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<document id="ED16CCC3F0688BA8A2AC972CAB9C4F31" ID-CLB-Dataset="21416" ID-DOI="10.5852/ejt.2020.690" ID-GBIF-Dataset="c6d0d802-2934-45b6-8561-bf4ff9930444" ID-Zenodo-Dep="3961787" ID-ZooBank="C5EA97FE-0FFE-44E5-91F9-DA2F7C3420A4" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595861068960" checkinUser="valdenar" docAuthor="Sheth, Sayali D., Ghate, Hemant V. &amp; Fikáček, Martin" docDate="2020" docId="646D87BB8E2EFE2A2E61FE80400A97C7" docLanguage="en" docName="ejt-690_sheth_ghate_fikacek.pdf" docOrigin="European Journal of Taxonomy 690" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Coelostoma lyratum Sheth &amp; Ghate &amp; Fikáček 2020, sp. nov." docType="treatment" docUuid="6E05380A-D3AA-410E-A974-D1CED984068A" docUuidSource="ZooBank" docVersion="9" lastPageNumber="17" masterDocId="9854FFC38E20FE3A2C22FF8A434E9604" masterDocTitle="Review of Coelostoma of the Indian subcontinent (Coleoptera: Hydrophilidae) Part 1: Coelostoma s. str. and Holocoelostoma" masterLastPageNumber="32" masterPageNumber="1" pageNumber="15" updateTime="1698843983510" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="8C150AAF92785A40E0470D31E7C64B46">
<mods:title id="73236CA762DA45BC0C6F1440ABABA182">Review of Coelostoma of the Indian subcontinent (Coleoptera: Hydrophilidae) Part 1: Coelostoma s. str. and Holocoelostoma</mods:title>
</mods:titleInfo>
<mods:name id="C1EC125BCACCBB07BFE55EA46984A9B8" type="personal">
<mods:role id="FC9503C099D5652A246D8164DA898668">
<mods:roleTerm id="CBDC792C7353067DB494D44BD09D4835">Author</mods:roleTerm>
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<mods:namePart id="AF8551281F5A3168854CB7E64C47AC33">Sheth, Sayali D.</mods:namePart>
<mods:nameIdentifier id="F3C6222F88A932DA8C01BFA935FBE547" type="LSID">1F5AD99F-7DDE-455A-9DA0-6E9C263F0A5E</mods:nameIdentifier>
<mods:affiliation id="5789235B5399762D21A469514A4C237D">Annasaheb Kulkarni Department of Biodiversity, Abasaheb Garware College, Pune, India.</mods:affiliation>
<mods:nameIdentifier id="0084C542DB101296E6444CFF7DA5388D" type="email">saylisheth@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="3174FA8929E7627C342BFB923991ACFA" type="personal">
<mods:role id="B600DDE738447D3084A613325ACED584">
<mods:roleTerm id="1F2099A14DDA2BEF83DB6C2EC498FCD9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9F332BDEB1B7B371461A289B2F781C2">Ghate, Hemant V.</mods:namePart>
<mods:nameIdentifier id="A26F207204B759BE2E80AADBB2FADE2B" type="LSID">26ED55BA-F5FF-4F89-98E8-348B506A4CCB</mods:nameIdentifier>
<mods:affiliation id="725E6695AE05CEA15A4293644D25E7CB">Department of Zoology, Modern College, Shivajinagar, Pune, India.</mods:affiliation>
<mods:nameIdentifier id="D47B3FFD8833309C7EC10C7EFB853F52" type="email">hemantghate@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="35E49BEF3BFC13D8D7592AC311221CCE">Fikáček, Martin</mods:namePart>
<mods:nameIdentifier id="DA5A988EF05A43A826D3B0BDEA385451" type="LSID">E1A02744-41A2-4B9C-A127-B78F0D812637</mods:nameIdentifier>
<mods:affiliation id="EB2E68E6FD5F26029FE23F715314C86E">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19300, Praha 9 – Horní Počernice, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844, Praha 2, Czech Republic.</mods:affiliation>
<mods:nameIdentifier id="5441DF37BC7201E01F7B0C044D9A4F30" type="email">mfikacek@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="FEDC9F4D0462B9D35AA647E1E738DA6F">text</mods:typeOfResource>
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<mods:title id="E0617C75A6503AF8C89A9BCD99130D1E">European Journal of Taxonomy</mods:title>
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<mods:part id="751EB1861B85BB5AF154A080D3F8DFCD">
<mods:date id="B5C92309A8A08070A714F96D3719E5C4">2020</mods:date>
<mods:detail id="04D76CFE52142FE8F487E6EC90F299B4" type="pubDate">
<mods:number id="9C6AC03FA56F6A689F85723970B8E59B">2020-07-24</mods:number>
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<mods:detail id="42AF81E42C9036EAD063A16B3BC14722" type="volume">
<mods:number id="A400AC0BB852845300DA1840D2ED18D6">690</mods:number>
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<mods:start id="BEF0B3D93CB574CC46CBAE4B689A48C9">1</mods:start>
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<treatment id="646D87BB8E2EFE2A2E61FE80400A97C7" ID-DOI="http://doi.org/10.5281/zenodo.4329887" ID-GBIF-Taxon="166003095" ID-Zenodo-Dep="4329887" LSID="urn:lsid:zoobank.org:act:6E05380A-D3AA-410E-A974-D1CED984068A" httpUri="http://treatment.plazi.org/id/646D87BB8E2EFE2A2E61FE80400A97C7" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="A4DE65268E2EFE342E61FE8040BF9721" box="[579,1009,266,293]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="EC7B36AD8E2EFE342E61FE8040BF9721" blockId="14.[360,1229,266,363]" box="[579,1009,266,293]" pageId="14" pageNumber="15">
<heading id="B73381C18E2EFE342E61FE8040BF9721" box="[579,1009,266,293]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="2BC44D2E8E2EFE342E61FE8040C19721" ID-CoL="8XXZZ" authority="Sheth &amp; Ghate &amp; Fikáček, 2020" authorityName="Sheth &amp; Ghate &amp; Fikáček" authorityYear="2020" box="[579,911,266,293]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lyratum" status="sp. nov.">
<emphasis id="DEB0EABF8E2EFE342E61FE80419F9721" bold="true" box="[579,721,266,293]" italics="true" pageId="14" pageNumber="15">Coelostoma</emphasis>
(s. str.)
<emphasis id="DEB0EABF8E2EFE342F12FE8040C19721" bold="true" box="[816,911,266,293]" italics="true" pageId="14" pageNumber="15">lyratum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C58357C48E2EFE342FB4FE8140BF9721" box="[918,1009,267,293]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4DE65268E2EFE342D4AFEA44016976F" pageId="14" pageNumber="15" type="description">
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<uri id="98553AAF8E2EFE342D4AFEA44783974D" box="[360,1229,302,329]" pageId="14" pageNumber="15">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="EC7B36AD8E2EFE342EF9FEDB4016976F" blockId="14.[360,1229,266,363]" box="[731,856,337,363]" pageId="14" pageNumber="15">
<figureCitation id="74FF2A288E2EFE342EF9FEDB4050976F" box="[731,798,337,363]" captionStart="Fig" captionStartId="14.[189,232,1885,1911]" captionTargetBox="[189,1379,907,1840]" captionTargetId="figure@14.[189,1396,890,1847]" captionTargetPageId="14" captionText="Fig. 6. Coelostoma (s. str.) lyratum sp. nov., holotype, ♂ (NMPC). A. Dorsal habitus. B. Lateral habitus. C. Frontal view. D. Thorax in ventral view. E. Dorsal punctation of head. F. Dorsal punctation of pronotum. G. Dorsal punctation of elytron. H. Abdominal ventrites. I. Detail of abdominal apex. J–K. Aedeagus: ventral (J) and dorsal (K)." figureDoi="http://doi.org/10.5281/zenodo.3961801" httpUri="https://zenodo.org/record/3961801/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
A–K
</paragraph>
</subSubSection>
<subSubSection id="A4DE65268E2EFE342C9FFE18419F946D" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="EC7B36AD8E2EFE342C9FFE18428797A8" blockId="14.[189,457,402,428]" box="[189,457,402,428]" pageId="14" pageNumber="15">
<heading id="B73381C18E2EFE342C9FFE18428797A8" bold="true" box="[189,457,402,428]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="DEB0EABF8E2EFE342C9FFE18428797A8" bold="true" box="[189,457,402,428]" pageId="14" pageNumber="15">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="EC7B36AD8E2EFE342C9FFE4B419F946D" blockId="14.[189,1399,449,617]" pageId="14" pageNumber="15">
The species is easily recognized based on its aedeagus which resembles those of
<taxonomicName id="2BC44D2E8E2EFE3428B1FE48460D97DF" authorityName="Jayaswal" authorityYear="1972" box="[1171,1347,449,476]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bhutanicum">
<emphasis id="DEB0EABF8E2EFE3428B1FE48460D97DF" box="[1171,1347,449,476]" italics="true" pageId="14" pageNumber="15">C. bhutanicum</emphasis>
</taxonomicName>
and
<taxonomicName id="2BC44D2E8E2EFE342C9FFE6F427697FA" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[189,312,484,511]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stultum">
<emphasis id="DEB0EABF8E2EFE342C9FFE6F427697FA" box="[189,312,484,511]" italics="true" pageId="14" pageNumber="15">C. stultum</emphasis>
</taxonomicName>
in its apically situated gonopore and parallel-sided median lobe. In contrast to both latter species, the outer margin of the parameres in strongly bisinuate in
<taxonomicName id="2BC44D2E8E2EFE342FF1FD82471F9426" authority="Sheth &amp; Ghate &amp; Fikáček, 2020" authorityName="Sheth &amp; Ghate &amp; Fikáček" authorityYear="2020" box="[979,1105,520,546]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lyratum" status="sp. nov.">
<emphasis id="DEB0EABF8E2EFE342FF1FD82471F9426" box="[979,1105,520,546]" italics="true" pageId="14" pageNumber="15">C. lyratum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C58357C48E2EFE34287BFD8247FB9426" box="[1113,1205,520,546]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(in contrast to a nearly continuously arcuate outer margin in
<taxonomicName id="2BC44D2E8E2EFE342EE9FDA140379441" authorityName="Jayaswal" authorityYear="1972" box="[715,889,555,581]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bhutanicum">
<emphasis id="DEB0EABF8E2EFE342EE9FDA140379441" box="[715,889,555,581]" italics="true" pageId="14" pageNumber="15">C. bhutanicum</emphasis>
</taxonomicName>
and
<taxonomicName id="2BC44D2E8E2EFE342F96FDA147639441" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[948,1069,555,581]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="stultum">
<emphasis id="DEB0EABF8E2EFE342F96FDA147639441" box="[948,1069,555,581]" italics="true" pageId="14" pageNumber="15">C. stultum</emphasis>
</taxonomicName>
) and the abdominal apex is simple, without emargination and stout setae.
</paragraph>
</subSubSection>
<subSubSection id="A4DE65268E2EFE342C9FFD1D475D94DB" box="[189,1043,663,736]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="EC7B36AD8E2EFE342C9FFD1D420A94B5" blockId="14.[189,324,663,689]" box="[189,324,663,689]" pageId="14" pageNumber="15">
<heading id="B73381C18E2EFE342C9FFD1D420A94B5" bold="true" box="[189,324,663,689]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="DEB0EABF8E2EFE342C9FFD1D420A94B5" bold="true" box="[189,324,663,689]" pageId="14" pageNumber="15">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="EC7B36AD8E2EFE342C9FFD4F475D94DB" blockId="14.[189,1043,709,736]" box="[189,1043,709,736]" pageId="14" pageNumber="15">The species name refers to the lyriform shape of the aedeagus; adjective.</paragraph>
</subSubSection>
<caption id="B8BB66258E2EFE342C9FF8D741FF91E5" ID-DOI="http://doi.org/10.5281/zenodo.3961801" ID-Zenodo-Dep="3961801" httpUri="https://zenodo.org/record/3961801/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1885,1911]" targetBox="[189,1379,907,1840]" targetPageId="14">
<paragraph id="EC7B36AD8E2EFE342C9FF8D741FF91E5" blockId="14.[189,1399,1883,2017]" pageId="14" pageNumber="15">
<emphasis id="DEB0EABF8E2EFE342C9FF8D742479173" bold="true" box="[189,265,1885,1911]" pageId="14" pageNumber="15">Fig. 6.</emphasis>
<taxonomicName id="2BC44D2E8E2EFE342D32F8D7411D9173" authority="Sheth &amp; Ghate &amp; Fikáček, 2020" authorityName="Sheth &amp; Ghate &amp; Fikáček" authorityYear="2020" box="[272,595,1885,1912]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lyratum" status="sp. nov.">
<emphasis id="DEB0EABF8E2EFE342D32F8D742D29173" box="[272,412,1885,1911]" italics="true" pageId="14" pageNumber="15">Coelostoma</emphasis>
(s. str.)
<emphasis id="DEB0EABF8E2EFE342DDAF8D7411D9173" box="[504,595,1885,1911]" italics="true" pageId="14" pageNumber="15">lyratum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C58357C48E2EFE342E78F8D441F8917C" box="[602,694,1886,1912]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NMPC).
<emphasis id="DEB0EABF8E2EFE342FE4F8D740959173" bold="true" box="[966,987,1885,1911]" pageId="14" pageNumber="15">A</emphasis>
. Dorsal habitus.
<emphasis id="DEB0EABF8E2EFE342882F8D747FA9173" bold="true" box="[1184,1204,1885,1911]" pageId="14" pageNumber="15">B</emphasis>
. Lateral habitus.
<emphasis id="DEB0EABF8E2EFE342C9FF80A439C919E" bold="true" box="[189,210,1920,1946]" pageId="14" pageNumber="15">C</emphasis>
. Frontal view.
<emphasis id="DEB0EABF8E2EFE342DB2F80A42EB919E" bold="true" box="[400,421,1920,1946]" pageId="14" pageNumber="15">D</emphasis>
. Thorax in ventral view.
<emphasis id="DEB0EABF8E2EFE342EC3F80A41BB919E" bold="true" box="[737,757,1920,1946]" pageId="14" pageNumber="15">E</emphasis>
. Dorsal punctation of head.
<emphasis id="DEB0EABF8E2EFE34287AF80A4724919E" bold="true" box="[1112,1130,1920,1946]" pageId="14" pageNumber="15">F</emphasis>
. Dorsal punctation of pronotum.
<emphasis id="DEB0EABF8E2EFE342D66F829421591B9" bold="true" box="[324,347,1955,1981]" pageId="14" pageNumber="15">G</emphasis>
. Dorsal punctation of elytron.
<emphasis id="DEB0EABF8E2EFE342EC2F82E41B991BA" bold="true" box="[736,759,1956,1982]" pageId="14" pageNumber="15">H</emphasis>
. Abdominal ventrites.
<emphasis id="DEB0EABF8E2EFE342837F82E476E91BA" bold="true" box="[1045,1056,1956,1982]" pageId="14" pageNumber="15">I</emphasis>
. Detail of abdominal apex.
<emphasis id="DEB0EABF8E2EFE342C9FF84D43BF91E5" bold="true" box="[189,241,1991,2017]" pageId="14" pageNumber="15">J–K</emphasis>
. Aedeagus: ventral (J) and dorsal (K).
</paragraph>
</caption>
<subSubSection id="A4DE65268E2FFE352C9FFE8140A7946C" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="EC7B36AD8E2FFE352C9FFE8142229721" blockId="15.[189,364,267,293]" box="[189,364,267,293]" pageId="15" pageNumber="16">
<heading id="B73381C18E2FFE352C9FFE8142229721" bold="true" box="[189,364,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="DEB0EABF8E2FFE352C9FFE8142229721" bold="true" box="[189,364,267,293]" pageId="15" pageNumber="16">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="EC7B36AD8E2FFE352CC8FEB042129750" blockId="15.[189,1398,314,410]" box="[234,348,314,340]" pageId="15" pageNumber="16">
<emphasis id="DEB0EABF8E2FFE352CC8FEB042129750" bold="true" box="[234,348,314,340]" pageId="15" pageNumber="16">
<typeStatus id="337F880F8E2FFE352CC8FEB042129750" box="[234,348,314,340]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FFED740DB979E" blockId="15.[189,1398,314,410]" pageId="15" pageNumber="16">
<materialsCitation id="5CAC3CF08E2FFE352C9FFED740DB979E" ID-GBIF-Occurrence="2818213329" collectedFrom="riverside" collectingDate="2006-05-22" collectionCode="NMPC" collectorName="Z. Kejval" country="India" elevation="40" latitude="15.917701" location="Maharashtra" longLatPrecision="1" longitude="73.81574" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
<collectingCountry id="94D3763D8E2FFE352C9FFED7425E9773" box="[189,272,349,375]" name="India" pageId="15" pageNumber="16">INDIA</collectingCountry>
<emphasis id="DEB0EABF8E2FFE352D35FED7426F9773" bold="true" box="[279,289,349,375]" pageId="15" pageNumber="16">•</emphasis>
<specimenCount id="FAC2FD248E2FFE352D0AFED142719773" box="[296,319,347,375]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
; “
<collectingRegion id="2E00F84F8E2FFE352D75FED742BE9773" box="[343,496,349,375]" country="India" name="Maharashtra" pageId="15" pageNumber="16">Maharashtra</collectingRegion>
,
<locationDeviation id="FDC92F8F8E2FFE352DD8FED740569773" box="[506,792,348,375]" pageId="15" pageNumber="16">
ca.
<quantity id="2B3C9B488E2FFE352E00FED641259773" box="[546,619,348,375]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="km" value="15.0">15 km</quantity>
of Savantvadi
</locationDeviation>
;
<geoCoordinate id="89F0506A8E2FFE352F01FED140CC9773" box="[803,898,347,375]" degrees="15" direction="north" minutes="55" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="15.916667">15º55ʹN</geoCoordinate>
<geoCoordinate id="89F0506A8E2FFE352FABFED140A69773" box="[905,1000,347,375]" degrees="75" direction="east" minutes="53" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="75.88333">75º53ʹE</geoCoordinate>
” [15º55ʹ3.72ʺ N, 73º48ʹ56.67ʺ E]; ca
<elevation id="67E9D19E8E2FFE352CFCFE0A4214979E" box="[222,346,384,410]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="m" value="40.0">
<quantity id="2B3C9B488E2FFE352CFCFE0A4254979E" box="[222,282,384,410]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="m" value="40.0">40 m</quantity>
a.s.l.
</elevation>
;
<date id="987A106D8E2FFE352D45FE0A4149979E" box="[359,519,384,410]" pageId="15" pageNumber="16" value="2006-05-22">
<collectingDate id="883EE9858E2FFE352D45FE0A4149979E" box="[359,519,384,410]" pageId="15" pageNumber="16" value="2006-05-22">22 May 2006</collectingDate>
</date>
;
<collectorName id="4131537B8E2FFE352E31FE0A41CC979E" box="[531,642,384,410]" pageId="15" pageNumber="16">Z. Kejval</collectorName>
leg.;
<collectedFrom id="5FD875508E2FFE352EE6FE0A4060979E" box="[708,814,384,410]" pageId="15" pageNumber="16">riverside</collectedFrom>
;
<collectionCode id="8AD5AE688E2FFE352F18FE0A40DF979E" box="[826,913,384,410]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="15" pageNumber="16">NMPC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="EC7B36AD8E2FFE352CC8FE4B422997DF" blockId="15.[189,1399,449,617]" box="[234,359,449,475]" pageId="15" pageNumber="16">
<emphasis id="DEB0EABF8E2FFE352CC8FE4B422997DF" bold="true" box="[234,359,449,475]" pageId="15" pageNumber="16">
<typeStatus id="337F880F8E2FFE352CC8FE4B422997DF" box="[234,359,449,475]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FFE6F40A7946C" blockId="15.[189,1399,449,617]" pageId="15" pageNumber="16">
<materialsCitation id="5CAC3CF08E2FFE352C9FFE6F470F97FB" ID-GBIF-Occurrence="2818213349" box="[189,1089,484,511]" collectingDate="2006-05-22" collectionCode="NMPC" collectorName="Z. Kejval" country="India" elevation="40" latitude="15.917701" location="Maharashtra" longLatPrecision="1" longitude="73.81574" pageId="15" pageNumber="16" specimenCount="3" stateProvince="Maharashtra" typeStatus="paratype">
<collectingCountry id="94D3763D8E2FFE352C9FFE6F424097FB" box="[189,270,485,511]" name="India" pageId="15" pageNumber="16">INDIA</collectingCountry>
•
<collectingRegion id="2E00F84F8E2FFE352D05FE6E42F497FA" box="[295,442,484,510]" country="India" name="Maharashtra" pageId="15" pageNumber="16">Maharashtra</collectingRegion>
•
<specimenCount id="FAC2FD248E2FFE352DF1FE6F416297FB" box="[467,556,485,511]" pageId="15" pageNumber="16">3 specs</specimenCount>
; same collection data as for holotype;
<collectionCode id="8AD5AE688E2FFE352FCCFE6F470F97FB" box="[1006,1089,485,511]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="15" pageNumber="16">NMPC</collectionCode>
</materialsCitation>
•
<materialsCitation id="5CAC3CF08E2FFE352878FE6F41569426" ID-GBIF-Occurrence="2818213311" collectingDate="2006-05-22" collectionCode="BMNH" collectorName="Z. Kejval" country="India" elevation="40" latitude="15.917701" location="Savantvadi" longLatPrecision="1" longitude="73.81574" pageId="15" pageNumber="16" specimenCount="2" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCount id="FAC2FD248E2FFE352878FE6F47FD97FB" box="[1114,1203,485,511]" pageId="15" pageNumber="16">2 specs</specimenCount>
; same collection data as for holotype;
<collectionCode id="8AD5AE688E2FFE352DE2FD8241569426" box="[448,536,520,546]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="15" pageNumber="16">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="5CAC3CF08E2FFE352E15FD8247FB9426" ID-GBIF-Occurrence="2818213327" box="[567,1205,520,546]" collectingDate="2006-05-22" collectionCode="SMNS" collectorName="Z. Kejval" country="India" elevation="40" latitude="15.917701" location="Savantvadi" longLatPrecision="1" longitude="73.81574" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCount id="FAC2FD248E2FFE352E15FD8241C19426" box="[567,655,520,546]" pageId="15" pageNumber="16" type="generic">1 spec.</specimenCount>
; same collection data as for holotype;
<collectionCode id="8AD5AE688E2FFE352847FD8247FB9426" box="[1125,1205,520,546]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="15" pageNumber="16">SMNS</collectionCode>
</materialsCitation>
•
<materialsCitation id="5CAC3CF08E2FFE3528F1FD8241139441" ID-GBIF-Occurrence="2818213315" collectingDate="2006-05-22" collectionCode="ZSI" collectorName="Z. Kejval" country="India" elevation="40" latitude="15.917701" location="Savantvadi" longLatPrecision="1" longitude="73.81574" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCount id="FAC2FD248E2FFE3528F1FD8246659426" box="[1235,1323,520,546]" pageId="15" pageNumber="16" type="generic">1 spec.</specimenCount>
; same collection data as for holotype;
<collectionCode id="8AD5AE688E2FFE352E13FDA141139441" box="[561,605,555,581]" pageId="15" pageNumber="16">ZSI</collectionCode>
</materialsCitation>
•
<materialsCitation id="5CAC3CF08E2FFE352E54FDA146399441" ID-GBIF-Occurrence="2818213301" box="[630,1399,555,581]" collectingDate="2006-05-22" collectionCode="NMPC" collectorName="Z. Kejval" country="India" elevation="40" latitude="15.917701" location="Savantvadi" longLatPrecision="1" longitude="73.81574" pageId="15" pageNumber="16" specimenCode="NCBS-BL018-019" specimenCount="2" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCount id="FAC2FD248E2FFE352E54FDA1419E9441" box="[630,720,555,581]" pageId="15" pageNumber="16">2 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="BC629ED68E2FFE3528B6FDA146399441" box="[1172,1399,555,581]" pageId="15" pageNumber="16">NCBS-BL018–019</specimenCode>
</materialsCitation>
•
<materialsCitation id="5CAC3CF08E2FFE352CEDFDC440AB946C" ID-GBIF-Occurrence="2818213308" box="[207,997,590,617]" collectingDate="2006-05-22" collectionCode="UASB" collectorName="Z. Kejval" country="India" elevation="40" latitude="15.917701" location="Savantvadi" longLatPrecision="1" longitude="73.81574" pageId="15" pageNumber="16" specimenCode="UASB 01923068-69" specimenCount="2" stateProvince="Maharashtra" typeStatus="paratype">
<specimenCount id="FAC2FD248E2FFE352CEDFDC44266946D" box="[207,296,590,617]" pageId="15" pageNumber="16">2 specs</specimenCount>
; same collection data as for holotype;
<specimenCode id="BC629ED68E2FFE352EC8FDC440AB946C" box="[746,997,590,616]" collectionCode="UASB" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="15" pageNumber="16">UASB 01923068–69</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A4DE65268E2FFE352C9FFD05429291AC" pageId="15" pageNumber="16" type="description">
<paragraph id="EC7B36AD8E2FFE352C9FFD05420094AD" blockId="15.[189,334,655,681]" box="[189,334,655,681]" pageId="15" pageNumber="16">
<heading id="B73381C18E2FFE352C9FFD05420094AD" bold="true" box="[189,334,655,681]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="DEB0EABF8E2FFE352C9FFD05420094AD" bold="true" box="[189,334,655,681]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FFD3541089561" blockId="15.[189,1399,702,869]" pageId="15" pageNumber="16">
FORM AND COLOUR. Body length
<quantity id="2B3C9B488E2FFE352E67FD34419894DD" box="[581,726,702,729]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.7" metricValueMin="4.2" pageId="15" pageNumber="16" unit="mm" value="4.45" valueMax="4.7" valueMin="4.2">4.2–4.7 mm</quantity>
(
<typeStatus id="337F880F8E2FFE352ECBFD34401C94DC" box="[745,850,702,728]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
<quantity id="2B3C9B488E2FFE352F7FFD3440F094DD" box="[861,958,702,729]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="16" unit="mm" value="4.5">4.5 mm</quantity>
), body width
<quantity id="2B3C9B488E2FFE352848FD3447B594DD" box="[1130,1275,702,729]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="15" pageNumber="16" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
(
<typeStatus id="337F880F8E2FFE35292CFD34463994DC" box="[1294,1399,702,728]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
3.0 mm). Body narrowly oval in dorsal view, moderately convex in lateral view. Head black, reddish brown near clypeus and anterior to eyes; pronotum and elytra uniformly dark brown to black with slightly paler margins, ventral surface blackish brown, legs reddish brown. Mouthparts and antennae pale brown, antennal club brown.
</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FFC0747F19237" blockId="15.[189,1399,908,1076]" pageId="15" pageNumber="16">HEAD. Dorsal punctation dense, consisting of simple punctures without associated ridges; trichobothria present; surface between punctures smooth.Anterior margin of clypeus arcuate, indistinctly emarginated medially. Eyes large, interocular distance ca 4.0 × the width of one eye in dorsal view; eye emarginate anteriorly. Labrum moderately sclerotized, largely exposed anterior of clypeus. Antenna with 9 antennomeres, club loosely segmented. Second maxillary palpomere moderately broad.</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FFBD142909305" blockId="15.[189,1399,1114,1281]" pageId="15" pageNumber="16">PROTHORAX. Pronotum bisinuate anteriorly, anterolateral corners obtuse; posterior margin moderately bisinuate, posterolateral corners rectangular.Anterior and lateral margins with distinct bead not extending to posterior margin. Pronotal punctation denser and finer than on head, consisting of simple punctures without associated ridges; surface between punctures smooth. Prosternum straight on anterior margin, weakly carinate mesally.</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FFAA3429193A8" blockId="15.[189,1399,1321,1453]" pageId="15" pageNumber="16">MESOTHORAX. Elytral punctation dense and moderately coarse, consisting of punctures without transverse ridges. Series of impressed punctures absent. Sutural stria impressed, present in apical two thirds. Mesoventral plate as long as wide, arrowhead-shaped, bluntly pointed anteriorly, posteriorly widely attached to metaventrite.</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FFA5E465D9030" blockId="15.[189,1399,1492,1589]" pageId="15" pageNumber="16">METATHORAX. Metaventrite raised medially, median part sparsely pubescent except posterior third and anterior part, which are bare; lateral portions pubescent. Anterior metaventral process narrowly projecting between mesocoxae; posterior process bifid. Wings well-developed (macropterous).</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FF9D6411E909D" blockId="15.[189,1398,1627,1689]" pageId="15" pageNumber="16">LEGS. Profemur with dense pubescence except in apical fifth; mesofemur and metafemur with very sparsely arranged short setae only.</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FF94A412C90F9" blockId="15.[189,1399,1728,1790]" pageId="15" pageNumber="16">ABDOMEN. All ventrites densely pubescent. First ventrite without carina. Posterior margin of last ventrite entire, without stout spines mesally.</paragraph>
<paragraph id="EC7B36AD8E2FFE352C9FF8AF429291AC" blockId="15.[189,1399,1829,1960]" pageId="15" pageNumber="16">
AEDEAGUS (
<figureCitation id="74FF2A288E2FFE352D66F8AF42C69144" box="[324,392,1829,1856]" captionStart="Fig" captionStartId="14.[189,232,1885,1911]" captionTargetBox="[189,1379,907,1840]" captionTargetId="figure@14.[189,1396,890,1847]" captionTargetPageId="14" captionText="Fig. 6. Coelostoma (s. str.) lyratum sp. nov., holotype, ♂ (NMPC). A. Dorsal habitus. B. Lateral habitus. C. Frontal view. D. Thorax in ventral view. E. Dorsal punctation of head. F. Dorsal punctation of pronotum. G. Dorsal punctation of elytron. H. Abdominal ventrites. I. Detail of abdominal apex. J–K. Aedeagus: ventral (J) and dorsal (K)." figureDoi="http://doi.org/10.5281/zenodo.3961801" httpUri="https://zenodo.org/record/3961801/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
J–K).
<quantity id="2B3C9B488E2FFE352DF3F8AF4174913B" box="[465,570,1829,1855]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="15" pageNumber="16" unit="mm" value="0.84">0.84 mm</quantity>
long. Median lobe of nearly the same width throughout; apex roundly cut off; gonopore situated at apex, widely semicircular. Parameres slightly longer than median lobe; heavily tri-sinuate on outer margin; apex acute at inner margin, rounded laterally. Phallobase small, slightly wider than long.
</paragraph>
</subSubSection>
<subSubSection id="A4DE65268E30FE2A2C9FFE8147949757" box="[189,1242,267,340]" pageId="16" pageNumber="17" type="biology_ecology">
<paragraph id="EC7B36AD8E30FE2A2C9FFE8142529721" blockId="16.[189,284,267,293]" box="[189,284,267,293]" pageId="16" pageNumber="17">
<heading id="B73381C18E30FE2A2C9FFE8142529721" bold="true" box="[189,284,267,293]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="DEB0EABF8E30FE2A2C9FFE8142529721" bold="true" box="[189,284,267,293]" pageId="16" pageNumber="17">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="EC7B36AD8E30FE2A2C9FFEB347949757" blockId="16.[189,1242,313,340]" box="[189,1242,313,340]" pageId="16" pageNumber="17">
Unknown. The labels of the
<typeStatus id="337F880F8E30FE2A2E28FEB041739750" box="[522,573,314,340]" pageId="16" pageNumber="17">type</typeStatus>
specimens indicate that they were found at the riverside.
</paragraph>
</subSubSection>
<subSubSection id="A4DE65268E30FE2A2C9FFEF1400A97C7" box="[189,836,379,452]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="EC7B36AD8E30FE2A2C9FFEF142189791" blockId="16.[189,342,379,405]" box="[189,342,379,405]" pageId="16" pageNumber="17">
<heading id="B73381C18E30FE2A2C9FFEF142189791" bold="true" box="[189,342,379,405]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="DEB0EABF8E30FE2A2C9FFEF142189791" bold="true" box="[189,342,379,405]" pageId="16" pageNumber="17">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="EC7B36AD8E30FE2A2C9FFE20400A97C7" blockId="16.[189,836,425,452]" box="[189,836,425,452]" pageId="16" pageNumber="17">
Known from the
<typeStatus id="337F880F8E30FE2A2DA4FE2042F797C0" box="[390,441,426,452]" pageId="16" pageNumber="17">type</typeStatus>
locality in southern
<collectingRegion id="2E00F84F8E30FE2A2E88FE23400F97C7" box="[682,833,425,451]" country="India" name="Maharashtra" pageId="16" pageNumber="17">Maharashtra</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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