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<document id="3107F80BAA3840E22D05A974F831F644" ID-DOI="10.11646/zootaxa.3858.1.1" ID-GBIF-Dataset="d6152690-b0f3-4f6f-9fea-05c3fb53690e" ID-ISSN="1175-5326" ID-Zenodo-Dep="287013" ID-ZooBank="D1CCF02B-4314-4537-A64F-0372715E3F93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460902618942" checkinUser="plazi" docAuthor="Fleming, Alan J., Wood, Monty, Smith, Alex, Hallwachs, Winnie &amp; Janzen, Daniel H." docDate="2014" docId="03D087FFB7588F45FF1AFB3AFADFFCC5" docLanguage="en" docName="zt03858p090.pdf" docOrigin="Zootaxa 3858 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Houghia matarritai Fleming &amp; Wood, sp. nov." docType="treatment" docVersion="7" lastPageNumber="71" masterDocId="FFE9FF87B71C8F03FF8DFF96FFD1FFFD" masterDocTitle="Revision of the New World species of Houghia Coquillett (Diptera, Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, Costa Rica" masterLastPageNumber="90" masterPageNumber="1" pageNumber="69" updateTime="1698360527715" updateUser="plazi">
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<mods:title id="B4A8F95F85C7538DA2137E1A256AA77B">Revision of the New World species of Houghia Coquillett (Diptera, Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, Costa Rica</mods:title>
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<mods:namePart id="78E0F8849640D431CF8B6E7A52CF3E56">Fleming, Alan J.</mods:namePart>
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<mods:namePart id="DE1A749BD1F166406160BF4F734B8152">Wood, Monty</mods:namePart>
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<mods:namePart id="A22C0025E02993D66F3FA3FE06F29E3C">Smith, Alex</mods:namePart>
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<mods:namePart id="1402F3BC7DB6B3B347D5166F7937CCC5">Hallwachs, Winnie</mods:namePart>
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<mods:namePart id="87E206C76CAC7E38AEA408DD0358F1A3">Janzen, Daniel H.</mods:namePart>
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<mods:date id="328EE4954E11D4EA8F7320FDFF3B6386">2014</mods:date>
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<treatment id="03D087FFB7588F45FF1AFB3AFADFFCC5" ID-DOI="http://doi.org/10.5281/zenodo.5695648" ID-GBIF-Taxon="127669713" ID-Zenodo-Dep="5695648" LSID="urn:lsid:plazi:treatment:03D087FFB7588F45FF1AFB3AFADFFCC5" httpUri="http://treatment.plazi.org/id/03D087FFB7588F45FF1AFB3AFADFFCC5" lastPageId="70" lastPageNumber="71" pageId="68" pageNumber="69">
<subSubSection id="C3636562B7588F47FF1AFB3AFEC0FB1A" pageId="68" pageNumber="69" type="nomenclature">
<paragraph id="8BC636E9B7588F47FF1AFB3AFD1CFB3B" blockId="68.[151,717,1196,1255]" box="[151,717,1196,1222]" pageId="68" pageNumber="69">
<heading id="D08E8185B7588F47FF1AFB3AFD1CFB3B" bold="true" box="[151,717,1196,1222]" fontSize="11" level="1" pageId="68" pageNumber="69" reason="1">
<emphasis id="B90DEAFBB7588F47FF1AFB3AFD1CFB3B" bold="true" box="[151,717,1196,1222]" pageId="68" pageNumber="69">
<taxonomicName id="4C794D6AB7588F47FF1AFB3AFDB2FB3B" ID-CoL="9HJ9X" authority="Fleming &amp; Wood" authorityName="Fleming &amp; Wood" box="[151,611,1196,1222]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="matarritai" status="sp. nov.">
<emphasis id="B90DEAFBB7588F47FF1AFB3AFE52FB3B" bold="true" box="[151,387,1196,1222]" italics="true" pageId="68" pageNumber="69">Houghia matarritai</emphasis>
Fleming &amp; Wood
</taxonomicName>
,
<taxonomicNameLabel id="A23E5780B7588F47FDFDFB3AFD1CFB3B" box="[624,717,1196,1222]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC636E9B7588F47FF1AFB58FEC0FB1A" blockId="68.[151,717,1196,1255]" box="[151,273,1230,1255]" pageId="68" pageNumber="69">
<figureCitation id="13422A6CB7588F47FF1AFB58FF37FB1A" box="[151,230,1230,1255]" captionStart="FIGURE 34" captionStartId="69.[151,250,1593,1615]" captionTargetBox="[340,1246,445,1572]" captionTargetId="figure@69.[340,1247,445,1572]" captionTargetPageId="69" captionText="FIGURE 34. General morphology of Houghia matarritai sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287047/files/figure.png" pageId="68" pageNumber="69">Fig. 34</figureCitation>
af
</paragraph>
</subSubSection>
<subSubSection id="C3636562B7588F47FF1AFA80FC55FA1E" pageId="68" pageNumber="69" type="diagnosis">
<paragraph id="8BC636E9B7588F47FF1AFA80FC55FA1E" blockId="68.[151,1437,1302,2014]" pageId="68" pageNumber="69">
<emphasis id="B90DEAFBB7588F47FF1AFA80FEC0FAD2" bold="true" box="[151,273,1302,1327]" pageId="68" pageNumber="69">Diagnosis.</emphasis>
<taxonomicName id="4C794D6AB7588F47FE90FA80FE2FFAD3" box="[285,510,1302,1327]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="matarritai">
<emphasis id="B90DEAFBB7588F47FE90FA80FE2FFAD3" box="[285,510,1302,1327]" italics="true" pageId="68" pageNumber="69">Houghia matarritai</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C794D6AB7588F47FD31FA81FCE6FAD2" box="[700,823,1302,1327]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="blancoi">
<emphasis id="B90DEAFBB7588F47FD31FA81FCE6FAD2" box="[700,823,1302,1327]" italics="true" pageId="68" pageNumber="69">H. blancoi</emphasis>
</taxonomicName>
group (
<taxonomicName id="4C794D6AB7588F47FC14FA81FBC5FAD2" box="[921,1044,1302,1327]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="blancoi">
<emphasis id="B90DEAFBB7588F47FC14FA81FBC5FAD2" box="[921,1044,1302,1327]" italics="true" pageId="68" pageNumber="69">H. blancoi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794D6AB7588F47FBAAFA81FB69FAD3" box="[1063,1208,1303,1326]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="romeroae">
<emphasis id="B90DEAFBB7588F47FBAAFA81FB69FAD3" box="[1063,1208,1303,1326]" italics="true" pageId="68" pageNumber="69">H. romeroae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794D6AB7588F47FB41FA81FABFFAD2" box="[1228,1390,1302,1327]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="chavarriae">
<emphasis id="B90DEAFBB7588F47FB41FA81FABFFAD2" box="[1228,1390,1302,1327]" italics="true" pageId="68" pageNumber="69">H. chavarriae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794D6AB7588F47FA0CFA81FF33FAAE" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="marini">
<emphasis id="B90DEAFBB7588F47FA0CFA81FF33FAAE" italics="true" pageId="68" pageNumber="69">H. marini</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C794D6AB7588F47FEAEFAAAFE6BFAAE" box="[291,442,1340,1363]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="matarritai">
<emphasis id="B90DEAFBB7588F47FEAEFAAAFE6BFAAE" box="[291,442,1340,1363]" italics="true" pageId="68" pageNumber="69">H. matarritai</emphasis>
</taxonomicName>
). It has few distinctive features except in the male terminalia and the presence of gold tomentosity covering in excess of 50% of the fronto-orbital plate. The eye is bare and the antennae are entirely black. The ground colour of the abdomen is black, and sex patches are confined to tergites 4 and 5. Gold tomentosity extends from the vertex to lower fronto-orbital setae, but the apex of the first flagellomere falls short of the lower facial margin by half the length of the pedicel (
<figureCitation id="13422A6CB7588F47FC98FA5CFCB8FA1E" box="[789,873,1482,1507]" captionStart="FIGURE 34" captionStartId="69.[151,250,1593,1615]" captionTargetBox="[340,1246,445,1572]" captionTargetId="figure@69.[340,1247,445,1572]" captionTargetPageId="69" captionText="FIGURE 34. General morphology of Houghia matarritai sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287047/files/figure.png" pageId="68" pageNumber="69">Fig. 34</figureCitation>
b).
</paragraph>
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<subSubSection id="C3636562B7588F46FF4AFA78FCD6F8C3" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="description">
<paragraph id="8BC636E9B7588F46FF4AFA78FF02FE7A" blockId="68.[151,1437,1302,2014]" lastBlockId="69.[151,1436,151,391]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
<emphasis id="B90DEAFBB7588F47FF4AFA78FE88F9FA" bold="true" box="[199,345,1518,1543]" pageId="68" pageNumber="69">Description.</emphasis>
Male. Antenna black. When viewed in profile, antenna arises distinctly above middle of eye. Length of first flagellomere almost extending to facial margin, (usually shorter by less than length of pedicel). Facial ridge bare except for a few (usually 35) decumbent small setae above vibrissa. Palpus pale, usually distinctly yellowish. Postgena behind postoccipital row, above level of lower facial margin, with a small patch of few black setae. Parafacial silver. Colour of fronto-orbital plate pale brassy to gold on its entire length from vertex to base of antenna (more than 50% coverage). Surface of fronto-orbital plate covered with small recumbent hairs, especially near margin of eye. Ocellar triangle appearing rounded anteriorly when viewed from above. Diameter of anterior ocellus less than diameter of base of adjacent ocellar seta. Ocellar setae arising beside, or slightly in front of, anterior ocellus. Eye with inconspicuous hairs. Postpronotum with 4 or 5 postpronotal setae. Dark stripes on either side of dorsocentral row of setae separated from one another by yellow tomentosity. Median and lateral stripes on either side of scutum separate from each other posteriorly. Postsutural dorsocentral setae 4. Anterior quadrant of anepisternum covered with short setae except for usually 3 to 5 distinctly larger setae. Katepisternum with three setae, the middle one always the smallest. Vein R1 bare dorsally. Legs ranging from reddish brown to yellow tinged but overall dark. Coxae dark usually concolourous with remainder of leg. Ground colour of dorsal surface of abdomen dark to black. Ground colour of ventral surface of abdomen entirely black. Sex patches present on tergites 4 and 5. Ground colour of sex patches shiny black. Terminalia: surstylus bayonet shaped, posterodorsal half bare, apex bearing many stout apical spines, tip with light inwardly apical curve when viewed dorsally. Cerci rounded, apex with blunt, hooked tip, ventral surface haired, separation between cerci deep narrow V shape, up to 85% as long as surstylus. Lobe of sternite 5 small and pointed apically, inner margin covered in dense tomentosity appearing darker than surrounding cuticle, internal edge inwardly curved, single long apical seta emanating from apex.
</paragraph>
<caption id="DF066661B7598F46FF1AF9AFFAE6F993" httpUri="https://zenodo.org/record/287047/files/figure.png" pageId="69" pageNumber="70" targetBox="[340,1246,445,1572]" targetPageId="69">
<paragraph id="8BC636E9B7598F46FF1AF9AFFAE6F993" blockId="69.[151,1437,1593,1646]" pageId="69" pageNumber="70">
<emphasis id="B90DEAFBB7598F46FF1AF9AFFECEF9B2" bold="true" box="[151,287,1593,1615]" pageId="69" pageNumber="70">FIGURE 34.</emphasis>
General morphology of
<taxonomicName id="4C794D6AB7598F46FD9AF9AFFD0CF9B2" box="[535,733,1593,1615]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="matarritai" status="sp. nov.">
<emphasis id="B90DEAFBB7598F46FD9AF9AFFD0CF9B2" box="[535,733,1593,1615]" italics="true" pageId="69" pageNumber="70">Houghia matarritai</emphasis>
</taxonomicName>
<emphasis id="B90DEAFBB7598F46FD6EF9AFFC9DF9B2" bold="true" box="[739,844,1593,1615]" pageId="69" pageNumber="70">
<taxonomicNameLabel id="A23E5780B7598F46FD6EF9AFFCE2F9B2" box="[739,819,1593,1615]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
a.
</emphasis>
Dorsal habitus.
<emphasis id="B90DEAFBB7598F46FC7CF9AFFBD4F9B2" bold="true" box="[1009,1029,1593,1615]" pageId="69" pageNumber="70">b.</emphasis>
Frontal view of head.
<emphasis id="B90DEAFBB7598F46FB6AF9AFFB28F9B2" bold="true" box="[1255,1273,1593,1615]" pageId="69" pageNumber="70">c.</emphasis>
Lateral habitus.
<emphasis id="B90DEAFBB7598F46FF1AF9CEFF7AF993" bold="true" box="[151,171,1624,1646]" pageId="69" pageNumber="70">d.</emphasis>
Posterior view of male terminalia.
<emphasis id="B90DEAFBB7598F46FD9CF9CEFDF3F993" bold="true" box="[529,546,1624,1646]" pageId="69" pageNumber="70">e.</emphasis>
Lateral view of male terminalia.
<emphasis id="B90DEAFBB7598F46FCF9F9CEFC53F993" bold="true" box="[884,898,1624,1646]" pageId="69" pageNumber="70">f.</emphasis>
Ventral view of sternite 5 + segments 7+8.
</paragraph>
</caption>
<paragraph id="8BC636E9B7598F46FF4AF903FCD6F8C3" blockId="69.[151,1436,1685,2034]" pageId="69" pageNumber="70">
<emphasis id="B90DEAFBB7598F46FF4AF903FEC0F953" bold="true" box="[199,273,1685,1710]" pageId="69" pageNumber="70">Hosts.</emphasis>
<taxonomicName id="4C794D6AB7598F46FE92F903FDD3F950" box="[287,514,1685,1710]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="matarritai">
<emphasis id="B90DEAFBB7598F46FE92F903FDD3F950" box="[287,514,1685,1710]" italics="true" pageId="69" pageNumber="70">Houghia matarritai</emphasis>
</taxonomicName>
has been reared 60 times, from caterpillars of 4 species of monocot-eating
<taxonomicName id="4C794D6AB7598F46FF1AF92FFEFFF92F" box="[151,302,1721,1746]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
(
<taxonomicName id="4C794D6AB7598F46FECDF92FFDCAF92F" authority="Westwood" authorityName="Westwood" box="[320,539,1721,1746]" class="Insecta" family="Nymphalidae" genus="Pierella" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEAFBB7598F46FECDF92FFE48F92F" box="[320,409,1721,1746]" italics="true" pageId="69" pageNumber="70">Pierella</emphasis>
Westwood
</taxonomicName>
,
<taxonomicName id="4C794D6AB7598F46FDAAF92DFD33F92F" authority="Hubner" authorityName="Hubner" box="[551,738,1721,1746]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEAFBB7598F46FDAAF92DFD52F92F" box="[551,643,1723,1746]" italics="true" pageId="69" pageNumber="70">Taygetis</emphasis>
Hübner
</taxonomicName>
) and 14 species of monocot-eating hesperiine
<taxonomicName id="4C794D6AB7598F46FB7AF92FFAACF92F" box="[1271,1405,1721,1746]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="family">Hesperiidae</taxonomicName>
in ACG dry forest and rain forest, out of a sample of 4000+ caterpillars. There is one exception: the single rearing from
<taxonomicName id="4C794D6AB7598F46FF5AF895FD2AF8E7" authority="Chacon" authorityName="Chacon" box="[215,763,1793,1818]" class="Insecta" family="Notodontidae" genus="Dunama" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="jessiebarronae">
<emphasis id="B90DEAFBB7598F46FF5AF895FE37F8E7" box="[215,486,1793,1818]" italics="true" pageId="69" pageNumber="70">Dunama jessiebarronae</emphasis>
Chacón (Notodontidae)
</taxonomicName>
was from a monocot-eating caterpillar. It has been reared only from this ecological grouping of ACG caterpillars.
</paragraph>
</subSubSection>
<subSubSection id="C3636562B7598F45FF4AF8DFFE7CFD55" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="8BC636E9B7598F46FF4AF8DFFA45F87B" blockId="69.[151,1436,1685,2034]" pageId="69" pageNumber="70">
<emphasis id="B90DEAFBB7598F46FF4AF8DFFEEAF89F" bold="true" box="[199,315,1865,1890]" pageId="69" pageNumber="70">
<typeStatus id="54C2884BB7598F46FF4AF8DFFEE7F89F" box="[199,310,1865,1890]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
.
</emphasis>
♂,
<collectionCode id="ED68AE2CB7598F46FEE2F8DCFE7DF89F" box="[367,428,1866,1890]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="69" pageNumber="70">CNC</collectionCode>
.
<typeStatus id="54C2884BB7598F46FE31F8DCFE22F89F" box="[444,499,1866,1890]" pageId="69" pageNumber="70">Type</typeStatus>
locality:
<collectingCountry id="F36E7679B7598F46FDEAF8DCFD38F89F" box="[615,745,1865,1890]" name="Costa Rica" pageId="69" pageNumber="70">Costa Rica</collectingCountry>
, Area de Conservación Guanacaste, Prov. Alajuela, Sector Rincon Rain Forest, Sendero Venado (10.89678°, -85.27001°),
<quantity id="4C819B0CB7598F46FCD0F8F8FC77F878" box="[861,934,1902,1926]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="69" pageNumber="70" unit="m" value="420.0">420 m</quantity>
,
<date id="FFC71029B7598F46FC3CF8F8FBE5F87B" box="[945,1076,1902,1926]" pageId="69" pageNumber="70" value="2008-08-12">12/08/2008</date>
, Jose Perez, DHJPAR0030019.
</paragraph>
<paragraph id="8BC636E9B7598F45FF4AF807FE7CFD55" blockId="69.[151,1436,1685,2034]" lastBlockId="70.[151,1436,151,824]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
<emphasis id="B90DEAFBB7598F46FF4AF807FE97F857" bold="true" box="[199,326,1937,1962]" pageId="69" pageNumber="70">
<typeStatus id="54C2884BB7598F46FF4AF807FE90F857" box="[199,321,1937,1962]" pageId="69" pageNumber="70" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D7FFD60B7598F46FEDEF804FE45F854" box="[339,404,1938,1962]" pageId="69" pageNumber="70" type="generic">70 ♂</specimenCount>
, 74 ♀ (
<collectionCode id="ED68AE2CB7598F46FE76F804FDE9F857" box="[507,568,1938,1962]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="69" pageNumber="70">CNC</collectionCode>
)
<collectingCountry id="F36E7679B7598F46FDC0F804FD01F857" box="[589,720,1937,1962]" name="Costa Rica" pageId="69" pageNumber="70">Costa Rica</collectingCountry>
, Prov. Guanacaste, ACG database codes: DHJPAR0008557, DHJPAR0011527, DHJPAR0029799, DHJPAR0019631, DHJPAR0016604, DHJPAR0008527, DHJPAR0014998, DHJPAR0016344, DHJPAR0011526, DHJPAR0011514, DHJPAR0011518, DHJPAR0011523, DHJPAR0016614, DHJPAR0008169, DHJPAR0008126, DHJPAR0008161, DHJPAR0042705, DHJPAR0008509, DHJPAR0006596, DHJPAR0011517, DHJPAR0048515, DHJPAR0011521, DHJPAR0008166, DHJPAR0008160, DHJPAR0008165, DHJPAR0008508, DHJPAR0008551, DHJPAR0008171, DHJPAR0030021, DHJPAR0008172, DHJPAR0008532, DHJPAR0027860, DHJPAR0008531, DHJPAR0011473, DHJPAR0006947, DHJPAR0014982, DHJPAR0010224, DHJPAR0017146, DHJPAR0011507, DHJPAR0007121, DHJPAR0024543, DHJPAR0011524, DHJPAR0008118, DHJPAR0011535, DHJPAR0016111, DHJPAR0008528, DHJPAR0011522, DHJPAR0011519, DHJPAR0016492, DHJPAR0008167, DHJPAR0008119, DHJPAR0008164, DHJPAR0048515, DHJPAR0011521, DHJPAR0011520, DHJPAR0008163, DHJPAR0008173, DHJPAR0008524, DHJPAR0008525, DHJPAR0010011, DHJPAR0008170, DHJPAR0008535, DHJPAR0011481, DHJPAR0008111, DHJPAR0011516, DHJPAR0008168, 05-SRNP-42616, 05-SRNP-42567, 05-SRNP-42576, 05-SRNP-42138, 05-SRNP-42142, 04-SRNP-40623, 06-SRNP-41954, 04- SRNP-41183, 06-SRNP-32172, 05-SRNP-42140, 05-SRNP-42460, 04-SRNP-60960, 05-SRNP-42575, 05-SRNP- 42424, 05-SRNP-42427, 05-SRNP-42461, 04-SRNP-14343, 03-SRNP-12350.1, 04-SRNP-40001, 04-SRNP- 60760, 05-SRNP-42138, 02-SRNP-42682, 04-SRNP-60438, 01-SRNP-17731, 05-SRNP-42141, 05-SRNP-42579, 04-SRNP-40624, 07-SRNP-41428, 06-SRNP-90, 06-SRNP-43579, 06-SRNP-41954, 06-SRNP-47507, 07-SRNP- 66183, 05-SRNP-42425.
</paragraph>
</subSubSection>
<subSubSection id="C3636562B75A8F45FF4AFD22FD33FCE8" pageId="70" pageNumber="71" type="etymology">
<paragraph id="8BC636E9B75A8F45FF4AFD22FD33FCE8" blockId="70.[151,1436,151,824]" pageId="70" pageNumber="71">
<emphasis id="B90DEAFBB75A8F45FF4AFD22FE9FFD30" bold="true" box="[199,334,692,717]" pageId="70" pageNumber="71">Etymology.</emphasis>
<taxonomicName id="4C794D6AB75A8F45FED2FD22FD97FD31" box="[351,582,692,717]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="matarritai">
<emphasis id="B90DEAFBB75A8F45FED2FD22FD97FD31" box="[351,582,692,717]" italics="true" pageId="70" pageNumber="71">Houghia matarritai</emphasis>
</taxonomicName>
is dedicated to Federico Matarrita of
<collectingCountry id="F36E7679B75A8F45FBBAFD22FB59FD30" box="[1079,1160,692,717]" name="Liberia" pageId="70" pageNumber="71">Liberia</collectingCountry>
in recognition of his development of the ACG web site (http://www.acguanacaste.ac.cr) as an encyclopedia of ACG history and activities, including the inventory of these tachinids.
</paragraph>
</subSubSection>
<subSubSection id="C3636562B75A8F45FF4AFC89FADFFCC5" box="[199,1294,799,824]" pageId="70" pageNumber="71" type="distribution">
<paragraph id="8BC636E9B75A8F45FF4AFC89FADFFCC5" blockId="70.[151,1436,151,824]" box="[199,1294,799,824]" pageId="70" pageNumber="71">
<emphasis id="B90DEAFBB75A8F45FF4AFC89FEB1FCC5" bold="true" box="[199,352,799,824]" pageId="70" pageNumber="71">Distribution.</emphasis>
<collectingCountry id="F36E7679B75A8F45FEEAFCB6FE35FCC5" box="[359,484,799,824]" name="Costa Rica" pageId="70" pageNumber="71">Costa Rica</collectingCountry>
, ACG, Prov. Guanacaste, dry forest and rain forest,
<quantity id="4C819B0CB75A8F45FBA5FC89FB48FCCA" box="[1064,1177,799,824]" metricMagnitude="2" metricUnit="m" metricValue="3.035" metricValueMax="5.9" metricValueMin="0.16999999999999998" pageId="70" pageNumber="71" unit="m" value="303.5" valueMax="590.0" valueMin="17.0">17590 m</quantity>
elevation.
</paragraph>
</subSubSection>
</treatment>
</document>