347 lines
39 KiB
XML
347 lines
39 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e66321" ID-GBIF-Dataset="df495ca3-4f40-481d-b6e6-c1835d81b0a0" ID-GBIF-Taxon="189469083" ID-Pensoft-Pub="1864-8312-79-485" ID-Pensoft-UUID="12D8C5F9E3F15CBCA235F81E56FA0BC8" ID-ZooBank="3C3B731E89B44D1B9E0347BED83F8530" LSID="urn:lsid:zoobank.org:act:D2951940-FBC8-45DB-B91B-5B968A11B1B8" ModsDocID="1864-8312-79-485" checkinTime="1633780201340" checkinUser="pensoft" docAuthor="Saraiva, Nicolas Eugenio de Vasconcelos, Hara, Marcos Ryotaro & DaSilva, Marcio Bernardino" docDate="2021" docId="3590AB43E999560898415D2E3131545B" docLanguage="en" docName="ArthropodSystPhyl 79: 485-507" docOrigin="Arthropod Systematics & amp; Phylogeny 79" docPubDate="2021-10-08" docSource="http://dx.doi.org/10.3897/asp.79.e66321" docTitle="Sertaneja falcata Saraiva & Hara & DaSilva 2021, sp. nov." docType="treatment" docUuid="D2951940-FBC8-45DB-B91B-5B968A11B1B8" docUuidSource="ZooBank" docVersion="3" id="12D8C5F9E3F15CBCA235F81E56FA0BC8" lastPageNumber="485" masterDocId="12D8C5F9E3F15CBCA235F81E56FA0BC8" masterDocTitle="Harvestmen in the semiarid: a new genus and three new species of Pachylinae (Opiliones: Gonyleptidae) from Caatinga dry vegetation, with a cladistic analysis" masterLastPageNumber="507" masterPageNumber="485" pageId="0" pageNumber="485" updateTime="1635284737390" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Harvestmen in the semiarid: a new genus and three new species of Pachylinae (Opiliones: Gonyleptidae) from Caatinga dry vegetation, with a cladistic analysis</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Saraiva, Nicolas Eugenio de Vasconcelos</mods:namePart>
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<mods:affiliation>Universidade Federal da Paraiba, Centro de Ciencias Exatas e da Natureza, Departamento de Sistematica e Ecologia, Programa de Pos-Graduacao em Ciencias Biologicas (Zoologia), Joao Pessoa, PB, Brazil; Nicolas E. de V. Saraiva [nicools. eugenio @ gmail. com]; Marcio B. DaSilva [1940 @ uol. com. br]</mods:affiliation>
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<mods:nameIdentifier type="email">nicools.eugenio@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hara, Marcos Ryotaro</mods:namePart>
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<mods:affiliation>Universidade de Sao Paulo, Escola de Artes, Ciencias e Humanidades (EACH), Sao Paulo, SP, Brazil; Marcos R. Hara [marcosrh @ usp. br]</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DaSilva, Marcio Bernardino</mods:namePart>
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<mods:affiliation>Universidade Federal da Paraiba, Centro de Ciencias Exatas e da Natureza, Departamento de Sistematica e Ecologia, Programa de Pos-Graduacao em Ciencias Biologicas (Zoologia), Joao Pessoa, PB, Brazil; Nicolas E. de V. Saraiva [nicools. eugenio @ gmail. com]; Marcio B. DaSilva [1940 @ uol. com. br]</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-10-08</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>79</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>485</mods:start>
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<mods:end>507</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.79.e66321</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.79.e66321</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-79-485</mods:identifier>
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<mods:identifier type="ZooBank">3C3B731E89B44D1B9E0347BED83F8530</mods:identifier>
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<mods:identifier type="Pensoft-UUID">12D8C5F9E3F15CBCA235F81E56FA0BC8</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="485" type="3. results">
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<treatment ID-GBIF-Taxon="189469083" LSID="urn:lsid:zoobank.org:act:D2951940-FBC8-45DB-B91B-5B968A11B1B8" httpUri="http://treatment.plazi.org/id/3590AB43E999560898415D2E3131545B" lastPageNumber="485" pageId="0" pageNumber="485">
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<subSubSection pageId="0" pageNumber="485" type="nomenclature">
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<paragraph pageId="0" pageNumber="485">
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3.3.5.
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<taxonomicName LSID="http://zoobank.org/D2951940-FBC8-45DB-B91B-5B968A11B1B8" authority="Saraiva & Hara & DaSilva, 2021" authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Sertaneja falcata" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="falcata" status="sp. nov.">Sertaneja falcata</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="485">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="description">
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<paragraph pageId="0" pageNumber="485">
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">Figs 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">, 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Sertaneja falcata sp. nov. (UFPB OP- 728), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure11" httpUri="https://binary.pensoft.net/fig/596984" pageId="0" pageNumber="485">, 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Photographs of live Sertaneja gen. nov. A: Female of S. crassitibialis sp. nov. (courtesy of Leonardo Sousa Carvalho); B: Male of S. bicuspidata sp. nov.; C, E: Male of S. falcata sp. nov.; D: Female of S. falcata sp. nov." figureDoi="10.3897/asp.79.e66321.figure12" httpUri="https://binary.pensoft.net/fig/596985" pageId="0" pageNumber="485">, 12C-E</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Geographic distribution of Sertaneja gen. nov. and biomes. A: northeastern Brazil detail; B: distribution compared to Pachylinae groups." figureDoi="10.3897/asp.79.e66321.figure13" httpUri="https://binary.pensoft.net/fig/596986" pageId="0" pageNumber="485">, 13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="type locality">
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<paragraph pageId="0" pageNumber="485">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="485">BRAZIL, Rio Grande do Norte: Portalegre, Cachoeira do Pinga.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="type material">
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<paragraph pageId="0" pageNumber="485">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Holotype.</emphasis>
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BRAZIL. Rio Grande do Norte: Portalegre, Cachoeira do Pinga, encosta da serra, elev. 431 m, 23-24.iv.2014, DaSilva M.B. and Saraiva N.E.V., ♂ (UFPB OP-481).
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<emphasis bold="true" pageId="0" pageNumber="485">Paratypes.</emphasis>
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same as for holotype, 3 ♂ (UFPB OP-482), 1 ♂ (UFPB OP-728) [dissected], 2 ♀ (UFPB OP-729) [female description], 1 ♂ (MZUSP 76647), 1 ♂ (MNRJ 60617).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="diagnosis">
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<paragraph pageId="0" pageNumber="485">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="485">
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Distinguished from other species by the presence of a pair of slightly enlarged paramedian tubercles on scute area III (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9A, C</figureCitation>
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), instead of spines; coxa IV bearing a short, conical, straight prodorsal apical apophysis and a long, curved, sickle-like retro-apical apophysis (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9D</figureCitation>
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); femur IV retro-lateral face with three central elongated apophysis (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10A, D</figureCitation>
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); and contains five pairs of MS C (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Sertaneja falcata sp. nov. (UFPB OP- 728), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure11" httpUri="https://binary.pensoft.net/fig/596984" pageId="0" pageNumber="485">11</figureCitation>
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). Most similar to
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<taxonomicName genus="S." lsidName="S. bicuspidata" pageId="0" pageNumber="485" rank="species" species="bicuspidata">
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<emphasis italics="true" pageId="0" pageNumber="485">S. bicuspidata</emphasis>
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</taxonomicName>
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sp. nov. by the slightly elongated trochanter IV; penis
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<normalizedToken originalValue="glans’">glans'</normalizedToken>
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stylus with a rounded dorso-apical projection; similar MS A and MS B; and MS D pairs close MS C group.
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</paragraph>
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<caption doi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Figure 9.</emphasis>
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<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja falcata" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="falcata">
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<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja falcata</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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,
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<emphasis bold="true" pageId="0" pageNumber="485">C</emphasis>
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-
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<emphasis bold="true" pageId="0" pageNumber="485">G</emphasis>
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) Male holotype (UFPB OP-481),
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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: habitus, dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">C</emphasis>
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: same, right lateral view;
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<emphasis bold="true" pageId="0" pageNumber="485">D</emphasis>
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: right coxa and trochanter IV in dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">E</emphasis>
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: right pedipalp in lateral view;
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<emphasis bold="true" pageId="0" pageNumber="485">F</emphasis>
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: same, mesal view;
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<emphasis bold="true" pageId="0" pageNumber="485">G</emphasis>
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: ocularium frontal view. (
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<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
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) Female paratype (UFPB OP-729) habitus, dorsal view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485" start="Figure 10" startId="F10">
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Figure 10.</emphasis>
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<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja falcata" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="falcata">
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<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja falcata</emphasis>
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</taxonomicName>
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sp. nov. Male holotype (UFPB OP-481) right trochanter-tibia IV.
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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: dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
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: prolateral view;
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<emphasis bold="true" pageId="0" pageNumber="485">C</emphasis>
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: ventral view;
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<emphasis bold="true" pageId="0" pageNumber="485">D</emphasis>
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: retro-lateral view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.79.e66321.figure11" httpUri="https://binary.pensoft.net/fig/596984" pageId="0" pageNumber="485" start="Figure 11" startId="F11">
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Figure 11.</emphasis>
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<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja falcata" order="Opiliones" pageId="0" pageNumber="485" rank="species" species="falcata">
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<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja falcata</emphasis>
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</taxonomicName>
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sp. nov. (UFPB OP-728), apical portion of penis.
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<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
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: dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
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: right lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="485" type="description">
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<paragraph pageId="0" pageNumber="485">Description.</paragraph>
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<paragraph pageId="0" pageNumber="485">
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<emphasis bold="true" pageId="0" pageNumber="485">Holotype MALE</emphasis>
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(UFPB OP-481; Figs
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10</figureCitation>
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):
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<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
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: DSL: 4.43; DSW: 4.87; LI: 8.91; LII: 19.48; LIII: 12.24; LIV: 16.53.
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<emphasis bold="true" pageId="0" pageNumber="485">Dorsum</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9A, C, G</figureCitation>
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): Prosoma with few tubercles. Front margin bearing a single small median tubercle. Ocularium twice as tall as the eye diameter, with a pair of divergent spines slightly projected anteriorly (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9C, G</figureCitation>
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). Two pairs of ozopores of similar size, anterior one placed laterally, and posterior one rounded and more dorsal; lateral channel developed and followed by oblong sensorial pegs that increase in size posteriorly. Scute area I with tubercles placed in a row along the longitudinal groove and scute groove II, with a pair of slightly enlarged paramedian tubercles (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9A</figureCitation>
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). Scute areas II-IV each with a transversal central row of enlarged tubercles increasing in size towards the middle, paramedian pair larger than the others. Scute areas III-IV with more conical tubercles compared to those on other areas. Lateral margin of DS with an external row of low, rounded tubercles slightly increasing in size posteriorly. Posterior margin of DS and free tergites I-III each with a row of conical tubercles. Anal operculum tuberculate.
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<emphasis bold="true" pageId="0" pageNumber="485">Venter</emphasis>
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: Coxa I tuberculate, median row of large ones, apical face with two prolateral and two retro-lateral tubercles. Coxae II-III with small tubercles, pro- and retro-apical tubercles reduced. Coxa IV tuberculate.
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<emphasis bold="true" pageId="0" pageNumber="485">Chelicerae</emphasis>
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: Segment I with ectodorsal row of three tubercles on posterior face of bulla. Fixed and movable fingers with five and three teeth, respectively.
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<emphasis bold="true" pageId="0" pageNumber="485">Pedipalps</emphasis>
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(Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9E, F</figureCitation>
|
||
): Trochanter inflated dorsally, ventral face with two setiferous tubercles, mesal largest. Femur with a basal setiferous tubercle, an external lateral row of 2-3 smaller setiferous tubercles on the middle 1/3 (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9E</figureCitation>
|
||
), and a mesal apical seta. Tibial setation: mesal IiIi, lateral IiIii. Tarsal setation: mesal IIi, lateral IiIi/Iii.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10</figureCitation>
|
||
): Coxae I-III each with one prodorsal and one retro-dorsal apophysis; coxa II retro-dorsal apophysis trifid and fused at the apex with coxa III prodorsal apophysis. Coxa IV large, covered by more acuminate tubercles than those of the rest of the body (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9A</figureCitation>
|
||
); with an obliquely inserted, conical prodorsal apical apophysis that is slightly curved downwards (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9C, D</figureCitation>
|
||
); and a sickle-shaped retro-apical apophysis, twice the size of proapical apophysis, curved out and downwards (Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Sertaneja falcata sp. nov. (A, C - G) Male holotype (UFPB OP- 481), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 729) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure9" httpUri="https://binary.pensoft.net/fig/596982" pageId="0" pageNumber="485">9D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10D</figureCitation>
|
||
). Trochanter III with a reduced retro-apical projection. Trochanter IV long, with pro- and a retro-lateral central blunt apophyses, the prolateral one slightly acuminated (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10A</figureCitation>
|
||
); apical portion with a long, conical retro-apical apophysis and a small proapical round tubercle (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10</figureCitation>
|
||
). Femur III slightly curved retrolaterally in dorsal view, with a slightly acuminated retro-apical dorsal tubercle; ventral surface with two rows of tubercles increasing in size subapically, prolateral row larger. Femur IV slightly curved inwards at the base, medio-dorsal row of tubercles twice the height of those covering the podomere, almost on the entire femur (except the sub-apical surface), basal and sub-apical tubercles more conspicuous (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10A, B, D</figureCitation>
|
||
); a pair of long, curved dorso-apical paralateral spines, retro-lateral spine largest and strongly curved upwards (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10A, D</figureCitation>
|
||
); retro-lateral face with three long, conical, central apophyses (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10A, D</figureCitation>
|
||
); ventral surface with a prolateral row of large tubercles alternating with small ones on apical half, and a retro-lateral row with round tubercles (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10B, C</figureCitation>
|
||
); ventro-apical portion with two paralateral large straight spiniform apophyses, proapical largest (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10C</figureCitation>
|
||
). Patella IV tuberculate, ventral surface covered by enlarged, acuminated tubercles, with a large proapical spine (twice the size of the surrounding ones) (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10B</figureCitation>
|
||
). Tibia IV with two ventral rows of tubercles increasing in size apically, pointing posteriorly (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10B, D</figureCitation>
|
||
). Metatarsus I slightly swollen. Tarsal counts: 6, 9, 6, 6.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Penis</emphasis>
|
||
(UFPB OP-728; Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Sertaneja falcata sp. nov. (UFPB OP- 728), apical portion of penis. A: dorsal view; B: right lateral view." figureDoi="10.3897/asp.79.e66321.figure11" httpUri="https://binary.pensoft.net/fig/596984" pageId="0" pageNumber="485">11</figureCitation>
|
||
): VP apex and lateral margins slightly concave. MS A set showing asymmetry between right (4) and left (3) sides (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10A, B</figureCitation>
|
||
); MS B almost undistinguishable from MS A but inserted more basally. Five pairs of conical, curved MS C, decreasing in size apically; two pairs of MS D close to MS C5, but slightly smaller and placed more dorsally; two pairs of small MS E, ventral to the MS C set (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sertaneja falcata sp. nov. Male holotype (UFPB OP- 481) right trochanter-tibia IV. A: dorsal view; B: prolateral view; C: ventral view; D: retro-lateral view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure10" httpUri="https://binary.pensoft.net/fig/596983" pageId="0" pageNumber="485">10B</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Coloration</emphasis>
|
||
: General coloration predominantly moderate yellow (87). DS scute areas, prosoma and margins, free tergites I-III moderate olive brown (95) granulation.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<emphasis bold="true" pageId="0" pageNumber="485">FEMALE</emphasis>
|
||
(UFPB OP-729; Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sertaneja crassitibialis sp. nov. (A, C - G) Male holotype (UFPB OP- 487), A: habitus, dorsal view; C: same, right lateral view; D: right coxa and trochanter IV in dorsal view; E: right pedipalp in lateral view; F: same, mesal view; G: ocularium frontal view. (B) Female paratype (UFPB OP- 727) habitus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/asp.79.e66321.figure6" httpUri="https://binary.pensoft.net/fig/596979" pageId="0" pageNumber="485">6B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Photographs of live Sertaneja gen. nov. A: Female of S. crassitibialis sp. nov. (courtesy of Leonardo Sousa Carvalho); B: Male of S. bicuspidata sp. nov.; C, E: Male of S. falcata sp. nov.; D: Female of S. falcata sp. nov." figureDoi="10.3897/asp.79.e66321.figure12" httpUri="https://binary.pensoft.net/fig/596985" pageId="0" pageNumber="485">12D</figureCitation>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
|
||
: DSL: 4.13; DSW: 4.32; LI: 8.83; LII: 16.57; LIII: 10.93; LIV: 14.40.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Dorsum</emphasis>
|
||
: Less armed compared to male. DS narrower. Lateral margin of DS with an external row of tubercles, posterior tubercles slightly larger than in male.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Pedipalps</emphasis>
|
||
: Pedipalp tibial setation: lateral IiIi.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
: Coxa IV narrower and shorter than in male, with a small prolateral apophysis, and a conical, straight retro-apical apophysis of same size. Trochanter IV narrower and shorter than in male, the prolateral basal apophysis as a large, blunt tubercle, and a small retro-lateral apophysis. Femur III lacking a proventral row of enlarged tubercles, reduced retro-dorsal apical tubercle. Femur IV unarmed, covered by small conical tubercles that slightly increase in size apically, with a pair of small dorso-apical spines of similar size. Patella-tibia IV unarmed. Tarsal counts: 6, 9-10, 6, 6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="485" type="variation">
|
||
<paragraph pageId="0" pageNumber="485">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Males</emphasis>
|
||
(n = 6):
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
|
||
: DSL 4.43-4.66; DSW 4.84-5.06; LI 8.91-10.21; LII 17.52-19.48; LIII 12.24-13.08; LIV 16.19-17.51.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Dorsum</emphasis>
|
||
: Front margin with or without tubercles. Ocularium with parallel or diverging spines, inclination of these ranging from near-horizontal to vertical.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Pedipalps</emphasis>
|
||
: Femur external lateral row with 2-3 sub-apical tubercles.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
: Femur IV dorso-apical spines with similar size to retro-lateral largest and strongly curved upwards; retro-lateral central conical apophyses sometimes with an additional apophysis of half the height of the remaining apophyses in the middle; apical half of proventral row bearing small tubercles alternated with large ones. Tarsal counts: 6, 10-12, 6, 4-6.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Females</emphasis>
|
||
(n = 2):
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Measurements</emphasis>
|
||
: DSL 4.13-4.27; DSW 4.32-4.46; LI 8.83-8.84; LII 16.16-16.57; LIII 10.93-11.62; LIV 10.09.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Pedipalps</emphasis>
|
||
: Pedipalp tibial setation: mesal IiIi, lateral IiIi. Tarsal setation: mesal IIi, lateral IiIi/IiIii.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Legs</emphasis>
|
||
: Tarsal counts: 6, 9-10, 6, 6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="485" type="etymology">
|
||
<paragraph pageId="0" pageNumber="485">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<normalizedToken originalValue="‘Falcata’">'Falcata'</normalizedToken>
|
||
is a Latin adjective (adj. nom. fem.) that means sickle-shaped and refers to the diagnostic retro-apical apophysis of male coxa IV of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="485" type="distribution">
|
||
<paragraph pageId="0" pageNumber="485">
|
||
Distribution (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Geographic distribution of Sertaneja gen. nov. and biomes. A: northeastern Brazil detail; B: distribution compared to Pachylinae groups." figureDoi="10.3897/asp.79.e66321.figure13" httpUri="https://binary.pensoft.net/fig/596986" pageId="0" pageNumber="485">13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="485">Known only from type locality. Caatinga dry forest of tableland slopes of interior Rio Grande do Norte.</paragraph>
|
||
<caption doi="10.3897/asp.79.e66321.figure12" httpUri="https://binary.pensoft.net/fig/596985" pageId="0" pageNumber="485" start="Figure 12" startId="F12">
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Figure 12.</emphasis>
|
||
Photographs of live
|
||
<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja" order="Opiliones" pageId="0" pageNumber="485" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja</emphasis>
|
||
</taxonomicName>
|
||
gen. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
|
||
: Female of
|
||
<taxonomicName genus="S." lsidName="S. crassitibialis" pageId="0" pageNumber="485" rank="species" species="crassitibialis">
|
||
<emphasis italics="true" pageId="0" pageNumber="485">S. crassitibialis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (courtesy of Leonardo Sousa Carvalho);
|
||
<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
|
||
: Male of
|
||
<taxonomicName genus="S." lsidName="S. bicuspidata" pageId="0" pageNumber="485" rank="species" species="bicuspidata">
|
||
<emphasis italics="true" pageId="0" pageNumber="485">S. bicuspidata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.;
|
||
<emphasis bold="true" pageId="0" pageNumber="485">C</emphasis>
|
||
,
|
||
<emphasis bold="true" pageId="0" pageNumber="485">E</emphasis>
|
||
: Male of
|
||
<taxonomicName genus="S." lsidName="S. falcata" pageId="0" pageNumber="485" rank="species" species="falcata">
|
||
<emphasis italics="true" pageId="0" pageNumber="485">S. falcata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.;
|
||
<emphasis bold="true" pageId="0" pageNumber="485">D</emphasis>
|
||
: Female of
|
||
<taxonomicName genus="S." lsidName="S. falcata" pageId="0" pageNumber="485" rank="species" species="falcata">
|
||
<emphasis italics="true" pageId="0" pageNumber="485">S. falcata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/asp.79.e66321.figure13" httpUri="https://binary.pensoft.net/fig/596986" pageId="0" pageNumber="485" start="Figure 13" startId="F13">
|
||
<paragraph pageId="0" pageNumber="485">
|
||
<emphasis bold="true" pageId="0" pageNumber="485">Figure 13.</emphasis>
|
||
Geographic distribution of
|
||
<taxonomicName authorityName="Saraiva & Hara & DaSilva" authorityYear="2021" family="Gonyleptidae" genus="Sertaneja" kingdom="Animalia" lsidName="Sertaneja" order="Opiliones" pageId="0" pageNumber="485" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="485">Sertaneja</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. and biomes.
|
||
<emphasis bold="true" pageId="0" pageNumber="485">A</emphasis>
|
||
: northeastern Brazil detail;
|
||
<emphasis bold="true" pageId="0" pageNumber="485">B</emphasis>
|
||
: distribution compared to
|
||
<taxonomicName authorityName="Sorensen" authorityYear="1884" lsidName="" pageId="0" pageNumber="485" rank="subFamily" subFamily="Pachylinae">Pachylinae</taxonomicName>
|
||
groups.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |