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<document id="80D480B21316749BF135B006E9B3E18D" ID-DOI="10.5281/zenodo.185153" ID-GBIF-Dataset="0b5ad8cc-9b48-4700-acc5-012d1707f720" ID-ISSN="1175-5326" ID-Zenodo-Dep="185153" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1459966635824" checkinUser="plazi" docAuthor="Nihei, Silvio Shigueo &amp; Carvalho, Claudio José Barros De" docDate="2009" docId="E60987AB2400E1639686F97CFC16F207" docLanguage="en" docName="zt01976p024.pdf" docOrigin="Zootaxa 1976" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2814bb39" docStyleName="zootaxa.2007.monograph" docTitle="Morellia nigricosta" docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="1A30FFD3240EE16C9611FFBDFF97F67C" masterDocTitle="The Muscini flies of the world (Diptera, Muscidae): identification key and generic diagnoses" masterLastPageNumber="24" masterPageNumber="1" pageNumber="15" updateTime="1698231170040" updateUser="plazi">
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<mods:title id="93168B08EE567FA18EFFA2D64E39C9F1">The Muscini flies of the world (Diptera, Muscidae): identification key and generic diagnoses</mods:title>
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<mods:namePart id="290223D402DA25D53532422D3CD72B7A">Nihei, Silvio Shigueo</mods:namePart>
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<mods:namePart id="5D0DAD13EAC6411CB32EBDAEBF8DAF5B">Carvalho, Claudio José Barros De</mods:namePart>
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<mods:date id="5D358C6326B99DF9A62A7B00B4FFB07B">2009</mods:date>
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<mods:number id="E2B215291261B858385A1A66EFC1FA38">1976</mods:number>
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<treatment id="E60987AB2400E1639686F97CFC16F207" ID-DOI="http://doi.org/10.5281/zenodo.5658505" ID-GBIF-Taxon="119377428" ID-Zenodo-Dep="5658505" LSID="urn:lsid:plazi:treatment:E60987AB2400E1639686F97CFC16F207" httpUri="http://treatment.plazi.org/id/E60987AB2400E1639686F97CFC16F207" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="26BA65362400E1629686F97CFE45F0A6" box="[151,466,1728,1755]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="6E1F36BD2400E1629686F97CFE45F0A6" blockId="14.[151,466,1728,1755]" box="[151,466,1728,1755]" pageId="14" pageNumber="15">
<heading id="355781D12400E1629686F97CFE45F0A6" bold="true" box="[151,466,1728,1755]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="5CD4EAAF2400E1629686F97CFE45F0A6" bold="true" box="[151,466,1728,1755]" pageId="14" pageNumber="15">
<taxonomicName id="A9A04D3E2400E1629686F97CFEE9F0A7" ID-CoL="44CZJ" box="[151,382,1729,1755]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigricosta">
<emphasis id="5CD4EAAF2400E1629686F97CFEE9F0A7" bold="true" box="[151,382,1729,1755]" italics="true" pageId="14" pageNumber="15">Morellia nigricosta</emphasis>
</taxonomicName>
-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="26BA65362400E1639686F8B6FB91F7B3" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="6E1F36BD2400E1639686F8B6FAFEF7DB" blockId="14.[151,1437,1803,2029]" lastBlockId="15.[151,1437,152,1147]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="5CD4EAAF2400E1629686F8B6FEB8F159" bold="true" box="[151,303,1803,1829]" pageId="14" pageNumber="15">Description.</emphasis>
Colouration metallic violaceous blue; wing with maculae. Female with proclinate fronto-orbital seta undeveloped; parafrons almost entirely setulose; interfrontal seta absent. Humerals 3; posthumerals developed or not; parahumeral undeveloped; presutural strong; intra-alar 1; intrapostalar present. Presutural acrostichals undeveloped; one postsutural pair. Prosternum setulose (
<figureCitation id="F69B2A382400E16295ADF83EFB94F1E1" box="[956,1027,1923,1949]" captionStart="FIGURES 1 – 8. 1" captionStartId="5.[151,269,1686,1710]" captionTargetBox="[228,1368,278,1625]" captionTargetId="figure@5.[219,1369,266,1625]" captionTargetPageId="5" captionText="FIGURES 1 – 8. 1. Head, Polietina orbitalis (Stein), female, lateral view. 2. Same, upper frontal view. 3. Head, Polietina major (Albuquerque), male, lateral view. 4. Head, Morellia (Xenomorellia) undescribed sp., female, frontal view. 5. Antenna, Pyrellia albocuprea Villeneuve, inner lateral view. 6. Prosternum, Biopyrellia bipuncta (Wiedemann), anteroventral view. 7. Prosternum, Morellia nigricosta Hough, anteroventral view. 8. Prosternum, Mitroplatia smaragdina (Séguy), anteroventral view. Figures 1 – 2 and 5 after Nihei and de Carvalho (2007 a); Figure 3 after Nihei (2002). Figures 6 – 8 on same scale." httpUri="https://zenodo.org/record/185154/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
). Katepisternals 1+2. Postalar wall bare; anterior and posterior suprasquamal ridges bare. Anepimeron and katepimeron setulose. Metakatepisternum setulose above coxa. Greater ampulla, proepisternum and meron bare. Posterior spiracle setulose on posterior margin. Wing with the basal portion of stem-vein bare dorsally and ventrally; apical portion of stemvein bare dorsally and ventrally; Rs node setulose dorsally and ventrally; R4+5 setulose dorsally, and ventrally bare or setulose; M bent forward towards R4+5. Lower calypter enlarged posteriorly, extending under base of scutellum. Subcostal sclerite bare. Fore tibia bare on posteroventral surface. Male mid femur without a preapical protuberance with setae on dorsal surface; without row of developed setae at mid third of anterior surface. Mid tibia bare on anterodorsal surface. Calcar absent. First abdominal sternite setulose. Male terminalia: cercal plate with the median and marginal spined processes on ventral surface; distiphallus without spinules.
</paragraph>
<paragraph id="6E1F36BD2401E16396D7FE08FB91F7B3" blockId="15.[151,1437,152,1147]" box="[198,1030,437,463]" pageId="15" pageNumber="16">
<emphasis id="5CD4EAAF2401E16396D7FE08FE05F7B3" bold="true" box="[198,402,437,463]" pageId="15" pageNumber="16">Included species</emphasis>
:
<taxonomicName id="A9A04D3E2401E16397B3FE0BFD3AF7B3" authority="Hough" authorityName="Hough" box="[418,685,437,463]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nigricosta">
<emphasis id="5CD4EAAF2401E16397B3FE0BFDD7F7B3" box="[418,576,438,463]" italics="true" pageId="15" pageNumber="16">M. nigricosta</emphasis>
(Hough)
</taxonomicName>
,
<taxonomicName id="A9A04D3E2401E16394A8FE0BFB95F7B3" authority="Pamplona" authorityName="Pamplona" box="[697,1026,437,463]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="xanthoptera">
<emphasis id="5CD4EAAF2401E16394A8FE0BFCFAF7B3" box="[697,877,438,463]" italics="true" pageId="15" pageNumber="16">M. xanthoptera</emphasis>
(Pamplona)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="26BA65362401E16396D4FE60FD0FF5A7" pageId="15" pageNumber="16" type="discussion">
<paragraph id="6E1F36BD2401E16396D4FE60FD0FF5A7" blockId="15.[151,1437,152,1147]" pageId="15" pageNumber="16">
<emphasis id="5CD4EAAF2401E16396D4FE60FEC3F78B" bold="true" box="[197,340,477,503]" pageId="15" pageNumber="16">Comments.</emphasis>
At least two Neotropical species have been recognized as belonging to this group:
<taxonomicName id="A9A04D3E2401E1639320FE63FED7F463" authority="Hough" authorityName="Hough" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nigricosta">
<emphasis id="5CD4EAAF2401E1639320FE63FF42F463" italics="true" pageId="15" pageNumber="16">M. nigricosta</emphasis>
(Hough)
</taxonomicName>
and
<taxonomicName id="A9A04D3E2401E163976DFDBBFD51F463" authority="Pamplona" authorityName="Pamplona" box="[380,710,517,543]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="xanthoptera">
<emphasis id="5CD4EAAF2401E163976DFDBBFDA6F463" box="[380,561,518,543]" italics="true" pageId="15" pageNumber="16">M. xanthoptera</emphasis>
(Pamplona)
</taxonomicName>
. This species-group was regarded as a genus-ranked taxon in the
<taxonomicName id="A9A04D3E2401E16396D4FD90FEB2F43B" box="[197,293,557,583]" class="Insecta" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="tribe" tribe="Muscini">Muscini</taxonomicName>
phylogeny (Nihei &amp; de
<bibRefCitation id="0A314B4C2401E1639444FD90FC88F43B" author="Carvalho" box="[597,799,557,583]" pageId="15" pageNumber="22" refString="Nihei, S. S. &amp; de Carvalho, C. J. B. (2007 a) Phylogeny and classification of Muscini (Diptera, Muscidae). Zoological Journal of the Linnean Society, 149, 493 - 532." type="journal article" year="2007" yearSuffix="a">Carvalho, 2007a</bibRefCitation>
), placed far from the large
<taxonomicName id="A9A04D3E2401E1639260FD93FB41F43B" box="[1137,1238,558,583]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E1639260FD93FB41F43B" box="[1137,1238,558,583]" italics="true" pageId="15" pageNumber="16">Morellia</emphasis>
</taxonomicName>
clade and more closely related to
<emphasis id="5CD4EAAF2401E1639763FDEBFE7DF413" box="[370,490,598,623]" italics="true" pageId="15" pageNumber="16">Ziminellia</emphasis>
and
<taxonomicName id="A9A04D3E2401E163943AFDEBFD26F413" box="[555,689,598,623]" class="Insecta" family="Muscidae" genus="Mitroplatia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E163943AFDEBFD26F413" box="[555,689,598,623]" italics="true" pageId="15" pageNumber="16">Mitroplatia</emphasis>
</taxonomicName>
, with which it composes the base of the clade giving rise to
<taxonomicName id="A9A04D3E2401E1639686FDC3FF72F4EB" box="[151,229,638,663]" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E1639686FDC3FF72F4EB" box="[151,229,638,663]" italics="true" pageId="15" pageNumber="16">Musca</emphasis>
</taxonomicName>
,
<taxonomicName id="A9A04D3E2401E16396E2FDC3FECCF4EB" box="[243,347,638,663]" class="Insecta" family="Muscidae" genus="Neomyia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E16396E2FDC3FECCF4EB" box="[243,347,638,663]" italics="true" pageId="15" pageNumber="16">Neomyia</emphasis>
</taxonomicName>
,
<taxonomicName id="A9A04D3E2401E163977BFDC3FE66F4EB" box="[362,497,638,663]" class="Insecta" family="Muscidae" genus="Curranosia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E163977BFDC3FE66F4EB" box="[362,497,638,663]" italics="true" pageId="15" pageNumber="16">Curranosia</emphasis>
</taxonomicName>
,
<taxonomicName id="A9A04D3E2401E16397EEFDC3FDCBF4EB" box="[511,604,638,663]" class="Insecta" family="Muscidae" genus="Pyrellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E16397EEFDC3FDCBF4EB" box="[511,604,638,663]" italics="true" pageId="15" pageNumber="16">Pyrellia</emphasis>
</taxonomicName>
,
<taxonomicName id="A9A04D3E2401E163947AFDC3FCB6F4EB" box="[619,801,638,663]" class="Insecta" family="Muscidae" genus="Sarcopromusca" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E163947AFDC3FCB6F4EB" box="[619,801,638,663]" italics="true" pageId="15" pageNumber="16">Sarcopromusca</emphasis>
</taxonomicName>
and
<taxonomicName id="A9A04D3E2401E1639548FDC3FC4CF4EB" box="[857,987,638,663]" class="Insecta" family="Muscidae" genus="Dasyphora" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E1639548FDC3FC4CF4EB" box="[857,987,638,663]" italics="true" pageId="15" pageNumber="16">Dasyphora</emphasis>
</taxonomicName>
. The phylogenetic characters supporting the
<taxonomicName id="A9A04D3E2401E16396E0FD1BFEFFF4C3" box="[241,360,678,703]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nigricosta">
<emphasis id="5CD4EAAF2401E16396E0FD1BFEFFF4C3" box="[241,360,678,703]" italics="true" pageId="15" pageNumber="16">nigricosta</emphasis>
</taxonomicName>
-group as a clade were the following (fig. 50 of Nihei &amp; de
<bibRefCitation id="0A314B4C2401E1639222FD18FB63F4C3" author="Carvalho" box="[1075,1268,677,703]" pageId="15" pageNumber="22" refString="Nihei, S. S. &amp; de Carvalho, C. J. B. (2007 a) Phylogeny and classification of Muscini (Diptera, Muscidae). Zoological Journal of the Linnean Society, 149, 493 - 532." type="journal article" year="2007" yearSuffix="a">Carvalho 2007a</bibRefCitation>
): katepimeron setulose; brown macula on humeral vein, apex of Sc and R1, apex of R2+3, and crossvein r-m; male mid femur without series of developed setae on median third of the anterior surface; and cercal plate with marginal spined process on ventral face. Here, a formal description of the
<taxonomicName id="A9A04D3E2401E1639589FC9CFB98F546" box="[920,1039,801,826]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nigricosta">
<emphasis id="5CD4EAAF2401E1639589FC9CFB98F546" box="[920,1039,801,826]" italics="true" pageId="15" pageNumber="16">nigricosta</emphasis>
</taxonomicName>
-group as a new genus is not provided as we consider it a premature proposal, at least until more detailed studies support and corroborate it as a valid generic taxon. Interestingly, the molecular analysis of
<bibRefCitation id="0A314B4C2401E1639563FCCCFBE6F5F7" box="[882,1137,881,907]" pageId="15" pageNumber="23" refString="Schuehli, G. S., de Carvalho, C. J. B. &amp; Wiegmann, B. M. (2007) Molecular phylogenetics of the Muscidae (Diptera: Calyptratae): new ideas in a congruence context. Invertebrate Systematics, 21, 263 - 278." type="journal article">Schuehli et al. (2007)</bibRefCitation>
indicated the non-monophyly of
<taxonomicName id="A9A04D3E2401E1639719FC24FEFAF5CE" box="[264,365,921,946]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="5CD4EAAF2401E1639719FC24FEFAF5CE" box="[264,365,921,946]" italics="true" pageId="15" pageNumber="16">Morellia</emphasis>
</taxonomicName>
as its two terminal representatives [
<taxonomicName id="A9A04D3E2401E163953DFC24FC72F5CE" box="[812,997,921,946]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="xanthoptera">
<emphasis id="5CD4EAAF2401E163953DFC24FC72F5CE" box="[812,997,921,946]" italics="true" pageId="15" pageNumber="16">M. xanthoptera</emphasis>
</taxonomicName>
and
<taxonomicName id="A9A04D3E2401E1639234FC24FAF9F5CF" authority="Wiedemann" authorityName="Wiedemann" box="[1061,1390,921,947]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ochricornis">
<emphasis id="5CD4EAAF2401E1639234FC24FB40F5CE" box="[1061,1239,921,946]" italics="true" pageId="15" pageNumber="16">M. ochricornis</emphasis>
Wiedemann
</taxonomicName>
(as
<emphasis id="5CD4EAAF2401E1639686FC7CFE81F5A6" box="[151,278,961,986]" italics="true" pageId="15" pageNumber="16">obscuripes</emphasis>
Bigot)] were not closely related.
</paragraph>
</subSubSection>
<subSubSection id="26BA65362401E16396D4FC55FD12F27F" box="[197,645,1000,1027]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="6E1F36BD2401E16396D4FC55FD12F27F" blockId="15.[151,1437,152,1147]" box="[197,645,1000,1027]" pageId="15" pageNumber="16">
<emphasis id="5CD4EAAF2401E16396D4FC55FEC9F27E" bold="true" box="[197,350,1000,1026]" pageId="15" pageNumber="16">Distribution</emphasis>
(2 species). Neotropical.
</paragraph>
</subSubSection>
<subSubSection id="26BA65362401E16396D4FBADFC16F207" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="6E1F36BD2401E16396D4FBADFC16F207" blockId="15.[151,1437,152,1147]" pageId="15" pageNumber="16">
<emphasis id="5CD4EAAF2401E16396D4FBADFEC2F256" bold="true" box="[197,341,1040,1066]" pageId="15" pageNumber="16">References.</emphasis>
<bibRefCitation id="0A314B4C2401E163974CFBACFD3FF257" author="Pamplona" box="[349,680,1041,1067]" pageId="15" pageNumber="22" refString="Pamplona, D. M. &amp; Couri, M. S. (1995) Morellia dendropanacis, a new species, and other species with spotted wings: characterization and comparison (Diptera: Muscidae: Muscinae). Proceedings of the Biological Society of Washington, 108, 451 - 457." type="journal article" year="1995">Pamplona and Couri (1995)</bibRefCitation>
, de
<bibRefCitation id="0A314B4C2401E16394C8FBACFB8DF257" author="Carvalho" box="[729,1050,1041,1067]" pageId="15" pageNumber="21" refString="de Carvalho, C. J. B. &amp; Couri, M. S. (2002) Part I. Basal groups. In: de Carvalho, C. J. B. (Ed.), Muscidae (Diptera) of the Neotropical Region: taxonomy. Editora Universidade Federal do Parana, Curitiba, pp. 17 - 132." type="book chapter" year="2002">Carvalho and Couri (2002)</bibRefCitation>
, Nihei and de
<bibRefCitation id="0A314B4C2401E16392D7FBACFA0EF257" author="Carvalho" box="[1222,1433,1041,1067]" pageId="15" pageNumber="22" refString="Nihei, S. S. &amp; de Carvalho, C. J. B. (2007 a) Phylogeny and classification of Muscini (Diptera, Muscidae). Zoological Journal of the Linnean Society, 149, 493 - 532." type="journal article" year="2007" yearSuffix="a">Carvalho (2007a)</bibRefCitation>
. [Note: No mention is made of a
<taxonomicName id="A9A04D3E2401E1639404FB84FD1BF22E" ID-CoL="44CZJ" box="[533,652,1081,1106]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nigricosta">
<emphasis id="5CD4EAAF2401E1639404FB84FD1BF22E" box="[533,652,1081,1106]" italics="true" pageId="15" pageNumber="16">nigricosta</emphasis>
</taxonomicName>
-group in these references, although taxonomic information and/or keys including
<taxonomicName id="A9A04D3E2401E163975BFBDCFE7FF206" ID-CoL="44CZJ" box="[330,488,1121,1146]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nigricosta">
<emphasis id="5CD4EAAF2401E163975BFBDCFE7FF206" box="[330,488,1121,1146]" italics="true" pageId="15" pageNumber="16">M. nigricosta</emphasis>
</taxonomicName>
and
<taxonomicName id="A9A04D3E2401E1639430FBDCFD41F206" ID-CoL="44D2F" box="[545,726,1121,1146]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="xanthoptera">
<emphasis id="5CD4EAAF2401E1639430FBDCFD41F206" box="[545,726,1121,1146]" italics="true" pageId="15" pageNumber="16">M. xanthoptera</emphasis>
</taxonomicName>
are provided.]
</paragraph>
</subSubSection>
</treatment>
</document>
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