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<document id="456A8C6E3126C30C7BC48EF44A80F3FC" ID-CLB-Dataset="124670" ID-DOI="10.11646/phytotaxa.558.2.4" ID-GBIF-Dataset="4f549d58-5299-4311-97d3-ec4b135b1d48" ID-ISSN="1179-3163" ID-Zenodo-Dep="7002961" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660733018858" checkinUser="plazi" docAuthor="Karaismailoğlu, Mehmet Cengiz, Güner, Özal &amp; Akçiçek, Ekrem" docDate="2022" docId="4B497F7CA702DB6EFF47FF4101EFFC8B" docLanguage="en" docName="phytotaxa.558.2.4.pdf" docOrigin="Phytotaxa 558 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.558.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Stachys L." docType="key" docVersion="7" lastPageNumber="215" masterDocId="B7700704A70EDB62FFCFFFDF0444FFDF" masterDocTitle="Characterization of the nutlet morpho-anatomical features of 12 Stachys taxa (Lamiaceae) from Turkey and its systematic practice" masterLastPageNumber="218" masterPageNumber="203" pageNumber="215" updateTime="1699367142723" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CE043011A19E17CB53A0F421C8C1A505">Characterization of the nutlet morpho-anatomical features of 12 Stachys taxa (Lamiaceae) from Turkey and its systematic practice</mods:title>
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<mods:namePart id="8BB9C18485093AA0F44E827FC158EBCF">Karaismailoğlu, Mehmet Cengiz</mods:namePart>
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<mods:affiliation id="DFA5DD4122FD554B3F79942A11033E2E">Department of Molecular Biology and Genetics, Faculty of Science, Bartın University, 74110, Bartın, Turkey biology _ 61 @ hotmail. com; https: // orcid. org / 0000 - 0002 - 6856 - 2742</mods:affiliation>
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<mods:namePart id="A07F4F9CA7BA8E0D085E5F381D6BD864">Güner, Özal</mods:namePart>
<mods:affiliation id="C5F58F0866FD2FD91964F21FF2827FC8">Harran District Directorate of National Education, Harran, Şanlıurfa, Turkey</mods:affiliation>
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<mods:namePart id="1C3FC30C3F59173A83B8DED0E06FA559">Akçiçek, Ekrem</mods:namePart>
<mods:affiliation id="94EDD8FC7658FDE891AA25F56E97C8DF">Department of Science Education, Faculty of Education, Dumlupınar University, Kütahya, Turkey</mods:affiliation>
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<mods:title id="341523176AD310BF5246A6BA13D3354B">Phytotaxa</mods:title>
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<mods:date id="7FF132C84E361DD04B35D77D6A9C2120">2022</mods:date>
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<treatment id="4B497F7CA702DB6EFF47FF4101EFFC8B" ID-DOI="http://doi.org/10.5281/zenodo.7010067" ID-GBIF-Taxon="199505155" ID-Zenodo-Dep="7010067" LSID="urn:lsid:plazi:treatment:4B497F7CA702DB6EFF47FF4101EFFC8B" httpUri="http://treatment.plazi.org/id/4B497F7CA702DB6EFF47FF4101EFFC8B" lastPageNumber="215" pageId="12" pageNumber="215">
<subSubSection id="8BFA9DE1A702DB6EFF47FF4100CFFF67" box="[136,1163,157,184]" pageId="12" pageNumber="215" type="nomenclature">
<paragraph id="C35FCE6AA702DB6EFF47FF4100CFFF67" blockId="12.[136,1163,157,184]" box="[136,1163,157,184]" pageId="12" pageNumber="215">
<heading id="98177906A702DB6EFF47FF4100CFFF67" bold="true" box="[136,1163,157,184]" fontSize="11" level="1" pageId="12" pageNumber="215" reason="1">
<emphasis id="F1941278A702DB6EFF47FF4100CFFF67" bold="true" box="[136,1163,157,184]" pageId="12" pageNumber="215">
Key to studied
<taxonomicName id="04E0B5E9A702DB6EFEF3FF4205D1FF67" ID-CoL="8W3RD" authority="L." authorityName="L." box="[316,405,157,184]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="genus">
<emphasis id="F1941278A702DB6EFEF3FF4205D1FF67" bold="true" box="[316,405,157,184]" italics="true" pageId="12" pageNumber="215">Stachys</emphasis>
</taxonomicName>
taxa, based on nutlet morphological and anatomical characters
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BFA9DE1A702DB6EFF47FF3D01EFFC8B" pageId="12" pageNumber="215" type="key">
<keyStep id="7814D6CFA702DB6EFF47FF3D01EFFECB" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FF3D01EFFF28" blockId="12.[136,1451,226,853]" box="[136,1451,226,248]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FF3D01EFFF28" box="[136,1451,226,248]" pageId="12" pageNumber="215">1. Nutlet color is brown, dark brown or dark brown-black ....................................................................................................................2</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FF2001EFFECB" blockId="12.[136,1451,226,853]" box="[136,1451,255,277]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FF2001EFFECB" box="[136,1451,255,277]" pageId="12" pageNumber="215">1. Nutlet color is taupe..........................................................................................................................................................................11</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FEC301EFFE91" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FEC301EFFEEE" blockId="12.[136,1451,226,853]" box="[136,1451,284,306]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FEC301EFFEEE" box="[136,1451,284,306]" pageId="12" pageNumber="215">2. Nutlet color is brown ..........................................................................................................................................................................3</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FEE601EFFE91" blockId="12.[136,1451,226,853]" box="[136,1451,313,335]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FEE601EFFE91" box="[136,1451,313,335]" pageId="12" pageNumber="215">2. Nutlet color is dark brown or dark brown-black ................................................................................................................................9</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FE8A01EEFE58" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FE8A01EFFEB5" blockId="12.[136,1451,226,853]" box="[136,1451,341,363]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FE8A01EFFEB5" box="[136,1451,341,363]" pageId="12" pageNumber="215">3. Nutlet shape is ovatus, ovatus-rectangularis or ovatus-triangularis ...................................................................................................4</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FEAD01EEFE58" blockId="12.[136,1451,226,853]" box="[136,1450,370,392]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FEAD01EEFE58" box="[136,1450,370,392]" pageId="12" pageNumber="215">
3. Nutlet shape is ellipticus............................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFB80FEAC01EEFE58" ID-CoL="7L3JG" box="[1103,1450,370,392]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="subSpecies" species="megalodonta" subSpecies="megalodonta">
<emphasis id="F1941278A702DB6EFB80FEAC00A0FE58" box="[1103,1252,370,392]" italics="true" pageId="12" pageNumber="215">S. megalodonta</emphasis>
subsp.
<emphasis id="F1941278A702DB6EFAE2FEAD01EEFE58" box="[1325,1450,370,391]" italics="true" pageId="12" pageNumber="215">megalodonta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FE5001EFFE1D" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FE5001EFFE7A" blockId="12.[136,1451,226,853]" box="[136,1451,399,421]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FE5001EFFE7A" box="[136,1451,399,421]" pageId="12" pageNumber="215">4. Ovatus-triangularis .............................................................................................................................................................................5</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FE7301EFFE1D" blockId="12.[136,1451,226,853]" box="[136,1451,428,450]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FE7301EFFE1D" box="[136,1451,428,450]" pageId="12" pageNumber="215">4. Ovatus, ovatus-rectangularis ..............................................................................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FE1701EEFE25" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FE1701EEFE01" blockId="12.[136,1451,226,853]" box="[136,1450,456,479]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FE1701EEFE01" box="[136,1450,456,479]" pageId="12" pageNumber="215">
5. Nutlet surface ornamentation
<typeStatus id="1C5B70C8A702DB6EFE3EFE16065EFE01" box="[497,538,457,478]" pageId="12" pageNumber="215">type</typeStatus>
is reticulate-foveate.....................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFA8AFE1601EEFE01" ID-CoL="4ZFYC" box="[1349,1450,457,478]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="sivasica">
<emphasis id="F1941278A702DB6EFA8AFE1601EEFE01" box="[1349,1450,457,478]" italics="true" pageId="12" pageNumber="215">S. sivasica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FE3A01EEFE25" blockId="12.[136,1451,226,853]" box="[136,1450,485,507]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FE3A01EEFE25" box="[136,1450,485,507]" pageId="12" pageNumber="215">
5. Nutlet surface ornamentation
<typeStatus id="1C5B70C8A702DB6EFE3EFE39065EFE24" box="[497,538,486,507]" pageId="12" pageNumber="215">type</typeStatus>
is verrucate ...........................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFAD6FE3901EEFE25" ID-CoL="4ZG2N" box="[1305,1450,485,507]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="tundjeliensis">
<emphasis id="F1941278A702DB6EFAD6FE3901EEFE25" box="[1305,1450,485,507]" italics="true" pageId="12" pageNumber="215">S. tundjeliensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FDDD01EFFDEB" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FDDD01EFFDC7" blockId="12.[136,1451,226,853]" box="[136,1451,514,536]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FDDD01EFFDC7" box="[136,1451,514,536]" pageId="12" pageNumber="215">
6. Nutlet surface ornamentation
<typeStatus id="1C5B70C8A702DB6EFE3EFDDC065EFDC7" box="[497,538,515,536]" pageId="12" pageNumber="215">type</typeStatus>
is reticulate or reticulate-areolate................................................................................................7
</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FDC001EFFDEB" blockId="12.[136,1451,226,853]" box="[136,1451,543,565]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FDC001EFFDEB" box="[136,1451,543,565]" pageId="12" pageNumber="215">
6. Nutlet surface ornamentation
<typeStatus id="1C5B70C8A702DB6EFE3EFDC0065EFDEB" box="[497,538,543,564]" pageId="12" pageNumber="215">type</typeStatus>
is verrucate or scalariform ..........................................................................................................8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FDE301EEFDB1" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FDE301EEFD8E" blockId="12.[136,1451,226,853]" box="[136,1450,572,594]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FDE301EEFD8E" box="[136,1450,572,594]" pageId="12" pageNumber="215">
7. Reticulate......................................................................................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFAD1FDE301EEFD8E" ID-CoL="4ZFQM" box="[1310,1450,572,593]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="mardinensis">
<emphasis id="F1941278A702DB6EFAD1FDE301EEFD8E" box="[1310,1450,572,593]" italics="true" pageId="12" pageNumber="215">S. mardinensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FD8601EEFDB1" blockId="12.[136,1451,226,853]" box="[136,1450,601,623]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FD8601EEFDB1" box="[136,1450,601,623]" pageId="12" pageNumber="215">
7. Reticulate-areolate...........................................................................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFAE1FD8601EEFDB1" box="[1326,1450,601,622]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="laetivirens">
<emphasis id="F1941278A702DB6EFAE1FD8601EEFDB1" box="[1326,1450,601,622]" italics="true" pageId="12" pageNumber="215">S. laetivirens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FDAA01EFFD78" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FDAA01EEFD55" blockId="12.[136,1451,226,853]" box="[136,1450,629,651]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FDAA01EEFD55" box="[136,1450,629,651]" pageId="12" pageNumber="215">
8. Scalariform .................................................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFB80FDA901EEFD55" ID-CoL="7L3JG" box="[1103,1450,629,651]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="subSpecies" species="megalodonta" subSpecies="megalodonta">
<emphasis id="F1941278A702DB6EFB80FDA900A0FD55" box="[1103,1252,629,651]" italics="true" pageId="12" pageNumber="215">S. megalodonta</emphasis>
subsp.
<emphasis id="F1941278A702DB6EFAE2FDAA01EEFD55" box="[1325,1450,629,650]" italics="true" pageId="12" pageNumber="215">megalodonta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FD4D01EFFD78" blockId="12.[136,1451,226,853]" box="[136,1451,658,680]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FD4D01EFFD78" box="[136,1451,658,680]" pageId="12" pageNumber="215">
8. Verrucate......................................................................................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFAD7FD4C01EFFD78" ID-CoL="4ZFCZ" box="[1304,1451,658,680]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="baytopiorum">
<emphasis id="F1941278A702DB6EFAD7FD4C01EFFD78" box="[1304,1451,658,680]" italics="true" pageId="12" pageNumber="215">S. baytopiorum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FD7001EFFD3E" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FD7001EEFD1B" blockId="12.[136,1451,226,853]" box="[136,1450,687,709]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FD7001EEFD1B" box="[136,1450,687,709]" pageId="12" pageNumber="215">
9. Nutlet shape is ellipticus-rectangularis...............................................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFA8EFD7001EEFD1B" ID-CoL="4ZFZ8" box="[1345,1450,687,708]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="subnuda">
<emphasis id="F1941278A702DB6EFA8EFD7001EEFD1B" box="[1345,1450,687,708]" italics="true" pageId="12" pageNumber="215">S. subnuda</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FD1301EFFD3E" blockId="12.[136,1451,226,853]" box="[136,1451,716,738]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FD1301EFFD3E" box="[136,1451,716,738]" pageId="12" pageNumber="215">9. Nutlet shape is ovatus or ovatus-rectangularis .................................................................................................................................10</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FD3601EFFCC4" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FD3601EFFD21" blockId="12.[136,1451,226,853]" box="[136,1451,745,767]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FD3601EFFD21" box="[136,1451,745,767]" pageId="12" pageNumber="215">
10. Nutlet surface ornamentation
<typeStatus id="1C5B70C8A702DB6EFE3EFD36065EFD21" box="[497,538,745,766]" pageId="12" pageNumber="215">type</typeStatus>
is alveolate .................................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFAF3FD3601EFFD21" ID-CoL="9V6QY" box="[1340,1451,745,766]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="siirtensis">
<emphasis id="F1941278A702DB6EFAF3FD3601EFFD21" box="[1340,1451,745,766]" italics="true" pageId="12" pageNumber="215">S. siirtensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FCDA01EFFCC4" blockId="12.[136,1451,226,853]" box="[136,1451,773,795]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FCDA01EFFCC4" box="[136,1451,773,795]" pageId="12" pageNumber="215">
10. Nutlet surface ornamentation
<typeStatus id="1C5B70C8A702DB6EFE3EFCD9065EFCC4" box="[497,538,774,795]" pageId="12" pageNumber="215">type</typeStatus>
is reticulate ................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFB19FCD901EFFCC4" ID-CoL="4ZG3G" authority="var. viscosa" box="[1238,1451,774,795]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="variety" species="viscosa" variety="viscosa">
<emphasis id="F1941278A702DB6EFB19FCD90171FCC4" box="[1238,1333,774,795]" italics="true" pageId="12" pageNumber="215">S. viscosa</emphasis>
var.
<emphasis id="F1941278A702DB6EFAABFCD901EFFCC4" box="[1380,1451,774,795]" italics="true" pageId="12" pageNumber="215">viscosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7814D6CFA702DB6EFF47FCFD01EFFC8B" pageId="12" pageNumber="215">
<paragraph id="C35FCE6AA702DB6EFF47FCFD01EEFCE8" blockId="12.[136,1451,226,853]" box="[136,1450,802,824]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FCFD01EEFCE8" box="[136,1450,802,824]" pageId="12" pageNumber="215">
11. Nutlet shape is ellipticus-rectangularis........................................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFADAFCFC01EEFCE8" ID-CoL="4ZFD3" box="[1301,1450,802,824]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="species" species="benthamiana">
<emphasis id="F1941278A702DB6EFADAFCFC01EEFCE8" box="[1301,1450,802,824]" italics="true" pageId="12" pageNumber="215">S. benthamiana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C35FCE6AA702DB6EFF47FCE001EFFC8B" blockId="12.[136,1451,226,853]" box="[136,1451,831,853]" pageId="12" pageNumber="215">
<keyLead id="78116D5FA702DB6EFF47FCE001EFFC8B" box="[136,1451,831,853]" pageId="12" pageNumber="215">
11. Nutlet shape is ovatus-rectangularis........................................................................................................
<taxonomicName id="04E0B5E9A702DB6EFB51FCE001EFFC8B" ID-CoL="4ZFSX" box="[1182,1451,831,853]" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="215" phylum="Tracheophyta" rank="variety" species="viscosa" variety="odontophylla">
<emphasis id="F1941278A702DB6EFB51FCE000B9FC8A" box="[1182,1277,831,853]" italics="true" pageId="12" pageNumber="215">S. viscosa</emphasis>
var.
<emphasis id="F1941278A702DB6EFAE3FCE001EFFC8B" box="[1324,1451,831,852]" italics="true" pageId="12" pageNumber="215">odontophylla</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>