopora clarionensis n. gen., n. sp. A, D, G, J, holotype, ZIRAS 1/50719, respectively showing an apical view of the fertile colony and profiles of the hooded ooeciostome; B, E, H, K, paratype 2, ZIRAS 3/50721, respectively showing an apical view of the gonozooid and profiles of a more-truncated ooeciostome; C, F, I, L, paratype 3, ZIRAS 4/50722, similar views of a colony with a more-open ooeciostome; M, specimen YMG413, Stn 287, mitriform shape of autozooecial orifice when seen in oblique lateral view; N, specimen GLD411, Stn 209, oblique view of autozooidal peristome and aperture showing spinose granulation of interior and exterior surfaces; O, P, paratype 1, ZIRAS 2/50720, respectively showing trabecula joining two peristomes and vertical view of autozooidal aperture. Scale bars: AF, 200 µm; GI, 100 µm; JP, 50 µm." httpUri="https://zenodo.org/record/1437917/files/figure.png" pageId="54" pageNumber="55">Fig. 34A, C</figureCitation>
).
</paragraph>
<paragraph id="DA0336F25604553109BEFE7788E8FD03" blockId="54.[151,1437,151,1040]" pageId="54" pageNumber="55">
Ancestrular zooid erect, not seen in isolation; earliest growth stage encountered comprising three very similar zooids (
<figureCitation id="42872A77560455310996FDAC8B3DFDBA" box="[239,375,548,573]" captionStart="FIGURE 35" captionStartId="53.[151,250,1910,1932]" captionTargetBox="[163,1428,193,1896]" captionTargetId="figure@53.[163,1428,193,1896]" captionTargetPageId="53" captionText="FIGURE 35. Calyssopora clarionensis n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. AD, specimen YMG1801, Stn 24, three-zooid colony; EH, specimen GLD408, Stn 144, fourfive-zooid stage; IL, specimen YMG407, Stn 134, seven-zooid stage with central cavity presumably representing incipient incubation chamber; MP, specimen GLD411, Stn 209, eight-zooid stage with paired central cavities. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437919/files/figure.png" pageId="54" pageNumber="55">Fig. 35AD</figureCitation>
) such that it is not clear which is ancestrular; protoecium not obvious. Elongation and outward curving of peristomes resulting in small vasiform colony (
<figureCitation id="42872A77560455310A48FDCF89FDFDE7" box="[817,951,583,608]" captionStart="FIGURE 35" captionStartId="53.[151,250,1910,1932]" captionTargetBox="[163,1428,193,1896]" captionTargetId="figure@53.[163,1428,193,1896]" captionTargetPageId="53" captionText="FIGURE 35. Calyssopora clarionensis n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. AD, specimen YMG1801, Stn 24, three-zooid colony; EH, specimen GLD408, Stn 144, fourfive-zooid stage; IL, specimen YMG407, Stn 134, seven-zooid stage with central cavity presumably representing incipient incubation chamber; MP, specimen GLD411, Stn 209, eight-zooid stage with paired central cavities. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437919/files/figure.png" pageId="54" pageNumber="55">Fig. 35FH</figureCitation>
) with budding of additional zooids lateroabfrontally between peristomes (
<figureCitation id="42872A77560455310B7FFDE488DFFD03" box="[518,661,620,645]" captionStart="FIGURE 35" captionStartId="53.[151,250,1910,1932]" captionTargetBox="[163,1428,193,1896]" captionTargetId="figure@53.[163,1428,193,1896]" captionTargetPageId="53" captionText="FIGURE 35. Calyssopora clarionensis n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. AD, specimen YMG1801, Stn 24, three-zooid colony; EH, specimen GLD408, Stn 144, fourfive-zooid stage; IL, specimen YMG407, Stn 134, seven-zooid stage with central cavity presumably representing incipient incubation chamber; MP, specimen GLD411, Stn 209, eight-zooid stage with paired central cavities. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437919/files/figure.png" pageId="54" pageNumber="55">Fig. 35M, N</figureCitation>
).
</paragraph>
<paragraph id="DA0336F25604553109BEFD078A97FCBF" blockId="54.[151,1437,151,1040]" pageId="54" pageNumber="55">
<emphasis id="E8C8EAE05604553109BEFD078B8FFD2F" bold="true" box="[199,453,655,680]" pageId="54" pageNumber="55">Measurements (mm).</emphasis>
<typeStatus id="050788505604553108B5FD07887DFD2F" box="[460,567,655,680]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
, ZIRAS 1/50719 (
<figureCitation id="42872A77560455310A7DFD078927FD2F" box="[772,877,655,680]" captionStart="FIGURE 33" captionStartId="51.[151,250,1901,1923]" captionTargetBox="[162,1427,193,1885]" captionTargetId="figure@51.[162,1427,193,1885]" captionTargetPageId="51" captionText="FIGURE 33. Calyssopora clarionensis n. gen., n. sp. General view of holotype and paratype colonies. A, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome; B, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome; C, paratype 2, ZIRAS 3/50721, with ooeciopore visible; D, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437915/files/figure.png" pageId="54" pageNumber="55">Fig. 33A</figureCitation>
): Colony height 1.94; capitulum 1.28 × 1.22; base at substratum 0.75 × 0.83; column diameter 0.470.76; ZL 0.6530.882 (0.784 ± 0.088) (
<emphasis id="E8C8EAE0560455310DFEFD3D8EDFFD4B" box="[1159,1173,693,716]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 8); PeL 0.2210.303 (0.255 ± 0.026) (
<emphasis id="E8C8EAE056045531082CFD518B29FD77" box="[341,355,729,752]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 8); PeD 0.2030.233 (0.218 ± 0.011) (
<emphasis id="E8C8EAE0560455310A49FD518974FD77" box="[816,830,729,752]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 8); ApL 0.1960.228 (0.211 ± 0.011) (
<emphasis id="E8C8EAE0560455310C74FD518F51FD77" box="[1293,1307,729,752]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 8); ApW 0.1550.192 (0.168 ± 0.013) (
<emphasis id="E8C8EAE0560455310896FD758BB7FC93" box="[495,509,765,788]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 8). Gonozooid (
<emphasis id="E8C8EAE0560455310BAAFD7588ABFC93" box="[723,737,765,788]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 1): GZL 0.483; GLW 0.491; OsL 0.348; OsW 0.226; OsH 0.207.
</paragraph>
<paragraph id="DA0336F25604553109BEFCCD8E08FCDB" blockId="54.[151,1437,151,1040]" box="[199,1090,836,861]" pageId="54" pageNumber="55">
Paratype 2, ZIRAS 3/50721 (
<figureCitation id="42872A77560455310B76FCCC883DFCDB" box="[527,631,836,861]" captionStart="FIGURE 33" captionStartId="51.[151,250,1901,1923]" captionTargetBox="[162,1427,193,1885]" captionTargetId="figure@51.[162,1427,193,1885]" captionTargetPageId="51" captionText="FIGURE 33. Calyssopora clarionensis n. gen., n. sp. General view of holotype and paratype colonies. A, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome; B, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome; C, paratype 2, ZIRAS 3/50721, with ooeciopore visible; D, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437915/files/figure.png" pageId="54" pageNumber="55">Fig. 33C</figureCitation>
): OpL 0.091 (
<emphasis id="E8C8EAE0560455310A6DFCCD8968FCDB" box="[788,802,837,860]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 1); OpW 0.108 (
<emphasis id="E8C8EAE0560455310A81FCCD8E4CFCDB" box="[1016,1030,837,860]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 1).
</paragraph>
<paragraph id="DA0336F25604553109BEFCE08E0EFC07" blockId="54.[151,1437,151,1040]" box="[199,1092,871,896]" pageId="54" pageNumber="55">
Paratype 3, ZIRAS 4/50722 (
<figureCitation id="42872A77560455310B76FCEF8832FC07" box="[527,632,871,896]" captionStart="FIGURE 33" captionStartId="51.[151,250,1901,1923]" captionTargetBox="[162,1427,193,1885]" captionTargetId="figure@51.[162,1427,193,1885]" captionTargetPageId="51" captionText="FIGURE 33. Calyssopora clarionensis n. gen., n. sp. General view of holotype and paratype colonies. A, holotype, ZIRAS 1/ 50719, with fully hooded ooeciostome; B, paratype 1, ZIRAS 2/50720, with mostly hooded ooecostome; C, paratype 2, ZIRAS 3/50721, with ooeciopore visible; D, paratype 3, ZIRAS 4/50722, with partially developed ooeciostome not yet concealing ooeciopore. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437915/files/figure.png" pageId="54" pageNumber="55">Fig. 33D</figureCitation>
): OpL 0.098 (
<emphasis id="E8C8EAE0560455310A6CFCE18969FC07" box="[789,803,873,896]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 1); OpW 0.124 (
<emphasis id="E8C8EAE0560455310A83FCE18E42FC07" box="[1018,1032,873,896]" italics="true" pageId="54" pageNumber="55">n</emphasis>
= 1).
</paragraph>
</subSubSection>
<subSubSection id="92A665795604553109BEFC048951FC4F" pageId="54" pageNumber="55" type="discussion">
<paragraph id="DA0336F25604553109BEFC048951FC4F" blockId="54.[151,1437,151,1040]" pageId="54" pageNumber="55">
<emphasis id="E8C8EAE05604553109BEFC048B71FC22" bold="true" box="[199,315,908,933]" pageId="54" pageNumber="55">Remarks.</emphasis>
<taxonomicName id="1DBC4D7156045531083BFC048815FC22" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[322,607,908,933]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="54" pageNumber="55" phylum="Bryozoa" rank="species" species="clarionensis" status="sp. nov.">
<emphasis id="E8C8EAE056045531083BFC048815FC22" box="[322,607,908,933]" italics="true" pageId="54" pageNumber="55">Calyssopora clarionensis</emphasis>
</taxonomicName>
<emphasis id="E8C8EAE0560455310B1FFC0488EEFC22" bold="true" box="[614,676,908,933]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="F3FB579B560455310B1FFC0488EEFC22" box="[614,676,908,933]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs most obviously from the
<typeStatus id="05078850560455310D6DFC058E0EFC22" box="[1044,1092,909,933]" pageId="54" pageNumber="55">type</typeStatus>
species in having a gonozooid with a curved ooeciostome and prickled zooidal interiors.
</paragraph>
</subSubSection>
<subSubSection id="92A665795604553109BEFC5C884FFB88" pageId="54" pageNumber="55" type="distribution">
<paragraph id="DA0336F25604553109BEFC5C884FFB88" blockId="54.[151,1437,151,1040]" pageId="54" pageNumber="55">
<emphasis id="E8C8EAE05604553109BEFC5C8B2AFC6A" bold="true" box="[199,352,980,1005]" pageId="54" pageNumber="55">Distribution.</emphasis>
Recorded from 13 stations within coordinates 12.84500
<geoCoordinate id="BF885035560455310A8CFC5C8ECDFC6B" box="[1013,1159,980,1005]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="14.33668">14.33668° N</geoCoordinate>
, 129.07338
<geoCoordinate id="BF885035560455310C65FC5C8AF8FB88" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="-134.46597">134.46597° W</geoCoordinate>
, at depth range
<quantity id="1D449B1756045531081BFC7F884AFB88" box="[354,512,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="4.956" metricValueMax="5.222" metricValueMin="4.6899999999999995" pageId="54" pageNumber="55" unit="m" value="4956.0" valueMax="5222.0" valueMin="4690.0">46905222 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>