133 lines
19 KiB
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133 lines
19 KiB
XML
<document id="F6F28F89F32ACF12C69F2087AED4318A" ID-DOI="10.5281/zenodo.5391729" ID-ISSN="1638-9387" ID-Zenodo-Dep="5391729" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630315468151" checkinUser="marcus" docAuthor="Wesener, Thomas & Wägele, Johann-Wolfgang" docDate="2008" docId="AA4514472B1DFFFCFEA9FF70FBB29C52" docLanguage="en" docName="Zoosystema.2008.30.1.5-85.pdf" docOrigin="Zoosystema 30 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Zoosphaerium libidinosum" docType="key" docVersion="4" lastPageNumber="22" masterDocId="567C6C3F2B0CFFEDFFCFFFA8FFDC9F45" masterDocTitle="The giant pill-millipedes of Madagascar: revision of the genus Zoosphaerium (Myriapoda, Diplopoda, Sphaerotheriida)" masterLastPageNumber="85" masterPageNumber="5" pageNumber="22" updateTime="1699223479737" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="DF833BC469C4943EA5E0E99BA895825E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="427D9C08FEE503FE10EA7B16521515B7">
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<mods:title id="5AE5D156F412457707085610C83D03A9">The giant pill-millipedes of Madagascar: revision of the genus Zoosphaerium (Myriapoda, Diplopoda, Sphaerotheriida)</mods:title>
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</mods:titleInfo>
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<mods:name id="63468FA2146A6EC56F48CAA8C144C0B0" type="personal">
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<mods:roleTerm id="F06666D2C26B4EB7841711B20D9F2900">Author</mods:roleTerm>
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<mods:namePart id="468851E13BBA2751E54025B005764CC3">Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:name id="1464213144A3FF890B5840B1845F8960" type="personal">
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<mods:role id="5272E674F8D888152AAFA7E5E755E6C9">
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<mods:roleTerm id="427035AA4453914993F511ED9A84C9DA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A1A2F4B27A431D0BC9F9B7B8C04F411E">Wägele, Johann-Wolfgang</mods:namePart>
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<mods:affiliation id="F6B9B62EEBF08574616188F5AB1CBC54">Lower Invertebrates, Museum Alexander Koenig ZFMK, Adenauerallee 160, D- 53113 Bonn (Germany) twesener @ uni-bonn. de</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="C868824556290CD50CFDDBAF9F33CD9F">text</mods:typeOfResource>
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<mods:titleInfo id="7F3DB5B3ACE1437D451D0012574D7CDA">
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<mods:title id="8717751B073E0CC8EB30645744DC0B13">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="E7AC9840507DB2C6A8900D366EA73774">
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<mods:date id="2A6BAD2477CF9BF45C59E25A5914A0B0">2008</mods:date>
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<mods:detail id="11C94ED4310B8B26658CE10A6847B671" type="volume">
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<mods:number id="00AA687CC7D493BCA968C2D58B952BCC">30</mods:number>
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</mods:detail>
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<mods:detail id="0F3992DEA294DE5D083604E64F593B5D" type="issue">
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<mods:number id="49748CE6AD147B2A064D69EEFF798846">1</mods:number>
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<mods:extent id="20D8744351C15CB49079B8A1A853E95D" unit="page">
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<mods:start id="0FF64F41BFE0FD39CC1CE91F0F00F640">5</mods:start>
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<mods:end id="CBE6AD4DFC401DDED49D2BE72D07C01F">85</mods:end>
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</mods:extent>
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<mods:classification id="24C59FDAA13385F320FE350D56FDBA68">journal article</mods:classification>
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<mods:identifier id="E3D4119C41D7A6800DC3FEDCF8AF79B6" type="DOI">10.5281/zenodo.5391729</mods:identifier>
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<mods:identifier id="8AD18E411BFC8C9FB06AC729BCBD999F" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="D59FE122232B9B5AAB2A62F308E8CC4F" type="Zenodo-Dep">5391729</mods:identifier>
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<treatment id="AA4514472B1DFFFCFEA9FF70FBB29C52" ID-DOI="http://doi.org/10.5281/zenodo.5467918" ID-Zenodo-Dep="5467918" LSID="urn:lsid:plazi:treatment:AA4514472B1DFFFCFEA9FF70FBB29C52" httpUri="http://treatment.plazi.org/id/AA4514472B1DFFFCFEA9FF70FBB29C52" lastPageNumber="22" pageId="17" pageNumber="22">
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<subSubSection id="6AF6F6DA2B1DFFFCFEA9FF70FC199FB5" box="[358,965,216,242]" pageId="17" pageNumber="22" type="nomenclature">
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<paragraph id="2253A5512B1DFFFCFEA9FF70FC199FB5" blockId="17.[358,965,216,242]" box="[358,965,216,242]" pageId="17" pageNumber="22">
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<heading id="791B123D2B1DFFFCFEA9FF70FC199FB5" box="[358,965,216,242]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="22" reason="4">
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KEY TO THE
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<taxonomicName id="E5ECDED22B1DFFFCFE20FF70FCC39FB5" ID-CoL="5D7HG" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[495,799,216,241]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="22" phylum="Arthropoda" rank="species" species="libidinosum">
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<emphasis id="109879432B1DFFFCFE20FF70FCC39FB5" box="[495,799,216,241]" italics="true" pageId="17" pageNumber="22">ZOOSPHAERIUM LIBIDINOSUM</emphasis>
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</taxonomicName>
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SPECIES- GROUP
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6AF6F6DA2B1DFFFCFF72FEA5FBB29C52" pageId="17" pageNumber="22" type="key">
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<keyStep id="9918BDF42B1DFFFCFF72FEA5FBB29ED2" pageId="17" pageNumber="22">
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<paragraph id="2253A5512B1DFFFCFF72FEA5FBB29E1B" blockId="17.[189,1134,268,407]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FEA5FBB29E1B" pageId="17" pageNumber="22">
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1. Inner margin of anal shield without locking carinae (in small, immature individuals traces of carinae can be visible). Operculum of female vulva well-rounded, not notched (
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<figureCitation id="BAD7B9D42B1DFFFCFBCCFE81FBBE9E04" box="[1027,1122,297,321]" captionStart="FIG" captionStartId="20.[113,124,1612,1629]" captionTargetBox="[135,1190,215,1575]" captionTargetPageId="20" captionText="FIG. 10. — Zoosphaerium libidinosum (de Saussure & Zehntner, 1897), SEM (BLF 5239): A-C right antenna; A, overview, lateral view; B, joint 6 with disc, apical view; C, sensory (?) cones on first joint; D-F gnathochilarium; D, posterior side; E, sensory cone lateral of left palpus; F, sensory cones on central located pads. Abbreviations: cP, central pads; P, palpus. Scale bars: A, 1 mm; B, 300 μm; C, 30 μm; D, 400 μm; E, F, 20 μm." figureDoi="http://doi.org/10.5281/zenodo.5391747" httpUri="https://zenodo.org/record/5391747/files/figure.png" pageId="17" pageNumber="22">Fig. 10C</figureCitation>
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). Tarsi slim, antenna with more than 10 sensory cones. Adults>
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<quantity id="E51408B42B1DFFFCFCA1FEEEFC669E18" box="[878,954,326,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="22" unit="mm" value="2.0">2 mm</quantity>
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...
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<taxonomicName id="E5ECDED22B1DFFFCFC14FEEFFBB29E1B" ID-CoL="5D7HG" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[987,1134,326,350]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="22" phylum="Arthropoda" rank="species" species="libidinosum">
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<emphasis id="109879432B1DFFFCFC14FEEFFBB29E1B" box="[987,1134,326,350]" italics="true" pageId="17" pageNumber="22">Z. libidinosum</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="2253A5512B1DFFFCFF72FECBFBB29ED2" blockId="17.[189,1134,268,407]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FECBFBB29ED2" pageId="17" pageNumber="22">
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— Inner margin of anal shield with 2 well-visible locking carinae (
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<figureCitation id="BAD7B9D42B1DFFFCFCA2FECBFC179E3E" box="[877,971,355,379]" captionStart="FIG" captionStartId="24.[113,124,1589,1606]" captionTargetBox="[120,1199,219,1542]" captionTargetId="figure-22@24.[115,1208,215,1550]" captionTargetPageId="24" captionText="FIG. 14. — Zoosphaerium priapus (de Saussure & Zehntner,1897):A, B, E, ♀ (FMNH 3911);A, coxa of 1st left leg with 1st stigma-carrying plate, posterior view; B, 9th left leg; E, subanal plate with washboard; C, D lectotype; C, closing ledges of anal shield; D, coxa of 2nd right leg with vulva;F-L, ♂ (FMNH 3911); F, coxa of 2nd left leg with gonopore;G-J left anterior telopod; G, anterior view; H, posterior view; I, inner view; J, lateral view; K, L right posterior telopod; K, anterior view; L, posterior view. Abbreviations: 12T, 12th tergite; AS, anal shield; Cx, coxa; LT, laterotergite; St, stigma-carrying plate. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391753" httpUri="https://zenodo.org/record/5391753/files/figure.png" pageId="17" pageNumber="22">Fig. 14C</figureCitation>
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). Apical margin of vulva operculum medially notched (
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<figureCitation id="BAD7B9D42B1DFFFCFDB7FED7FD0B9ED2" box="[632,727,383,407]" captionStart="FIG" captionStartId="24.[113,124,1589,1606]" captionTargetBox="[120,1199,219,1542]" captionTargetId="figure-22@24.[115,1208,215,1550]" captionTargetPageId="24" captionText="FIG. 14. — Zoosphaerium priapus (de Saussure & Zehntner,1897):A, B, E, ♀ (FMNH 3911);A, coxa of 1st left leg with 1st stigma-carrying plate, posterior view; B, 9th left leg; E, subanal plate with washboard; C, D lectotype; C, closing ledges of anal shield; D, coxa of 2nd right leg with vulva;F-L, ♂ (FMNH 3911); F, coxa of 2nd left leg with gonopore;G-J left anterior telopod; G, anterior view; H, posterior view; I, inner view; J, lateral view; K, L right posterior telopod; K, anterior view; L, posterior view. Abbreviations: 12T, 12th tergite; AS, anal shield; Cx, coxa; LT, laterotergite; St, stigma-carrying plate. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391753" httpUri="https://zenodo.org/record/5391753/files/figure.png" pageId="17" pageNumber="22">Fig. 14D</figureCitation>
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) ....................................................... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="9918BDF42B1DFFFCFF72FE1BFBB29EAD" pageId="17" pageNumber="22">
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<paragraph id="2253A5512B1DFFFCFF72FE1BFBB29E8E" blockId="17.[189,1134,435,488]" box="[189,1134,435,459]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FE1BFBB29E8E" box="[189,1134,435,459]" pageId="17" pageNumber="22">2. Hindmost point of anal shield extended, slightly protruding posteriorly .................... 3</keyLead>
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</paragraph>
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<paragraph id="2253A5512B1DFFFCFF72FE78FBB29EAD" blockId="17.[189,1134,435,488]" box="[189,1134,463,488]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FE78FBB29EAD" box="[189,1134,463,488]" pageId="17" pageNumber="22">— Hindmost end of anal shield well-rounded ................................................................. 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="9918BDF42B1DFFFCFF72FDABFBB29D11" pageId="17" pageNumber="22">
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<paragraph id="2253A5512B1DFFFCFF72FDABFBB29D72" blockId="17.[189,1134,515,596]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FDABFBB29D72" pageId="17" pageNumber="22">
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3. Posterior locking carina of anal shield more than 3 times longer than anterior carina ..................................................................................................................
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<taxonomicName id="E5ECDED22B1DFFFCFC3DFD88FBB29D72" ID-CoL="5D7GH" authorityName="Wesener & Sierwald" authorityYear="2005" box="[1010,1134,543,567]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="22" phylum="Arthropoda" rank="species" species="arborealis">
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<emphasis id="109879432B1DFFFCFC3DFD88FBB29D72" box="[1010,1134,543,567]" italics="true" pageId="17" pageNumber="22">Z. arborealis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="2253A5512B1DFFFCFF72FD95FBB29D11" blockId="17.[189,1134,515,596]" box="[189,1134,572,596]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FD95FBB29D11" box="[189,1134,572,596]" pageId="17" pageNumber="22">
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— Posterior locking carina less than 3 times longer than anterior one ..........
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<taxonomicName id="E5ECDED22B1DFFFCFC2AFD95FBB29D11" ID-CoL="5D7HD" baseAuthorityName="Broelemann" baseAuthorityYear="1922" box="[997,1134,572,596]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="22" phylum="Arthropoda" rank="species" species="lambertoni">
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<emphasis id="109879432B1DFFFCFC2AFD95FBB29D11" box="[997,1134,572,596]" italics="true" pageId="17" pageNumber="22">Z. lambertoni</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="9918BDF42B1DFFFCFF72FDD8FBB29C52" pageId="17" pageNumber="22">
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<paragraph id="2253A5512B1DFFFCFF72FDD8FBB29D84" blockId="17.[189,1135,624,791]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FDD8FBB29D84" pageId="17" pageNumber="22">
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4. Brown-blackish coloration pattern. More than 10 sensorial cones on antenna (
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<figureCitation id="BAD7B9D42B1DFFFCFBCBFDD8FBBE9DCD" box="[1028,1122,624,648]" captionStart="FIG" captionStartId="6.[113,124,1567,1584]" captionTargetBox="[160,1177,218,1520]" captionTargetId="figure-22@6.[113,1209,215,1520]" captionTargetPageId="6" captionText="FIG. 1. — Zoosphaerium neptunus (Butler, 1872): A-C ♂ (MNHN CB020); A, habitus; B, coxa of 1st left leg with 1st stigma-carrying plate; C, posterior body end with anal shield, ventral view, last three leg pairs and telopods removed; D, closing ledges of anal shield (holotype of Z. digitale n. syn.); E, closing ledges of anal shield (lectotype of Z. pygidiale n. syn.); F, G, holotype; F, 9th left leg; G, operculum of vulva; H, coxa of 2nd left leg with vulva (paralectotype of Z. pygidiale n. syn.); I, coxa of 2nd left leg with gonopore (MNHN CB020). Abbreviations:12T, 12th tergite; AS, anal shield; Cx, coxa; LT, laterotergite; O, operculum; St, stigma-carrying plate. Scale bars: A, 10 mm; B-I, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391731" httpUri="https://zenodo.org/record/5391731/files/figure.png" pageId="17" pageNumber="22">Fig. 1 C</figureCitation>
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). Legs short, covered with long hairs (
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<figureCitation id="BAD7B9D42B1DFFFCFD92FD24FD659DE0" box="[605,697,652,677]" captionStart="FIG" captionStartId="24.[113,124,1589,1606]" captionTargetBox="[120,1199,219,1542]" captionTargetId="figure-22@24.[115,1208,215,1550]" captionTargetPageId="24" captionText="FIG. 14. — Zoosphaerium priapus (de Saussure & Zehntner,1897):A, B, E, ♀ (FMNH 3911);A, coxa of 1st left leg with 1st stigma-carrying plate, posterior view; B, 9th left leg; E, subanal plate with washboard; C, D lectotype; C, closing ledges of anal shield; D, coxa of 2nd right leg with vulva;F-L, ♂ (FMNH 3911); F, coxa of 2nd left leg with gonopore;G-J left anterior telopod; G, anterior view; H, posterior view; I, inner view; J, lateral view; K, L right posterior telopod; K, anterior view; L, posterior view. Abbreviations: 12T, 12th tergite; AS, anal shield; Cx, coxa; LT, laterotergite; St, stigma-carrying plate. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391753" httpUri="https://zenodo.org/record/5391753/files/figure.png" pageId="17" pageNumber="22">Fig. 14B</figureCitation>
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). Third joint of posterior telopods without small hairs, second joint basally with one large, membranous lobe (
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<figureCitation id="BAD7B9D42B1DFFFCFC4AFD01FC039D84" box="[901,991,681,705]" captionStart="FIG" captionStartId="24.[113,124,1589,1606]" captionTargetBox="[120,1199,219,1542]" captionTargetId="figure-22@24.[115,1208,215,1550]" captionTargetPageId="24" captionText="FIG. 14. — Zoosphaerium priapus (de Saussure & Zehntner,1897):A, B, E, ♀ (FMNH 3911);A, coxa of 1st left leg with 1st stigma-carrying plate, posterior view; B, 9th left leg; E, subanal plate with washboard; C, D lectotype; C, closing ledges of anal shield; D, coxa of 2nd right leg with vulva;F-L, ♂ (FMNH 3911); F, coxa of 2nd left leg with gonopore;G-J left anterior telopod; G, anterior view; H, posterior view; I, inner view; J, lateral view; K, L right posterior telopod; K, anterior view; L, posterior view. Abbreviations: 12T, 12th tergite; AS, anal shield; Cx, coxa; LT, laterotergite; St, stigma-carrying plate. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391753" httpUri="https://zenodo.org/record/5391753/files/figure.png" pageId="17" pageNumber="22">Fig. 14K</figureCitation>
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) ...
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<taxonomicName id="E5ECDED22B1DFFFCFBC4FD02FBB29D84" ID-CoL="5D7HP" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[1035,1134,682,705]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="22" phylum="Arthropoda" rank="species" species="priapus">
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<emphasis id="109879432B1DFFFCFBC4FD02FBB29D84" box="[1035,1134,682,705]" italics="true" pageId="17" pageNumber="22">Z. priapus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="2253A5512B1DFFFCFF72FD6EFBB29C52" blockId="17.[189,1135,624,791]" pageId="17" pageNumber="22">
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<keyLead id="991D06642B1DFFFCFF72FD6EFBB29C52" pageId="17" pageNumber="22">
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— Shiny green coloration pattern (Fig. A). Four sensorial cones on antenna. Legs elongated. Second and third joints of posterior telopods covered with numerous minute hairs. Process of second joint basally without membranous lobe (
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<figureCitation id="BAD7B9D42B1DFFFCFD3BFD57FC829C52" box="[756,862,767,791]" captionStart="FIG" captionStartId="30.[113,124,1589,1606]" captionTargetBox="[116,1204,217,1533]" captionTargetId="figure-22@30.[115,1208,215,1550]" captionTargetPageId="30" captionText="FIG. 17. — Zoosphaerium anomalum (de Saussure & Zehntner,1902),holotype:A, habitus,body length 20.2 mm; B, coxa of 1st right leg with 1st stigma-carrying plate,posterior view;C, closing ledges of anal shield;D, coxa of 2nd right leg with gonopore;E-H, left anterior telopod; E, anterior view; F, posterior view; G, lateral view; H, inner view; I, J left posterior telopod; I, anterior view; J, posterior view. Abbreviations: 12T, 12th tergite; AS, anal shield; Cx, coxa; St, stigma-carrying plate; x, inner process of joint 3 of anterior telopods; y, lateral process of joint 3 of anterior telopods; z, process of joint 2 of anterior telopods. Scale bars: A, 5 mm; B-J, 1mm." figureDoi="http://doi.org/10.5281/zenodo.5391759" httpUri="https://zenodo.org/record/5391759/files/figure.png" pageId="17" pageNumber="22">Fig. 17I, J</figureCitation>
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) ................
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<taxonomicName id="E5ECDED22B1DFFFCFC29FCA8FBB29C52" ID-CoL="5D7GG" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1902" box="[998,1134,767,791]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="22" phylum="Arthropoda" rank="species" species="anomalum">
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<emphasis id="109879432B1DFFFCFC29FCA8FBB29C52" box="[998,1134,767,791]" italics="true" pageId="17" pageNumber="22">Z. anomalum</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |