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<document id="E2AE4F95B31103D90E8119C99D79502C" ID-CLB-Dataset="46269" ID-DOI="10.11646/zootaxa.3774.4.6" ID-GBIF-Dataset="59798127-2d8c-4259-8adf-faa2356239a1" ID-ISSN="1175-5326" ID-Zenodo-Dep="230940" ID-ZooBank="CB88DDD0-BBFE-4E06-8E89-6B4FA4E5BF1B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460393583345" checkinUser="plazi" docAuthor="Grismer, L. Lee, Wood, Perry L., Onn, Chan Kin, Anuar, Shahrul &amp; Muin, Mohd Abdul" docDate="2014" docId="03D087F5EA4BCF50FF21FA1DFC9BCA7D" docLanguage="en" docName="zt03774p394.pdf" docOrigin="Zootaxa 3774 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus metropolis Grismer, Wood, Onn, Anuar &amp; Muin, 2014, new species" docType="treatment" docVersion="10" lastPageNumber="391" masterDocId="FFE9FF8DEA4DCF5BFFB6FFC1FFE0CB3D" masterDocTitle="Cyrts in the city: A new Bent-toed Gecko (Genus Cyrtodactylus) is the only endemic species of vertebrate from Batu Caves, Selangor, Peninsular Malaysia" masterLastPageNumber="394" masterPageNumber="381" pageNumber="387" updateTime="1698298495441" updateUser="ExternalLinkService">
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<mods:title id="511C84ACECCF8EAD9CE2DA1F0DF71EA9">Cyrts in the city: A new Bent-toed Gecko (Genus Cyrtodactylus) is the only endemic species of vertebrate from Batu Caves, Selangor, Peninsular Malaysia</mods:title>
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<mods:name id="F5E5CF9E2DE4D458696FC020AFBD0DAF" type="personal">
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<mods:namePart id="A9429B0CFD42D77B98F748B3A11AD111">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="7F2A97A747A03A4639D1F1E3A2264ED8">Wood, Perry L.</mods:namePart>
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<mods:namePart id="E5D9CD716FC791D55FDE19C8782770C3">Onn, Chan Kin</mods:namePart>
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<mods:namePart id="AFD79AE025E8F5CE9A7929000EC8B9B6">Anuar, Shahrul</mods:namePart>
</mods:name>
<mods:name id="F4CC97AD5C967E73F43FD43AF6627DD4" type="personal">
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<mods:roleTerm id="B5670385BC06137A905AD7612D28A6D8">Author</mods:roleTerm>
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<mods:namePart id="B4C4687AB6ACD896ED27C25CFF1E30B0">Muin, Mohd Abdul</mods:namePart>
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<mods:title id="EBF4586CF862BE34B9CB999AE57163D3">Zootaxa</mods:title>
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<mods:date id="52047EA89EDAAD7EDE0D74E8CFE411DF">2014</mods:date>
<mods:detail id="7DE5B588A147A69395C72244E7F7A74D" type="volume">
<mods:number id="DC0A271615250C344FE78B1164081463">3774</mods:number>
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<mods:number id="7BA137DC342A491B27195E25BF0CE219">4</mods:number>
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<mods:identifier id="BE95EF9CF3496D6798E64E369CC49991" type="DOI">10.11646/zootaxa.3774.4.6</mods:identifier>
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<treatment id="03D087F5EA4BCF50FF21FA1DFC9BCA7D" ID-DOI="http://doi.org/10.5281/zenodo.5616032" ID-GBIF-Taxon="119417473" ID-Zenodo-Dep="5616032" LSID="urn:lsid:plazi:treatment:03D087F5EA4BCF50FF21FA1DFC9BCA7D" httpUri="http://treatment.plazi.org/id/03D087F5EA4BCF50FF21FA1DFC9BCA7D" lastPageId="11" lastPageNumber="391" pageId="6" pageNumber="387">
<subSubSection id="C3636568EA4BCF5DFF21FA1DFF0BCD01" pageId="6" pageNumber="387" type="nomenclature">
<paragraph id="8BC636E3EA4BCF5DFF21FA1DFDBACECA" blockId="6.[151,602,1500,1597]" box="[151,602,1500,1527]" pageId="6" pageNumber="387">
<heading id="D08E818FEA4BCF5DFF21FA1DFDBACECA" bold="true" box="[151,602,1500,1527]" fontSize="11" level="1" pageId="6" pageNumber="387" reason="1">
<emphasis id="B90DEAF1EA4BCF5DFF21FA1DFDBACECA" bold="true" box="[151,602,1500,1527]" pageId="6" pageNumber="387">
<taxonomicName id="4C794D60EA4BCF5DFF21FA1DFE23CECB" box="[151,451,1500,1526]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA4BCF5DFF21FA1DFE23CECB" bold="true" box="[151,451,1500,1526]" italics="true" pageId="6" pageNumber="387">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA4BCF5DFE7DFA1CFDBACECA" box="[459,602,1501,1527]" pageId="6" pageNumber="387" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC636E3EA4BCF5DFF21F9C1FF0BCD01" blockId="6.[151,602,1500,1597]" pageId="6" pageNumber="387">
Batu Caves Bent-toed Gecko (
<figureCitation id="13422A66EA4BCF5DFF29F9E5FF02CD01" box="[159,226,1572,1597]" captionStart="FIGURE 3" captionStartId="8.[151,250,1301,1323]" captionTargetBox="[151,1436,176,1279]" captionTargetId="figure@8.[151,1436,169,1279]" captionTargetPageId="8" captionText="FIGURE 3. Cyrtodactylus metropolis sp. nov. Upper; holotype LSUHC 11347. Lower left; holotype LSUHC 11343. Lower right; paratype LSUHC 11342." httpUri="https://zenodo.org/record/230943/files/figure.png" pageId="6" pageNumber="387">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3636568EA4BCF5DFF21F9ADFD9FCD9D" pageId="6" pageNumber="387" type="reference_group">
<paragraph id="8BC636E3EA4BCF5DFF21F9ADFD9FCD9D" blockId="6.[151,840,1644,1696]" pageId="6" pageNumber="387">
<treatmentCitationGroup id="AB6911CDEA4BCF5DFF21F9ADFD9FCD9D" pageId="6" pageNumber="387">
<taxonomicName id="4C794D60EA4BCF5DFF21F9ADFD9ACDBF" authority="Boulenger 1903:148" authorityName="Boulenger" authorityPageNumber="148" authorityYear="1903" box="[151,634,1644,1666]" class="Reptilia" family="Phyllodactylidae" genus="Gymnodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="B90DEAF1EA4BCF5DFF21F9ADFE43CDBF" box="[151,419,1644,1666]" italics="true" pageId="6" pageNumber="387">Gymnodactylus pulchellus</emphasis>
<treatmentCitation id="0AD810F2EA4BCF5DFE1FF9ADFD9ACDBF" author="Boulenger" box="[425,634,1644,1666]" page="148" pageId="6" pageNumber="387" year="1903">
<bibRefCitation id="EFE84B12EA4BCF5DFE1FF9ADFDB0CDBF" author="Boulenger" box="[425,592,1644,1666]" pageId="6" pageNumber="387" refString="Boulenger, G. A. (1903) Report on the Batrachians and Reptiles. In: Annandale, N. &amp; Robinson, H. C. (Eds.), Fasciculi Malayensis, anthropological and zoological results of an expedition to Perak and the Siamese Malay States, 1901 - 1902, undertaken by Nelson Annandale and Herbert C Robinson. Zoology, Part I. Williams and Norgate, London, pp. 131 - 176" type="book chapter" year="1903">Boulenger 1903</bibRefCitation>
:148
</treatmentCitation>
</taxonomicName>
,
<treatmentCitation id="0AD810F2EA4BCF5DFD33F9ADFCA8CDBF" author="Boulenger" box="[645,840,1644,1666]" page="37" pageId="6" pageNumber="387" year="1912">
<bibRefCitation id="EFE84B12EA4BCF5DFD33F9ADFCCBCDBF" author="Boulenger" box="[645,811,1644,1666]" pageId="6" pageNumber="387" refString="Boulenger, G. A. (1912) A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Reptilia and Batrachia. Taylor &amp; Francis, London, 294 pp." type="book" year="1912">Boulenger 1912</bibRefCitation>
:37
</treatmentCitation>
<taxonomicName id="4C794D60EA4BCF5DFF21F94BFEC5CD9D" box="[151,293,1674,1696]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="387" phylum="Chordata" rank="genus">
<emphasis id="B90DEAF1EA4BCF5DFF21F94BFEC5CD9D" box="[151,293,1674,1696]" italics="true" pageId="6" pageNumber="387">Cyrtodactylus</emphasis>
</taxonomicName>
sp.
<treatmentCitation id="0AD810F2EA4BCF5DFEF8F94BFD9FCD9D" author="Moseley" box="[334,639,1674,1696]" page="79" pageId="6" pageNumber="387" year="2012">
<bibRefCitation id="EFE84B12EA4BCF5DFEF8F94BFD81CD9D" author="Moseley" box="[334,609,1674,1696]" pageId="6" pageNumber="387" refString="Moseley, M., Lim, T. W. &amp; Lim, T. T. (2012) Fauna reported from Batu caves, Selangor, Malaysia: annotated checklist and bibliography. Cave and Karst Science, 39, 77 - 92." type="journal article" year="2012">Moseley, Lim &amp; Lim 2012</bibRefCitation>
:79
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3636568EA4BCF5DFF21F90CFAB4CC6F" pageId="6" pageNumber="387" type="materials_examined">
<paragraph id="8BC636E3EA4BCF5DFF21F90CFE55CC13" blockId="6.[151,1436,1741,2018]" pageId="6" pageNumber="387">
<typeStatus id="54C28841EA4BCF5DFF21F90CFEE4CDD8" box="[151,260,1741,1765]" pageId="6" pageNumber="387" type="holotype">
<emphasis id="B90DEAF1EA4BCF5DFF21F90CFEE4CDD8" bold="true" box="[151,260,1741,1765]" pageId="6" pageNumber="387">Holotype</emphasis>
</typeStatus>
. Adult male (
<collectionCode id="ED68AE26EA4BCF5DFE2AF90FFE1ACDDB" box="[412,506,1742,1766]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="6" pageNumber="387">LSUHC</collectionCode>
11347) from base of the Batu Caves massif east of Temple Cave, Gombak District, Selangor, Peninsular
<collectingCountry id="F36E7673EA4BCF5DFE27F930FE18CC37" box="[401,504,1777,1802]" name="Malaysia" pageId="6" pageNumber="387">Malaysia</collectingCountry>
(3°14’.54”
<geoCoordinate id="EE4D5024EA4BCF5DFDCEF930FCA4CC34" box="[632,836,1777,1802]" direction="north" orientation="latitude" pageId="6" pageNumber="387" precision="1" value="101.68396">
N
<geoCoordinate id="EE4D5024EA4BCF5DFD2BF930FCA4CC34" box="[669,836,1777,1801]" direction="east" orientation="longitude" pageId="6" pageNumber="387" precision="1" value="101.68396">101°41’02.24”</geoCoordinate>
</geoCoordinate>
E;
<quantity id="4C819B06EA4BCF5DFCDFF930FC4ACC34" box="[873,938,1777,1802]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="387" unit="m" value="75.0">75 m</quantity>
) collected by Chan Kin Onn and L. Lee Grismer on
<date id="FFC71023EA4BCF5DFEAAF8D4FE51CC13" box="[284,433,1813,1838]" pageId="6" pageNumber="387" value="2013-07-16">16 July 2013</date>
.
</paragraph>
<paragraph id="8BC636E3EA4BCF5DFF71F8F8FAB4CC6F" blockId="6.[151,1436,1741,2018]" box="[199,1364,1849,1874]" pageId="6" pageNumber="387">
<emphasis id="B90DEAF1EA4BCF5DFF71F8F8FEA5CC6C" bold="true" box="[199,325,1849,1873]" pageId="6" pageNumber="387">
<typeStatus id="54C28841EA4BCF5DFF71F8F8FEDFCC6C" box="[199,319,1849,1873]" pageId="6" pageNumber="387" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Female
<typeStatus id="54C28841EA4BCF5DFE10F8FBFDF1CC6F" box="[422,529,1850,1874]" pageId="6" pageNumber="387" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED68AE26EA4BCF5DFD96F8F8FD9ECC6C" box="[544,638,1849,1873]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="6" pageNumber="387">LSUHC</collectionCode>
11342–43, 11345) bear the same collection data as the
<typeStatus id="54C28841EA4BCF5DFB5CF8F8FAB0CC6F" box="[1258,1360,1849,1874]" pageId="6" pageNumber="387" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3636568EA4BCF52FF71F89DFEC4CAC8" lastPageId="9" lastPageNumber="390" pageId="6" pageNumber="387" type="diagnosis">
<paragraph id="8BC636E3EA4BCF5CFF71F89DFC95CA7D" blockId="6.[151,1436,1741,2018]" lastBlockId="7.[151,1436,151,320]" lastPageId="7" lastPageNumber="388" pageId="6" pageNumber="387">
<emphasis id="B90DEAF1EA4BCF5DFF71F89DFEA1CC48" bold="true" box="[199,321,1884,1909]" pageId="6" pageNumber="387">Diagnosis.</emphasis>
<taxonomicName id="4C794D60EA4BCF5DFEFAF89CFD8ACC48" box="[332,618,1885,1909]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA4BCF5DFEFAF89CFD8ACC48" box="[332,618,1885,1909]" italics="true" pageId="6" pageNumber="387">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA4BCF5DFDC2F89CFD31CC48" box="[628,721,1885,1909]" pageId="6" pageNumber="387" rank="species">
<emphasis id="B90DEAF1EA4BCF5DFDC2F89CFD31CC48" bold="true" box="[628,721,1885,1909]" pageId="6" pageNumber="387">sp. nov.</emphasis>
</taxonomicNameLabel>
is differentiated from its Sundaland congeners by having the unique combination of a maximum SVL of
<quantity id="4C819B06EA4BCF5DFD05F840FCC1CCA4" box="[691,801,1921,1945]" metricMagnitude="-2" metricUnit="m" metricValue="8.22" pageId="6" pageNumber="387" unit="mm" value="82.2">82.2 mm</quantity>
; nine or 10 supralabials; seven or eight infralabials; moderately tuberculated body and limbs; 32–34 paravertebral tubercles; 37–44 ventral scales; abrupt contact between large and small postfemoral scales; enlarged femoral scales; 20 or 21 subdigital lamellae; no femoral pores; no deep precloacal groove; nine enlarged precloacal pore-bearing scales in males; no enlarged median subcaudals; tubercles on the anterior portion of the dorsal side of the tail; no reticulated pattern on head; having a blotched dorsal pattern; and lacking the paired, dark, semi-lunar shaped blotches on the upper nape prominently outlined in white. These and other characters are scored for all Sundaland species (Girsmer
<emphasis id="B90DEAF1EA4ACF5CFB1FFEC4FB03CA21" box="[1193,1251,260,284]" italics="true" pageId="7" pageNumber="388">et al.</emphasis>
2012a;
<tableCitation id="C6FB0358EA4ACF5CFA89FEC5FA73CA21" box="[1343,1427,260,285]" captionStart="TABLE 4" captionStartId="6.[151,235,584,606]" captionTargetBox="[159,1386,726,1399]" captionTargetPageId="6" captionText="TABLE 4. Uncorrected p-distances for species in the C. sworderi and C. semenanjungensis complex calculated using MEGA 5 (Tamura et al. 2010)." httpUri="http://table.plazi.org/id/DF06666BEA4BCF5DFF21FD89FE8DC9B8" pageId="7" pageNumber="388" tableUuid="DF06666BEA4BCF5DFF21FD89FE8DC9B8">Table 4</tableCitation>
) and for all species in the
<taxonomicName id="4C794D60EA4ACF5CFE19FEE9FD7BCA02" box="[431,667,296,319]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="semenanjungensis">
<emphasis id="B90DEAF1EA4ACF5CFE19FEE9FD7BCA02" box="[431,667,296,319]" italics="true" pageId="7" pageNumber="388">C. semenanjungensis</emphasis>
</taxonomicName>
complex (
<tableCitation id="C6FB0358EA4ACF5CFCA7FEE6FC87CA7D" box="[785,871,295,320]" captionStart="TABLE 5" captionStartId="7.[151,239,362,385]" captionText="TABLE 5. Diagnostic characters (bold) separating species of the Cyrtodactylus semenanjungensis complex from C. metropolis sp. nov." pageId="7" pageNumber="388">Table 5</tableCitation>
).
</paragraph>
<caption id="DF06666BEA4ACF5CFF21FEABFE84CA98" pageId="7" pageNumber="388">
<paragraph id="8BC636E3EA4ACF5CFF21FEABFE84CA98" blockId="7.[151,1436,361,421]" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFF21FEABFEEDCABC" bold="true" box="[151,269,362,385]" pageId="7" pageNumber="388">TABLE 5.</emphasis>
Diagnostic characters (bold) separating species of the
<taxonomicName id="4C794D60EA4ACF5CFCC5FEA8FB35CABC" box="[883,1237,361,385]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="semenanjungensis">
<emphasis id="B90DEAF1EA4ACF5CFCC5FEA8FB35CABC" box="[883,1237,361,385]" italics="true" pageId="7" pageNumber="388">Cyrtodactylus semenanjungensis</emphasis>
</taxonomicName>
complex from
<taxonomicName id="4C794D60EA4ACF5CFA32FEABFEE8CA98" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA4ACF5CFA32FEABFEE8CA98" italics="true" pageId="7" pageNumber="388">C. metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA4ACF5CFEB8FE4FFE84CA98" box="[270,356,398,421]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BC636E3EA4ACF5CFDAFFE03FDCECAFB" box="[537,558,450,454]" pageId="7" pageNumber="388">.</paragraph>
<paragraph id="8BC636E3EA4ACF5CFDAFFE06FDCECAD2" blockId="7.[159,1371,455,807]" box="[537,558,455,495]" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFDAFFE06FDCECAD2" bold="true" box="[537,558,455,495]" pageId="7" pageNumber="388">nov</emphasis>
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFDAFFE34FDCECAC7" blockId="7.[159,1371,455,807]" box="[537,558,501,506]" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFDAFFE34FDCECAC7" bold="true" box="[537,558,501,506]" pageId="7" pageNumber="388">.</emphasis>
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFDAFFE3BFDCEC92E" blockId="7.[159,1371,455,807]" box="[537,558,506,531]" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFDAFFE3BFDCEC92E" bold="true" box="[537,558,506,531]" pageId="7" pageNumber="388">sp</emphasis>
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFDAFFDD8FDCEC9B9" blockId="7.[159,1371,455,807]" box="[537,558,537,644]" pageId="7" pageNumber="388">
<taxonomicName id="4C794D60EA4ACF5CFDAFFDD8FDCEC9B9" box="[537,558,537,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="metropolis">
<emphasis id="B90DEAF1EA4ACF5CFDAFFDD8FDCEC9B9" box="[537,558,537,644]" italics="true" pageId="7" pageNumber="388">metropolis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFD30FE3EFCC0C9B8" blockId="7.[159,1371,455,807]" box="[646,800,511,645]" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFD30FE3EFCC0C9B8" box="[646,800,511,645]" italics="true" pageId="7" pageNumber="388">
<taxonomicName id="4C794D60EA4ACF5CFD30FE3EFD7BC9B8" box="[646,667,511,645]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="seribuatensis">seribuatensis</taxonomicName>
<taxonomicName id="4C794D60EA4ACF5CFCBDFDE3FCC0C9B8" box="[779,800,546,645]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="batucolus">batucolus</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFC26FE0CFAD2C9B8" blockId="7.[159,1371,455,807]" box="[912,1330,461,645]" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFC26FE0CFAD2C9B8" box="[912,1330,461,645]" italics="true" pageId="7" pageNumber="388">
<taxonomicName id="4C794D60EA4ACF5CFC26FE0CFC45C9B8" box="[912,933,461,645]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="semenanjungensis">semenanjungensis</taxonomicName>
<taxonomicName id="4C794D60EA4ACF5CFBA2FDF3FBC9C9B8" box="[1044,1065,562,645]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="majulah">majulah</taxonomicName>
<taxonomicName id="4C794D60EA4ACF5CFB2FFDDFFB4EC9B8" box="[1177,1198,542,645]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="pantiensis">pantiensis</taxonomicName>
<taxonomicName id="4C794D60EA4ACF5CFAABFDE9FAD2C9B8" box="[1309,1330,552,645]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="payacola">payacola</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFF29FD54FABBCCCD" blockId="7.[159,1371,455,807]" lastBlockId="7.[151,1436,1510,2032]" pageId="7" pageNumber="388">
Supralabials 9,10 8–10 9–
<date id="FFC71023EA4ACF5CFC93FD54FCDCC996" box="[805,828,661,683]" pageId="7" pageNumber="388" value="2011-08-11" valueMax="2011-08-15" valueMin="2011-08-11">11</date>
<emphasis id="B90DEAF1EA4ACF5CFC39FD54FC2BC996" bold="true" box="[911,971,661,683]" pageId="7" pageNumber="388">11–15</emphasis>
8,
<date id="FFC71023EA4ACF5CFB93FD54FB30C996" box="[1061,1232,661,683]" pageId="7" pageNumber="388" value="2011-09-10">9 10,11</date>
10 Infralabials 7,
<date id="FFC71023EA4ACF5CFD9DFD7FFD59C9E9" box="[555,697,702,724]" pageId="7" pageNumber="388" value="2010-07-08">8 7–10</date>
9–
<date id="FFC71023EA4ACF5CFC93FD7FFCDCC9E9" box="[805,828,702,724]" pageId="7" pageNumber="388" value="2011-10-07" valueMax="2011-12-07" valueMin="2011-10-07">11</date>
<emphasis id="B90DEAF1EA4ACF5CFC39FD7FFC2EC9E9" bold="true" box="[911,974,702,724]" pageId="7" pageNumber="388">10–12</emphasis>
7,
<date id="FFC71023EA4ACF5CFB90FD7FFB24C9E9" box="[1062,1220,702,724]" pageId="7" pageNumber="388" value="2010-08-09">
8
<emphasis id="B90DEAF1EA4ACF5CFB2EFD7FFB24C9E9" bold="true" box="[1176,1220,702,724]" pageId="7" pageNumber="388">9,10</emphasis>
</date>
8–10 Paravertebral tubercles 32–34 27–35 30–35 32–37
<emphasis id="B90DEAF1EA4ACF5CFBA5FD29FABBC9C3" bold="true" box="[1043,1371,744,766]" pageId="7" pageNumber="388">39–46 35–37 37–39</emphasis>
Ventral scales 37–44 28–39 40–45 48–53 43–51 40–45 37–50
<emphasis id="B90DEAF1EA4ACF5CFF71FA27FE05CEC2" bold="true" box="[199,485,1510,1535]" pageId="7" pageNumber="388">
Description of
<typeStatus id="54C28841EA4ACF5CFECEFA26FE3FCEC2" box="[376,479,1511,1535]" pageId="7" pageNumber="388" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male SVL
<quantity id="4C819B06EA4ACF5CFD19FA26FCF6CEC3" box="[687,790,1511,1535]" metricMagnitude="-2" metricUnit="m" metricValue="8.22" pageId="7" pageNumber="388" unit="mm" value="82.2">82.2 mm</quantity>
; head moderate in length (HL/SVL 0.28) and wide (HW/ HL 0.65), flat (HD/HL 0.41), distinct from neck, and triangular in dorsal profile; lores weakly inflated, prefrontal region deeply concave, canthus rostralis rounded; snout elongate (ES/HL 0.43), rounded in dorsal profile; eye large (ED/HL 0.24); ear opening elliptical, obliquely oriented, moderate in size (EL/HL 0.07); eye to ear distance greater than diameter of eye; rostral rectangular, partially divided dorsally by an inverted Y-shaped furrow, bordered posteriorly by large left and right supranasals and two smaller internasals, laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by four small postnasals and ventrally by first supralabial; 10 (R,L) rectangular supralabials extending to just beyond upturn of labial margins tapering abruptly below midpoint of eye, first supralabial largest; seven (R,L) infralabials tapering smoothly posteriorly to beyond orbit; scales of rostrum and lores slightly raised, slightly larger than granular scales on top of head and occiput; patch of skin missing from top of head; scales of occiput intermixed with slightly enlarged tubercles; dorsal superciliaries not elongate or keeled; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapezoidal postmentals which contact medially for 50% of their length posterior to mental; one row of slightly enlarged, elongate sublabials extending posteriorly to 5th infralabial; gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales.
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFF29FCFBFABCC89E" pageId="7" pageNumber="388">
<table id="F979C443EA4A30A4FF29FCFBFA9CC89E" box="[159,1404,826,931]" gridcols="6" gridrows="3" pageId="7" pageNumber="388">
<tr id="354934A1EA4A30A4FF29FCFBFA9CC86D" box="[159,1404,826,848]" gridrow="0" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FCFBFD25C86D" box="[159,709,826,848]" gridcol="0" gridrow="0" pageId="7" pageNumber="388">
Enlarged median subcaudal scales No
<emphasis id="B90DEAF1EA4ACF5CFD30FCFBFD48C86D" bold="true" box="[646,680,826,848]" pageId="7" pageNumber="388">Ye s</emphasis>
</th>
<th id="76985DDDEA4A30A4FCBCFCFBFCA9C86D" box="[778,841,826,848]" gridcol="1" gridrow="0" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFCBCFCFBFCCCC86D" bold="true" box="[778,812,826,848]" pageId="7" pageNumber="388">Ye s</emphasis>
</th>
<th id="76985DDDEA4A30A4FC39FCFBFC2EC86D" box="[911,974,826,848]" gridcol="2" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FBA5FCFBFBB2C86D" box="[1043,1106,826,848]" gridcol="3" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FB2EFCFBFB31C86D" box="[1176,1233,826,848]" gridcol="4" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FAAAFCFBFA9CC86D" box="[1308,1404,826,848]" gridcol="5" gridrow="0" pageId="7" pageNumber="388">No</th>
</tr>
<tr id="354934A1EA4A30A4FF29FCA5FA9CC847" box="[159,1404,868,890]" gridrow="1" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FCA5FD25C847" box="[159,709,868,890]" gridcol="0" gridrow="1" pageId="7" pageNumber="388">Postfemoral scale contact abrupt Yes Yes</th>
<td id="76985DDDEA4A30A4FCBCFCA5FCA9C847" box="[778,841,868,890]" gridcol="1" gridrow="1" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FC39FCA5FC2EC847" box="[911,974,868,890]" gridcol="2" gridrow="1" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FBA5FCA5FBB2C847" box="[1043,1106,868,890]" gridcol="3" gridrow="1" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFBA5FCA5FBD2C847" bold="true" box="[1043,1074,868,890]" pageId="7" pageNumber="388">No</emphasis>
</td>
<td id="76985DDDEA4A30A4FB2EFCA5FB31C847" box="[1176,1233,868,890]" gridcol="4" gridrow="1" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FAAAFCA5FA9CC847" box="[1308,1404,868,890]" gridcol="5" gridrow="1" pageId="7" pageNumber="388">Yes</td>
</tr>
<tr id="354934A1EA4A30A4FF29FC4CFA9CC89E" box="[159,1404,909,931]" gridrow="2" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FC4CFD25C89E" box="[159,709,909,931]" gridcol="0" gridrow="2" pageId="7" pageNumber="388">4th toe lamellae 20,21 17–22</th>
<td id="76985DDDEA4A30A4FCBCFC4CFCA9C89E" box="[778,841,909,931]" gridcol="1" gridrow="2" pageId="7" pageNumber="388">21–23</td>
<td id="76985DDDEA4A30A4FC39FC4CFC2EC89E" box="[911,974,909,931]" gridcol="2" gridrow="2" pageId="7" pageNumber="388">17–21</td>
<td id="76985DDDEA4A30A4FBA5FC4CFBB2C89E" box="[1043,1106,909,931]" gridcol="3" gridrow="2" pageId="7" pageNumber="388">20–23</td>
<td id="76985DDDEA4A30A4FB2EFC4CFB31C89E" box="[1176,1233,909,931]" gridcol="4" gridrow="2" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFB2EFC4CFB31C89E" bold="true" box="[1176,1233,909,931]" pageId="7" pageNumber="388">22,23</emphasis>
</td>
<td id="76985DDDEA4A30A4FAAAFC4CFA9CC89E" box="[1308,1404,909,931]" gridcol="5" gridrow="2" pageId="7" pageNumber="388">18–21</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC636E3EA4ACF5CFF29FC76FAC3CEB5" pageId="7" pageNumber="388">
<table id="F979C443EA4A30A4FF29FC77FA9CCE9A" box="[159,1404,950,1447]" gridcols="8" gridrows="11" pageId="7" pageNumber="388">
<tr id="354934A1EA4A30A4FF29FC77FA9CC8F1" box="[159,1404,950,972]" gridrow="0" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FC77FE10C8F1" box="[159,496,950,972]" gridcol="0" gridrow="0" pageId="7" pageNumber="388">Deep precloacal groove</th>
<th id="76985DDDEA4A30A4FDAEFC77FD86C8F1" box="[536,614,950,972]" gridcol="1" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FD30FC77FD06C8F1" box="[646,742,950,972]" gridcol="2" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FCBCFC77FC8AC8F1" box="[778,874,950,972]" gridcol="3" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FC39FC77FC0BC8F1" box="[911,1003,950,972]" gridcol="4" gridrow="0" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFC39FC77FC0BC8F1" bold="true" box="[911,1003,950,972]" pageId="7" pageNumber="388">Variable</emphasis>
</th>
<th id="76985DDDEA4A30A4FBA5FC77FB93C8F1" box="[1043,1139,950,972]" gridcol="5" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FB21FC77FB18C8F1" box="[1175,1272,950,972]" gridcol="6" gridrow="0" pageId="7" pageNumber="388">No</th>
<th id="76985DDDEA4A30A4FAAAFC77FA9CC8F1" box="[1308,1404,950,972]" gridcol="7" gridrow="0" pageId="7" pageNumber="388">no</th>
</tr>
<tr id="354934A1EA4A30A4FF29FC21FA9CC8CB" box="[159,1404,992,1014]" gridrow="1" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FC21FE10C8CB" box="[159,496,992,1014]" gridcol="0" gridrow="1" pageId="7" pageNumber="388">No. of precloacal pores</th>
<td id="76985DDDEA4A30A4FDAEFC21FD86C8CB" box="[536,614,992,1014]" gridcol="1" gridrow="1" pageId="7" pageNumber="388">0</td>
<emphasis id="B90DEAF1EA4ACF5CFD30FC21FCD6C8CB" bold="true" box="[646,822,992,1014]" pageId="7" pageNumber="388">
<td id="76985DDDEA4A30A4FD30FC21FD06C8CB" box="[646,742,992,1014]" gridcol="2" gridrow="1" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFD30FC21FD25C8CB" bold="true" box="[646,709,992,1014]" pageId="7" pageNumber="388">10–12</emphasis>
</td>
<td id="76985DDDEA4A30A4FCBCFC21FC8AC8CB" box="[778,874,992,1014]" gridcol="3" gridrow="1" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFCBCFC21FCD6C8CB" bold="true" box="[778,822,992,1014]" pageId="7" pageNumber="388">9,10</emphasis>
</td>
</emphasis>
<td id="76985DDDEA4A30A4FC39FC21FC0BC8CB" box="[911,1003,992,1014]" gridcol="4" gridrow="1" pageId="7" pageNumber="388">0</td>
<td id="76985DDDEA4A30A4FBA5FC21FB93C8CB" box="[1043,1139,992,1014]" gridcol="5" gridrow="1" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFBA5FC21FBA5C8CB" bold="true" box="[1043,1093,992,1014]" pageId="7" pageNumber="388">7–11</emphasis>
</td>
<td id="76985DDDEA4A30A4FB21FC21FB18C8CB" box="[1175,1272,992,1014]" gridcol="6" gridrow="1" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFB2EFC21FB57C8CB" bold="true" box="[1176,1207,992,1014]" pageId="7" pageNumber="388">8,9</emphasis>
</td>
<td id="76985DDDEA4A30A4FAAAFC21FA9CC8CB" box="[1308,1404,992,1014]" gridcol="7" gridrow="1" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFAAAFC21FABCC8CB" bold="true" box="[1308,1372,992,1014]" pageId="7" pageNumber="388">12–15</emphasis>
</td>
</tr>
<tr id="354934A1EA4A30A4FF29FBC8FA9CCF22" box="[159,1404,1033,1055]" gridrow="2" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FBC8FE10CF22" box="[159,496,1033,1055]" gridcol="0" gridrow="2" pageId="7" pageNumber="388">Enlarged precloacal scales</th>
<td id="76985DDDEA4A30A4FDAEFBC8FD86CF22" box="[536,614,1033,1055]" gridcol="1" gridrow="2" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FD30FBC8FD06CF22" box="[646,742,1033,1055]" gridcol="2" gridrow="2" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FCBCFBC8FC8ACF22" box="[778,874,1033,1055]" gridcol="3" gridrow="2" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FC39FBC8FC0BCF22" box="[911,1003,1033,1055]" gridcol="4" gridrow="2" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FBA5FBC8FB93CF22" box="[1043,1139,1033,1055]" gridcol="5" gridrow="2" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FB21FBC8FB18CF22" box="[1175,1272,1033,1055]" gridcol="6" gridrow="2" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FAAAFBC8FA9CCF22" box="[1308,1404,1033,1055]" gridcol="7" gridrow="2" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFAAAFBC8FADBCF22" bold="true" box="[1308,1339,1033,1055]" pageId="7" pageNumber="388">No</emphasis>
</td>
</tr>
<tr id="354934A1EA4A30A4FF29FBF3FA9CCF75" box="[159,1404,1074,1096]" gridrow="3" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FBF3FE10CF75" box="[159,496,1074,1096]" gridcol="0" gridrow="3" pageId="7" pageNumber="388">No. of femoral pores</th>
<td id="76985DDDEA4A30A4FDAEFBF3FD86CF75" box="[536,614,1074,1096]" gridcol="1" gridrow="3" pageId="7" pageNumber="388">0</td>
<emphasis id="B90DEAF1EA4ACF5CFD30FBF3FCD0CF75" bold="true" box="[646,816,1074,1096]" pageId="7" pageNumber="388">
<td id="76985DDDEA4A30A4FD30FBF3FD06CF75" box="[646,742,1074,1096]" gridcol="2" gridrow="3" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFD30FBF3FD25CF75" bold="true" box="[646,709,1074,1096]" pageId="7" pageNumber="388">14–17</emphasis>
</td>
<td id="76985DDDEA4A30A4FCBCFBF3FC8ACF75" box="[778,874,1074,1096]" gridcol="3" gridrow="3" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFCBCFBF3FCD0CF75" bold="true" box="[778,816,1074,1096]" pageId="7" pageNumber="388">6–9</emphasis>
</td>
</emphasis>
<td id="76985DDDEA4A30A4FC39FBF3FC0BCF75" box="[911,1003,1074,1096]" gridcol="4" gridrow="3" pageId="7" pageNumber="388">0</td>
<td id="76985DDDEA4A30A4FBA5FBF3FB93CF75" box="[1043,1139,1074,1096]" gridcol="5" gridrow="3" pageId="7" pageNumber="388">0</td>
<td id="76985DDDEA4A30A4FB21FBF3FB18CF75" box="[1175,1272,1074,1096]" gridcol="6" gridrow="3" pageId="7" pageNumber="388">0</td>
<td id="76985DDDEA4A30A4FAAAFBF3FA9CCF75" box="[1308,1404,1074,1096]" gridcol="7" gridrow="3" pageId="7" pageNumber="388">0</td>
</tr>
<tr id="354934A1EA4A30A4FF29FB9DFA9CCF4F" box="[159,1404,1116,1138]" gridrow="4" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FB9DFE10CF4F" box="[159,496,1116,1138]" gridcol="0" gridrow="4" pageId="7" pageNumber="388">Enlarged femoral scales</th>
<td id="76985DDDEA4A30A4FDAEFB9DFD86CF4F" box="[536,614,1116,1138]" gridcol="1" gridrow="4" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FD30FB9DFD06CF4F" box="[646,742,1116,1138]" gridcol="2" gridrow="4" pageId="7" pageNumber="388">Yes</td>
<td id="76985DDDEA4A30A4FCBCFB9DFC8ACF4F" box="[778,874,1116,1138]" gridcol="3" gridrow="4" pageId="7" pageNumber="388">Yes</td>
<emphasis id="B90DEAF1EA4ACF5CFC39FB9DFB57CF4F" bold="true" box="[911,1207,1116,1138]" pageId="7" pageNumber="388">
<td id="76985DDDEA4A30A4FC39FB9DFC0BCF4F" box="[911,1003,1116,1138]" gridcol="4" gridrow="4" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFC39FB9DFC4ECF4F" bold="true" box="[911,942,1116,1138]" pageId="7" pageNumber="388">No</emphasis>
</td>
<td id="76985DDDEA4A30A4FBA5FB9DFB93CF4F" box="[1043,1139,1116,1138]" gridcol="5" gridrow="4" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFBA5FB9DFBD2CF4F" bold="true" box="[1043,1074,1116,1138]" pageId="7" pageNumber="388">No</emphasis>
</td>
<td id="76985DDDEA4A30A4FB21FB9DFB18CF4F" box="[1175,1272,1116,1138]" gridcol="6" gridrow="4" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFB2EFB9DFB57CF4F" bold="true" box="[1176,1207,1116,1138]" pageId="7" pageNumber="388">No</emphasis>
</td>
</emphasis>
<td id="76985DDDEA4A30A4FAAAFB9DFA9CCF4F" box="[1308,1404,1116,1138]" gridcol="7" gridrow="4" pageId="7" pageNumber="388">Yes</td>
</tr>
<tr id="354934A1EA4A30A4FF29FB44FA9CCF87" box="[159,1404,1157,1210]" gridrow="5" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FB44FE10CF87" box="[159,496,1157,1210]" gridcol="0" gridrow="5" pageId="7" pageNumber="388">Enlarged femorocloacal scales continuous</th>
<td id="76985DDDEA4A30A4FDAEFB44FD86CF87" box="[536,614,1157,1210]" gridcol="1" gridrow="5" pageId="7" pageNumber="388">yes</td>
<td id="76985DDDEA4A30A4FD30FB44FD06CF87" box="[646,742,1157,1210]" gridcol="2" gridrow="5" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFD30FB44FD45CFA6" bold="true" box="[646,677,1157,1179]" pageId="7" pageNumber="388">No</emphasis>
</td>
<td id="76985DDDEA4A30A4FCBCFB44FC8ACF87" box="[778,874,1157,1210]" gridcol="3" gridrow="5" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFCBCFB44FCC9CFA6" bold="true" box="[778,809,1157,1179]" pageId="7" pageNumber="388">No</emphasis>
</td>
<td id="76985DDDEA4A30A4FC39FB44FC0BCF87" box="[911,1003,1157,1210]" gridcol="4" gridrow="5" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FBA5FB44FB93CF87" box="[1043,1139,1157,1210]" gridcol="5" gridrow="5" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FB21FB44FB18CF87" box="[1175,1272,1157,1210]" gridcol="6" gridrow="5" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FAAAFB44FA9CCF87" box="[1308,1404,1157,1210]" gridcol="7" gridrow="5" pageId="7" pageNumber="388">Yes</td>
</tr>
<tr id="354934A1EA4A30A4FF29FB0CFA9CCFDE" box="[159,1404,1229,1251]" gridrow="6" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FB0CFE10CFDE" box="[159,496,1229,1251]" gridcol="0" gridrow="6" pageId="7" pageNumber="388">Proximal light caudal bands</th>
<td id="76985DDDEA4A30A4FDAEFB0CFD86CFDE" box="[536,614,1229,1251]" gridcol="1" gridrow="6" pageId="7" pageNumber="388">Narrow</td>
<td id="76985DDDEA4A30A4FD30FB0CFD06CFDE" box="[646,742,1229,1251]" gridcol="2" gridrow="6" pageId="7" pageNumber="388">Narrow</td>
<td id="76985DDDEA4A30A4FCBCFB0CFC8ACFDE" box="[778,874,1229,1251]" gridcol="3" gridrow="6" pageId="7" pageNumber="388">Narrow</td>
<td id="76985DDDEA4A30A4FC39FB0CFC0BCFDE" box="[911,1003,1229,1251]" gridcol="4" gridrow="6" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFC39FB0CFC28CFDE" bold="true" box="[911,968,1229,1251]" pageId="7" pageNumber="388">Wide</emphasis>
</td>
<td id="76985DDDEA4A30A4FBA5FB0CFB93CFDE" box="[1043,1139,1229,1251]" gridcol="5" gridrow="6" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFBA5FB0CFBACCFDE" bold="true" box="[1043,1100,1229,1251]" pageId="7" pageNumber="388">Wide</emphasis>
</td>
<td id="76985DDDEA4A30A4FB21FB0CFB18CFDE" box="[1175,1272,1229,1251]" gridcol="6" gridrow="6" pageId="7" pageNumber="388">Narrow</td>
<td id="76985DDDEA4A30A4FAAAFB0CFA9CCFDE" box="[1308,1404,1229,1251]" gridcol="7" gridrow="6" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFAAAFB0CFAB5CFDE" bold="true" box="[1308,1365,1229,1251]" pageId="7" pageNumber="388">Wide</emphasis>
</td>
</tr>
<tr id="354934A1EA4A30A4FF29FB37FA9CCE31" box="[159,1404,1270,1292]" gridrow="7" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FB37FE10CE31" box="[159,496,1270,1292]" gridcol="0" gridrow="7" pageId="7" pageNumber="388">No. of light bands on original tail</th>
<td id="76985DDDEA4A30A4FDAEFB37FD86CE31" box="[536,614,1270,1292]" gridcol="1" gridrow="7" pageId="7" pageNumber="388">9</td>
<td id="76985DDDEA4A30A4FD30FB37FD06CE31" box="[646,742,1270,1292]" gridcol="2" gridrow="7" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFD30FB37FD25CE31" bold="true" box="[646,709,1270,1292]" pageId="7" pageNumber="388">13–15</emphasis>
</td>
<td id="76985DDDEA4A30A4FCBCFB37FC8ACE31" box="[778,874,1270,1292]" gridcol="3" gridrow="7" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFCBCFB37FCAACE31" bold="true" box="[778,842,1270,1292]" pageId="7" pageNumber="388">14–16</emphasis>
</td>
<td id="76985DDDEA4A30A4FC39FB37FC0BCE31" box="[911,1003,1270,1292]" gridcol="4" gridrow="7" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFC39FB37FC47CE31" bold="true" box="[911,935,1270,1292]" pageId="7" pageNumber="388">11</emphasis>
</td>
<td id="76985DDDEA4A30A4FBA5FB37FB93CE31" box="[1043,1139,1270,1292]" gridcol="5" gridrow="7" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFBA5FB37FBCDCE31" bold="true" box="[1043,1069,1270,1292]" pageId="7" pageNumber="388">16</emphasis>
</td>
<td id="76985DDDEA4A30A4FB21FB37FB18CE31" box="[1175,1272,1270,1292]" gridcol="6" gridrow="7" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFB2EFB37FB37CE31" bold="true" box="[1176,1239,1270,1292]" pageId="7" pageNumber="388">13–15</emphasis>
</td>
<td id="76985DDDEA4A30A4FAAAFB37FA9CCE31" box="[1308,1404,1270,1292]" gridcol="7" gridrow="7" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFAAAFB37FAB5CE31" bold="true" box="[1308,1365,1270,1292]" pageId="7" pageNumber="388">13,14</emphasis>
</td>
</tr>
<tr id="354934A1EA4A30A4FF29FAE1FA9CCE0B" box="[159,1404,1312,1334]" gridrow="8" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FAE1FE10CE0B" box="[159,496,1312,1334]" gridcol="0" gridrow="8" pageId="7" pageNumber="388">Dorsal pattern</th>
<td id="76985DDDEA4A30A4FDAEFAE1FD86CE0B" box="[536,614,1312,1334]" gridcol="1" gridrow="8" pageId="7" pageNumber="388">Banded</td>
<td id="76985DDDEA4A30A4FD30FAE1FD06CE0B" box="[646,742,1312,1334]" gridcol="2" gridrow="8" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFD30FAE1FD06CE0B" bold="true" box="[646,742,1312,1334]" pageId="7" pageNumber="388">Blotched</emphasis>
</td>
<td id="76985DDDEA4A30A4FCBCFAE1FC8ACE0B" box="[778,874,1312,1334]" gridcol="3" gridrow="8" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFCBCFAE1FC8ACE0B" bold="true" box="[778,874,1312,1334]" pageId="7" pageNumber="388">Blotched</emphasis>
</td>
<td id="76985DDDEA4A30A4FC39FAE1FC0BCE0B" box="[911,1003,1312,1334]" gridcol="4" gridrow="8" pageId="7" pageNumber="388">Banded</td>
<td id="76985DDDEA4A30A4FBA5FAE1FB93CE0B" box="[1043,1139,1312,1334]" gridcol="5" gridrow="8" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFBA5FAE1FB93CE0B" bold="true" box="[1043,1139,1312,1334]" pageId="7" pageNumber="388">Blotched</emphasis>
</td>
<td id="76985DDDEA4A30A4FB21FAE1FB18CE0B" box="[1175,1272,1312,1334]" gridcol="6" gridrow="8" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFB2EFAE1FB18CE0B" bold="true" box="[1176,1272,1312,1334]" pageId="7" pageNumber="388">Blotched</emphasis>
</td>
<td id="76985DDDEA4A30A4FAAAFAE1FA9CCE0B" box="[1308,1404,1312,1334]" gridcol="7" gridrow="8" pageId="7" pageNumber="388">
<emphasis id="B90DEAF1EA4ACF5CFAAAFAE1FA9CCE0B" bold="true" box="[1308,1404,1312,1334]" pageId="7" pageNumber="388">Blotched</emphasis>
</td>
</tr>
<tr id="354934A1EA4A30A4FF29FA88FA9CCE62" box="[159,1404,1353,1375]" gridrow="9" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FA88FE10CE62" box="[159,496,1353,1375]" gridcol="0" gridrow="9" pageId="7" pageNumber="388">No. of dorsal body bands</th>
<td id="76985DDDEA4A30A4FDAEFA88FD86CE62" box="[536,614,1353,1375]" gridcol="1" gridrow="9" pageId="7" pageNumber="388">4,5</td>
<td id="76985DDDEA4A30A4FD30FA88FD06CE62" box="[646,742,1353,1375]" gridcol="2" gridrow="9" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FCBCFA88FC8ACE62" box="[778,874,1353,1375]" gridcol="3" gridrow="9" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FC39FA88FC0BCE62" box="[911,1003,1353,1375]" gridcol="4" gridrow="9" pageId="7" pageNumber="388">4–6</td>
<td id="76985DDDEA4A30A4FBA5FA88FB93CE62" box="[1043,1139,1353,1375]" gridcol="5" gridrow="9" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FB21FA88FB18CE62" box="[1175,1272,1353,1375]" gridcol="6" gridrow="9" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FAAAFA88FA9CCE62" box="[1308,1404,1353,1375]" gridcol="7" gridrow="9" pageId="7" pageNumber="388">/</td>
</tr>
<tr id="354934A1EA4A30A4FF29FAB3FA9CCE9A" box="[159,1404,1394,1447]" gridrow="10" pageId="7" pageNumber="388">
<th id="76985DDDEA4A30A4FF29FAB3FE10CE9A" box="[159,496,1394,1447]" gridcol="0" gridrow="10" pageId="7" pageNumber="388">Hatching/juvenles with white tail tips</th>
<td id="76985DDDEA4A30A4FDAEFAB3FD86CE9A" box="[536,614,1394,1447]" gridcol="1" gridrow="10" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FD30FAB3FD06CE9A" box="[646,742,1394,1447]" gridcol="2" gridrow="10" pageId="7" pageNumber="388">No</td>
<td id="76985DDDEA4A30A4FCBCFAB3FC8ACE9A" box="[778,874,1394,1447]" gridcol="3" gridrow="10" pageId="7" pageNumber="388">No</td>
<td id="76985DDDEA4A30A4FC39FAB3FC0BCE9A" box="[911,1003,1394,1447]" gridcol="4" gridrow="10" pageId="7" pageNumber="388">No</td>
<td id="76985DDDEA4A30A4FBA5FAB3FB93CE9A" box="[1043,1139,1394,1447]" gridcol="5" gridrow="10" pageId="7" pageNumber="388">Variable</td>
<td id="76985DDDEA4A30A4FB21FAB3FB18CE9A" box="[1175,1272,1394,1447]" gridcol="6" gridrow="10" pageId="7" pageNumber="388">/</td>
<td id="76985DDDEA4A30A4FAAAFAB3FA9CCE9A" box="[1308,1404,1394,1447]" gridcol="7" gridrow="10" pageId="7" pageNumber="388">/</td>
</tr>
</table>
</paragraph>
<caption id="DF06666BEA45CF53FF21FAD4FE30CE77" httpUri="https://zenodo.org/record/230943/files/figure.png" pageId="8" pageNumber="389" targetBox="[151,1436,176,1279]" targetPageId="8">
<paragraph id="8BC636E3EA45CF53FF21FAD4FE30CE77" blockId="8.[151,1435,1301,1354]" pageId="8" pageNumber="389">
<emphasis id="B90DEAF1EA45CF53FF21FAD4FEF4CE16" bold="true" box="[151,276,1301,1323]" pageId="8" pageNumber="389">FIGURE 3.</emphasis>
<taxonomicName id="4C794D60EA45CF53FEAAFAD4FDFDCE16" box="[284,541,1301,1323]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA45CF53FEAAFAD4FDFDCE16" box="[284,541,1301,1323]" italics="true" pageId="8" pageNumber="389">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA45CF53FD93FAD4FD98CE16" box="[549,632,1301,1323]" pageId="8" pageNumber="389" rank="species">
<emphasis id="B90DEAF1EA45CF53FD93FAD4FD98CE16" bold="true" box="[549,632,1301,1323]" pageId="8" pageNumber="389">sp. nov.</emphasis>
</taxonomicNameLabel>
Upper; holotype LSUHC 11347. Lower left; holotype LSUHC 11343. Lower right; paratype LSUHC 11342.
</paragraph>
</caption>
<paragraph id="8BC636E3EA45CF53FF71FAB0FB05CD03" blockId="8.[151,1436,1393,2030]" pageId="8" pageNumber="389">Body relatively short (AG/SVL 0.44) with poorly defined ventrolateral folds; dorsal scales small, granular interspersed with moderately sized, conical, semi-regularly arranged, weakly keeled tubercles; tubercles extend from occiput onto base of tail but no farther; tubercles on occiput and nape very small, those on body largest; approximately 22 longitudinal rows of tubercles at midbody between ventrolateral, body folds; 34 paravertebral tubercles on body; 44 flat, imbricate, ventral scales between ventrolateral, body folds, ventral scales larger than dorsal scales; nine large, pore-bearing, precloacal scales; no deep precloacal groove or depression.</paragraph>
<paragraph id="8BC636E3EA45CF53FF71F988FD61CCD3" blockId="8.[151,1436,1393,2030]" pageId="8" pageNumber="389">Forelimbs moderate in stature, relatively short (FL/SVL 0.16); granular scales of forearm larger than those on body, not interspersed with tubercles; palmar scales rounded, slightly raised; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae transversely expanded proximal to joint inflections, more granular distal to inflection; digits slightly more narrow distal to inflections; claws well-developed, claw base sheathed by a dorsal and ventral scale; hind limbs more robust than forelimbs, moderate in length (TBL/SVL 0.18), covered dorsally by granular scales interspersed with large, conical tubercles and anteriorly by flat, slightly larger scales; ventral scales of thigh flat, imbricate, larger than dorsals; ventral tibial scales flat, imbricate; no rows of enlarged, flat, imbricate, femoral scales; small postfemoral scales do not form an abrupt union with large ventral scales of posteroventral margin of thigh; plantar scales low, flat; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae transversely expanded proximal to inflections, more granular distal to inflections, digits more narrow distal to inflections; 21 subdigital lamellae on right 4th toe, 20 on left; claws well-developed, base of claw sheathed by a dorsal and ventral scale.</paragraph>
<paragraph id="8BC636E3EA44CF52FF71FF56FCA4CA20" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="390">
Tail moderate in proportions,
<quantity id="4C819B06EA44CF52FDA1FF56FD9BCB92" box="[535,635,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="9.15" pageId="9" pageNumber="390" unit="mm" value="91.5">91.5 mm</quantity>
in length, original, 7.0 mm in width at base, tapering to a point; dorsal scales of base of tail granular becoming flatter posteriorly; no median row of transversely enlarged subcaudal scales; subcaudal scales larger than dorsal caudal scales; no cauda tubercles; two enlarged, postcloacal tubercles at base of tail on hemipenal swelling; all postcloacal scales flat.
</paragraph>
<paragraph id="8BC636E3EA44CF52FF71FEE6FEC4CAC8" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="390">
<emphasis id="B90DEAF1EA44CF52FF71FEE6FE18CA7D" bold="true" box="[199,504,295,320]" pageId="9" pageNumber="390">
Coloration in life (
<figureCitation id="13422A66EA44CF52FE15FEE6FE09CA7D" box="[419,489,295,320]" captionStart="FIGURE 3" captionStartId="8.[151,250,1301,1323]" captionTargetBox="[151,1436,176,1279]" captionTargetId="figure@8.[151,1436,169,1279]" captionTargetPageId="8" captionText="FIGURE 3. Cyrtodactylus metropolis sp. nov. Upper; holotype LSUHC 11347. Lower left; holotype LSUHC 11343. Lower right; paratype LSUHC 11342." httpUri="https://zenodo.org/record/230943/files/figure.png" pageId="9" pageNumber="390">Fig. 3</figureCitation>
).
</emphasis>
Dorsal ground color of head, neck, trunk, limbs, and tail brown; beige mottling on head and rostrum; beige postorbital stripe extending to above ear opening; one short, narrow, beige band on nape; four beige, narrow bands between limb insertions continuing onto tail and encircling it; second and third bands bifurcate on the right and left side, respectively; irregularly shaped bands on limbs; ventral surfaces of head, neck, body and limbs beige with fine black stippling on each scale; subcaudal region gray, encircled by white bands; iris dark bronze.
</paragraph>
</subSubSection>
<caption id="DF06666BEA44CF52FF21FDE5FBEBC963" ID-Table-UUID="DF06666BEA44CF52FF21FDE5FBEBC963" httpUri="http://table.plazi.org/id/DF06666BEA44CF52FF21FDE5FBEBC963" pageId="9" pageNumber="390" targetBox="[159,1416,636,2011]" targetIsTable="true" targetPageId="9">
<paragraph id="8BC636E3EA44CF52FF21FDE5FBEBC963" blockId="9.[151,1436,547,606]" pageId="9" pageNumber="390">
<emphasis id="B90DEAF1EA44CF52FF21FDE5FEEDC906" bold="true" box="[151,269,548,571]" pageId="9" pageNumber="390">TABLE 6.</emphasis>
Morphological characters and selected maesurements (in mm) of the type series of
<taxonomicName id="4C794D60EA44CF52FB3BFDE2FA7CC906" box="[1165,1436,547,571]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA44CF52FB3BFDE2FA7CC906" box="[1165,1436,547,571]" italics="true" pageId="9" pageNumber="390">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA44CF52FF21FD86FF0EC963" box="[151,238,583,606]" pageId="9" pageNumber="390" rank="species">
<emphasis id="B90DEAF1EA44CF52FF21FD86FF0EC963" bold="true" box="[151,238,583,606]" pageId="9" pageNumber="390">sp. nov.</emphasis>
</taxonomicNameLabel>
/=data unavailable. Abbreviations are listed in the Materials and Methods.
</paragraph>
</caption>
<paragraph id="8BC636E3EA44CF52FF29FD0FFAEFCCE6" pageId="9" pageNumber="390">
<table id="F979C443EA4430A4FF29FDBDFA68CCE6" box="[159,1416,636,2011]" gridcols="5" gridrows="11" pageId="9" pageNumber="390">
<tr id="354934A1EA4430A4FF29FDBDFA68C9D9" box="[159,1416,636,740]" gridrow="0" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29FDBDFD5AC9D9" box="[159,698,636,740]" gridcol="0" gridrow="0" pageId="9" pageNumber="390">Sex</th>
<th id="76985DDDEA4430A4FD67FDBDFC88C9D9" box="[721,872,636,740]" gridcol="1" gridrow="0" pageId="9" pageNumber="390">LSUHC 11347 holotype F</th>
<th id="76985DDDEA4430A4FC30FDBDFBFDC9D9" box="[902,1053,636,740]" gridcol="2" gridrow="0" pageId="9" pageNumber="390">LSUHC 11342 paratype M</th>
<th id="76985DDDEA4430A4FB8DFDBDFB32C9D9" box="[1083,1234,636,740]" gridcol="3" gridrow="0" pageId="9" pageNumber="390">LSUHC 11343 paratype M</th>
<th id="76985DDDEA4430A4FB46FDBDFA68C9D9" box="[1264,1416,636,740]" gridcol="4" gridrow="0" pageId="9" pageNumber="390">LSUHC 11345 paratype M</th>
</tr>
<tr id="354934A1EA4430A4FF29FD39FA68C85D" box="[159,1416,760,864]" gridrow="1" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29FD39FD5AC85D" box="[159,698,760,864]" gridcol="0" gridrow="1" pageId="9" pageNumber="390">Supralabials Infralabials Paravertebral tubercles</th>
<td id="76985DDDEA4430A4FD67FD39FC88C85D" box="[721,872,760,864]" gridcol="1" gridrow="1" pageId="9" pageNumber="390">10 7 34</td>
<td id="76985DDDEA4430A4FC30FD39FBFDC85D" box="[902,1053,760,864]" gridcol="2" gridrow="1" pageId="9" pageNumber="390">9 8 32</td>
<td id="76985DDDEA4430A4FB8DFD39FB32C85D" box="[1083,1234,760,864]" gridcol="3" gridrow="1" pageId="9" pageNumber="390">10 8 33</td>
<td id="76985DDDEA4430A4FB46FD39FA68C85D" box="[1264,1416,760,864]" gridcol="4" gridrow="1" pageId="9" pageNumber="390">9 8 32</td>
</tr>
<tr id="354934A1EA4430A4FF29FCB5FA68C8E1" box="[159,1416,884,988]" gridrow="2" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29FCB5FD5AC8E1" box="[159,698,884,988]" gridcol="0" gridrow="2" pageId="9" pageNumber="390">Ventral scales Postfemoral scale contact abrupt 4th toe lamellae</th>
<td id="76985DDDEA4430A4FD67FCB5FC88C8E1" box="[721,872,884,988]" gridcol="1" gridrow="2" pageId="9" pageNumber="390">44 YES 21</td>
<td id="76985DDDEA4430A4FC30FCB5FBFDC8E1" box="[902,1053,884,988]" gridcol="2" gridrow="2" pageId="9" pageNumber="390">37 YES 20</td>
<td id="76985DDDEA4430A4FB8DFCB5FB32C8E1" box="[1083,1234,884,988]" gridcol="3" gridrow="2" pageId="9" pageNumber="390">42 YES 20</td>
<td id="76985DDDEA4430A4FB46FCB5FA68C8E1" box="[1264,1416,884,988]" gridcol="4" gridrow="2" pageId="9" pageNumber="390">42 YES 21</td>
</tr>
<tr id="354934A1EA4430A4FF29FC31FA68CF65" box="[159,1416,1008,1112]" gridrow="3" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29FC31FD5ACF65" box="[159,698,1008,1112]" gridcol="0" gridrow="3" pageId="9" pageNumber="390">Precloacal pores present (1) or absent (0) Enlarged precloacal scales Femorocloacal scales</th>
<td id="76985DDDEA4430A4FD67FC31FC88CF65" box="[721,872,1008,1112]" gridcol="1" gridrow="3" pageId="9" pageNumber="390">0 YES 37</td>
<td id="76985DDDEA4430A4FC30FC31FBFDCF65" box="[902,1053,1008,1112]" gridcol="2" gridrow="3" pageId="9" pageNumber="390">0 YES 39</td>
<td id="76985DDDEA4430A4FB8DFC31FB32CF65" box="[1083,1234,1008,1112]" gridcol="3" gridrow="3" pageId="9" pageNumber="390">0 YES 36</td>
<td id="76985DDDEA4430A4FB46FC31FA68CF65" box="[1264,1416,1008,1112]" gridcol="4" gridrow="3" pageId="9" pageNumber="390">0 YES 41</td>
</tr>
<tr id="354934A1EA4430A4FF29FBADFA68CE21" box="[159,1416,1132,1308]" gridrow="4" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29FBADFD5ACE21" box="[159,698,1132,1308]" gridcol="0" gridrow="4" pageId="9" pageNumber="390">Light caudal bands narrow (1) or wide (0) Light bands on original tail Dorsal pattern stiped (S), composed of longitudinally arranged spots (LS), or banded (B) Body bands</th>
<td id="76985DDDEA4430A4FD67FBADFC88CE21" box="[721,872,1132,1308]" gridcol="1" gridrow="4" pageId="9" pageNumber="390">0 / B 5</td>
<td id="76985DDDEA4430A4FC30FBADFBFDCE21" box="[902,1053,1132,1308]" gridcol="2" gridrow="4" pageId="9" pageNumber="390">0 / B 4</td>
<td id="76985DDDEA4430A4FB8DFBADFB32CE21" box="[1083,1234,1132,1308]" gridcol="3" gridrow="4" pageId="9" pageNumber="390">0 / B 5</td>
<td id="76985DDDEA4430A4FB46FBADFA68CE21" box="[1264,1416,1132,1308]" gridcol="4" gridrow="4" pageId="9" pageNumber="390">0 / B 5</td>
</tr>
<tr id="354934A1EA4430A4FF29FAF1FA68CEA5" box="[159,1416,1328,1432]" gridrow="5" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29FAF1FD5ACEA5" box="[159,698,1328,1432]" gridcol="0" gridrow="5" pageId="9" pageNumber="390">Hatch./juv. with white tail tip SVL TL</th>
<td id="76985DDDEA4430A4FD67FAF1FC88CEA5" box="[721,872,1328,1432]" gridcol="1" gridrow="5" pageId="9" pageNumber="390">/ 82.2 91.5</td>
<td id="76985DDDEA4430A4FC30FAF1FBFDCEA5" box="[902,1053,1328,1432]" gridcol="2" gridrow="5" pageId="9" pageNumber="390">/ 60.6 59.5</td>
<td id="76985DDDEA4430A4FB8DFAF1FB32CEA5" box="[1083,1234,1328,1432]" gridcol="3" gridrow="5" pageId="9" pageNumber="390">/ 79.8 83.4</td>
<td id="76985DDDEA4430A4FB46FAF1FA68CEA5" box="[1264,1416,1328,1432]" gridcol="4" gridrow="5" pageId="9" pageNumber="390">/ 78.2 97.9</td>
</tr>
<tr id="354934A1EA4430A4FF29FA6DFA68CD29" box="[159,1416,1452,1556]" gridrow="6" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29FA6DFD5ACD29" box="[159,698,1452,1556]" gridcol="0" gridrow="6" pageId="9" pageNumber="390">TW FL TBL</th>
<td id="76985DDDEA4430A4FD67FA6DFC88CD29" box="[721,872,1452,1556]" gridcol="1" gridrow="6" pageId="9" pageNumber="390">7 12.9 14.7</td>
<td id="76985DDDEA4430A4FC30FA6DFBFDCD29" box="[902,1053,1452,1556]" gridcol="2" gridrow="6" pageId="9" pageNumber="390">5.6 10.1 11.2</td>
<td id="76985DDDEA4430A4FB8DFA6DFB32CD29" box="[1083,1234,1452,1556]" gridcol="3" gridrow="6" pageId="9" pageNumber="390">7.3 13.0 15.5</td>
<td id="76985DDDEA4430A4FB46FA6DFA68CD29" box="[1264,1416,1452,1556]" gridcol="4" gridrow="6" pageId="9" pageNumber="390">7.1 13.2 15.4</td>
</tr>
<tr id="354934A1EA4430A4FF29F9E9FA68CDAD" box="[159,1416,1576,1680]" gridrow="7" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29F9E9FD5ACDAD" box="[159,698,1576,1680]" gridcol="0" gridrow="7" pageId="9" pageNumber="390">AG HL HW</th>
<td id="76985DDDEA4430A4FD67F9E9FC88CDAD" box="[721,872,1576,1680]" gridcol="1" gridrow="7" pageId="9" pageNumber="390">36.2 23.0 15.1</td>
<td id="76985DDDEA4430A4FC30F9E9FBFDCDAD" box="[902,1053,1576,1680]" gridcol="2" gridrow="7" pageId="9" pageNumber="390">25.3 16.9 11.2</td>
<td id="76985DDDEA4430A4FB8DF9E9FB32CDAD" box="[1083,1234,1576,1680]" gridcol="3" gridrow="7" pageId="9" pageNumber="390">34.1 22.8 15.1</td>
<td id="76985DDDEA4430A4FB46F9E9FA68CDAD" box="[1264,1416,1576,1680]" gridcol="4" gridrow="7" pageId="9" pageNumber="390">35.0 22.0 15.0</td>
</tr>
<tr id="354934A1EA4430A4FF29F965FA68CC31" box="[159,1416,1700,1804]" gridrow="8" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29F965FD5ACC31" box="[159,698,1700,1804]" gridcol="0" gridrow="8" pageId="9" pageNumber="390">HD ED EE</th>
<td id="76985DDDEA4430A4FD67F965FC88CC31" box="[721,872,1700,1804]" gridcol="1" gridrow="8" pageId="9" pageNumber="390">9.5 5.5 6.5</td>
<td id="76985DDDEA4430A4FC30F965FBFDCC31" box="[902,1053,1700,1804]" gridcol="2" gridrow="8" pageId="9" pageNumber="390">6.8 3.8 4.8</td>
<td id="76985DDDEA4430A4FB8DF965FB32CC31" box="[1083,1234,1700,1804]" gridcol="3" gridrow="8" pageId="9" pageNumber="390">9.6 5.5 6.2</td>
<td id="76985DDDEA4430A4FB46F965FA68CC31" box="[1264,1416,1700,1804]" gridcol="4" gridrow="8" pageId="9" pageNumber="390">9.5 5.2 6.3</td>
</tr>
<tr id="354934A1EA4430A4FF29F8E1FA68CCB5" box="[159,1416,1824,1928]" gridrow="9" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29F8E1FD5ACCB5" box="[159,698,1824,1928]" gridcol="0" gridrow="9" pageId="9" pageNumber="390">ES EN IO</th>
<td id="76985DDDEA4430A4FD67F8E1FC88CCB5" box="[721,872,1824,1928]" gridcol="1" gridrow="9" pageId="9" pageNumber="390">9.9 7.8 4.2</td>
<td id="76985DDDEA4430A4FC30F8E1FBFDCCB5" box="[902,1053,1824,1928]" gridcol="2" gridrow="9" pageId="9" pageNumber="390">7.4 5.7 3.0</td>
<td id="76985DDDEA4430A4FB8DF8E1FB32CCB5" box="[1083,1234,1824,1928]" gridcol="3" gridrow="9" pageId="9" pageNumber="390">10.0 7.6 4.2</td>
<td id="76985DDDEA4430A4FB46F8E1FA68CCB5" box="[1264,1416,1824,1928]" gridcol="4" gridrow="9" pageId="9" pageNumber="390">10.1 7.6 3.8</td>
</tr>
<tr id="354934A1EA4430A4FF29F85DFA68CCE6" box="[159,1416,1948,2011]" gridrow="10" pageId="9" pageNumber="390">
<th id="76985DDDEA4430A4FF29F85DFD5ACCE6" box="[159,698,1948,2011]" gridcol="0" gridrow="10" pageId="9" pageNumber="390">EL IN</th>
<td id="76985DDDEA4430A4FD67F85DFC88CCE6" box="[721,872,1948,2011]" gridcol="1" gridrow="10" pageId="9" pageNumber="390">1.7 2.5</td>
<td id="76985DDDEA4430A4FC30F85DFBFDCCE6" box="[902,1053,1948,2011]" gridcol="2" gridrow="10" pageId="9" pageNumber="390">1.4 2.0</td>
<td id="76985DDDEA4430A4FB8DF85DFB32CCE6" box="[1083,1234,1948,2011]" gridcol="3" gridrow="10" pageId="9" pageNumber="390">1.7 2.5</td>
<td id="76985DDDEA4430A4FB46F85DFA68CCE6" box="[1264,1416,1948,2011]" gridcol="4" gridrow="10" pageId="9" pageNumber="390">2.3 2.4</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3636568EA47CF51FF71FF56FEBDCA20" pageId="10" pageNumber="391" type="description">
<paragraph id="8BC636E3EA47CF51FF71FF56FEBDCA20" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="391">
<emphasis id="B90DEAF1EA47CF51FF71FF56FEDFCB8D" bold="true" box="[199,319,151,176]" pageId="10" pageNumber="391">Variation.</emphasis>
The general coloration and pattern of the
<typeStatus id="54C28841EA47CF51FC86FF59FC7CCB8D" box="[816,924,152,176]" pageId="10" pageNumber="391" type="paratype">paratypes</typeStatus>
closely match that of the
<typeStatus id="54C28841EA47CF51FB67FF56FAD3CB8D" box="[1233,1331,151,176]" pageId="10" pageNumber="391" type="holotype">holotype</typeStatus>
(
<figureCitation id="13422A66EA47CF51FAF0FF56FA6DCB8D" box="[1350,1421,151,176]" captionStart="FIGURE 3" captionStartId="8.[151,250,1301,1323]" captionTargetBox="[151,1436,176,1279]" captionTargetId="figure@8.[151,1436,169,1279]" captionTargetPageId="8" captionText="FIGURE 3. Cyrtodactylus metropolis sp. nov. Upper; holotype LSUHC 11347. Lower left; holotype LSUHC 11343. Lower right; paratype LSUHC 11342." httpUri="https://zenodo.org/record/230943/files/figure.png" pageId="10" pageNumber="391">Fig. 3</figureCitation>
). Dorsal bands of the
<typeStatus id="54C28841EA47CF51FE34FF7CFE0DCBE8" box="[386,493,189,213]" pageId="10" pageNumber="391" type="paratype">paratypes</typeStatus>
do not bifurcate. LSUHC 11345 has a regenerated tail bearing a unicolor grayish coloration. LSUHC 11342 is a subadult female and has a more bold color pattern (
<figureCitation id="13422A66EA47CF51FB81FF1EFB9CCBC5" box="[1079,1148,223,248]" captionStart="FIGURE 3" captionStartId="8.[151,250,1301,1323]" captionTargetBox="[151,1436,176,1279]" captionTargetId="figure@8.[151,1436,169,1279]" captionTargetPageId="8" captionText="FIGURE 3. Cyrtodactylus metropolis sp. nov. Upper; holotype LSUHC 11347. Lower left; holotype LSUHC 11343. Lower right; paratype LSUHC 11342." httpUri="https://zenodo.org/record/230943/files/figure.png" pageId="10" pageNumber="391">Fig. 3</figureCitation>
). Meristic differences are shown in
<tableCitation id="C6FB0358EA47CF51FEB2FEC5FEB9CA20" box="[260,345,260,285]" captionStart="TABLE 6" captionStartId="9.[151,242,548,571]" captionTargetBox="[159,1416,636,2011]" captionTargetPageId="9" captionText="TABLE 6. Morphological characters and selected maesurements (in mm) of the type series of Cyrtodactylus metropolis sp. nov. / = data unavailable. Abbreviations are listed in the Materials and Methods." httpUri="http://table.plazi.org/id/DF06666BEA44CF52FF21FDE5FBEBC963" pageId="10" pageNumber="391" tableUuid="DF06666BEA44CF52FF21FDE5FBEBC963">Table 6</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3636568EA47CF51FF71FEE6FAF6CC2E" pageId="10" pageNumber="391" type="distribution">
<paragraph id="8BC636E3EA47CF51FF71FEE6FB58CA58" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="391">
<emphasis id="B90DEAF1EA47CF51FF71FEE6FE80CA7D" bold="true" box="[199,352,295,320]" pageId="10" pageNumber="391">Distribution.</emphasis>
<taxonomicName id="4C794D60EA47CF51FEDDFEE9FD6BCA7D" box="[363,651,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA47CF51FEDDFEE9FD6BCA7D" box="[363,651,296,320]" italics="true" pageId="10" pageNumber="391">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA47CF51FD23FEE9FD13CA7D" box="[661,755,296,320]" pageId="10" pageNumber="391" rank="species">
<emphasis id="B90DEAF1EA47CF51FD23FEE9FD13CA7D" bold="true" box="[661,755,296,320]" pageId="10" pageNumber="391">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the Batu Caves massif (
<figureCitation id="13422A66EA47CF51FABDFEE6FAB4CA7D" box="[1291,1364,295,320]" captionStart="FIGURE 4" captionStartId="10.[151,250,1510,1532]" captionTargetBox="[290,1299,661,1475]" captionTargetId="figure@10.[243,1344,661,1489]" captionTargetPageId="10" captionText="FIGURE 4. Distribution of the species of the Cyrtodactylus semenanjungensis complex." httpUri="https://zenodo.org/record/230944/files/figure.png" pageId="10" pageNumber="391">Fig. 4</figureCitation>
). It is expected not to be found elsewhere, as there is no known limestone forest in the near vicinity.
</paragraph>
<paragraph id="8BC636E3EA47CF51FF71FEB1FC10C95D" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="391">
<emphasis id="B90DEAF1EA47CF51FF71FEB1FE68CAB5" bold="true" box="[199,392,367,392]" pageId="10" pageNumber="391">Natural history.</emphasis>
Nineteen lizards were observed at night on karst and in the surrounding limestone forest vegetation up to two meters above the ground (
<figureCitation id="13422A66EA47CF51FD0BFE55FCE4CA90" box="[701,772,404,429]" captionStart="FIGURE 5" captionStartId="11.[151,250,1300,1322]" captionTargetBox="[151,1435,375,1277]" captionTargetId="figure@11.[151,1436,375,1277]" captionTargetPageId="11" captionText="FIGURE 5. Habitat structure near the type locality of Cyrtodactylus metropolis sp. nov." httpUri="https://zenodo.org/record/230945/files/figure.png" pageId="10" pageNumber="391">Fig. 5</figureCitation>
).
<taxonomicName id="4C794D60EA47CF51FCAAFE55FBDACA91" box="[796,1082,404,428]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA47CF51FCAAFE55FBDACA91" box="[796,1082,404,428]" italics="true" pageId="10" pageNumber="391">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA47CF51FBF2FE55FB40CA91" box="[1092,1184,404,428]" pageId="10" pageNumber="391" rank="species">
<emphasis id="B90DEAF1EA47CF51FBF2FE55FB40CA91" bold="true" box="[1092,1184,404,428]" pageId="10" pageNumber="391">sp. nov.</emphasis>
</taxonomicNameLabel>
is an extremely wary species that flees at the slightest provocation. In nearly all cases, lizards retreated to the base of a tree or rock and sought refuge in accumulated debris. One lizard ran nearly two meters along a narrow branch to reach the base of a fern, although other apparently suitable retreats were much closer. This suggests that the lizard had spatial awareness of its surroundings and periodically used that particular retreat. Juveniles were observed but hatchings and gravid females were not, indicating that this species breeds prior to July.
</paragraph>
<caption id="DF06666BEA47CF51FF21FA27FBFCCEC1" box="[151,1052,1510,1532]" httpUri="https://zenodo.org/record/230944/files/figure.png" pageId="10" pageNumber="391" targetBox="[290,1299,661,1475]" targetPageId="10">
<paragraph id="8BC636E3EA47CF51FF21FA27FBFCCEC1" blockId="10.[151,1052,1510,1532]" box="[151,1052,1510,1532]" pageId="10" pageNumber="391">
<emphasis id="B90DEAF1EA47CF51FF21FA27FEF3CEC1" bold="true" box="[151,275,1510,1532]" pageId="10" pageNumber="391">FIGURE 4.</emphasis>
Distribution of the species of the
<taxonomicName id="4C794D60EA47CF51FDDCFA27FC57CEC1" box="[618,951,1510,1532]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="semenanjungensis">
<emphasis id="B90DEAF1EA47CF51FDDCFA27FC57CEC1" box="[618,951,1510,1532]" italics="true" pageId="10" pageNumber="391">Cyrtodactylus semenanjungensis</emphasis>
</taxonomicName>
complex.
</paragraph>
</caption>
<paragraph id="8BC636E3EA47CF51FF71F9E2FAF6CC2E" blockId="10.[151,1437,1571,2028]" pageId="10" pageNumber="391">
The abundance of
<taxonomicName id="4C794D60EA47CF51FE17F9E2FD20CD06" box="[417,704,1571,1595]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA47CF51FE17F9E2FD20CD06" box="[417,704,1571,1595]" italics="true" pageId="10" pageNumber="391">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA47CF51FD7CF9E2FCC7CD06" box="[714,807,1571,1595]" pageId="10" pageNumber="391" rank="species">
<emphasis id="B90DEAF1EA47CF51FD7CF9E2FCC7CD06" bold="true" box="[714,807,1571,1595]" pageId="10" pageNumber="391">sp. nov.</emphasis>
</taxonomicNameLabel>
on the exterior of the Batu Caves massif and its rare occurrence in Dark Cave suggests it is not a cave-adapted species but will on occasion enter deeper regions. We have observed the same behavior in the limestone forest species
<taxonomicName id="4C794D60EA47CF51FCC9F9ADFD7CCD95" authority="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Norhayati, Bauer, Wangkulangkul" authorityName="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Norhayati, Bauer, Wangkulangkul" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="astrum">
<emphasis id="B90DEAF1EA47CF51FCC9F9ADFC10CDBE" box="[895,1008,1644,1667]" italics="true" pageId="10" pageNumber="391">C. astrum</emphasis>
Grismer, Wood, Quah, Anuar, Muin, Sumontha, Norhayati, Bauer, Wangkulangkul
</taxonomicName>
, &amp; Pauwels,
<taxonomicName id="4C794D60EA47CF51FC85F951FBFECDF1" authority="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Norhayati, Bauer, Wangkulangkul" authorityName="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Norhayati, Bauer, Wangkulangkul" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="langkawiensis">
<emphasis id="B90DEAF1EA47CF51FC85F951FC14CD9A" box="[819,1012,1679,1703]" italics="true" pageId="10" pageNumber="391">C. langkawiensis</emphasis>
Grismer, Wood, Quah, Anuar,
<bibRefCitation id="EFE84B12EA47CF51FAEEF94EFBFECDF1" pageId="10" pageNumber="391" refString="Grismer, L. L., Wood, P. L. Jr., Quah, E. S. H., Shahrul, A., Muin, M. A., Sumontha, M., Norhayati, A., Bauer, A. M., Wangkulangkul, S., Grismer, J. L. &amp; Pauwels, O. S. G. (2012 a) A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa, 3520, 1 - 55." type="journal article">
Muin, Sumontha, Norhayati, Bauer, Wangkulangkul, &amp; Pauwels, (Grismer
<emphasis id="B90DEAF1EA47CF51FC2AF975FC34CDF6" box="[924,980,1715,1739]" italics="true" pageId="10" pageNumber="391">et al.</emphasis>
2012)
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C794D60EA47CF51FBE3F975FE34CDCD" authority="Grismer, Belabut, Quah, Chan, Wood" authorityName="Grismer, Belabut, Quah, Chan, Wood" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="guakanthanensis">
<emphasis id="B90DEAF1EA47CF51FBE3F975FAD2CDF6" box="[1109,1330,1715,1739]" italics="true" pageId="10" pageNumber="391">C. guakanthanensis</emphasis>
Grismer, Belabut, Quah, Chan, Wood
</taxonomicName>
, &amp; Hasim (
<bibRefCitation id="EFE84B12EA47CF51FDE4F916FCD2CDCD" author="Grismer" box="[594,818,1751,1776]" pageId="10" pageNumber="391" refString="Grismer, L. L., Belabut, D. M., Quah, S. H., Chan, K. O., Wood, P. L. Jr. &amp; Hashim, R. (2014) A new species of karst forestadapted Bent-toed Gecko (Genus Cyrtodactylus) belonging to the C. sworderi complex from a threatened karst forest in Perak, Peninsular Malaysia. Zootaxa, 3755 (5), 434 - 446. http: // dx. doi. org / 10.11646 / zootaxa. 3755.5.3" type="journal article" year="2014">
Grismer
<emphasis id="B90DEAF1EA47CF51FD01F919FD0FCDD2" box="[695,751,1751,1775]" italics="true" pageId="10" pageNumber="391">et al.</emphasis>
2014
</bibRefCitation>
). It is likely the cave ecosystems are too depauperate in prey resources and vegetation to support the density levels of these species observed outside the caves.
</paragraph>
</subSubSection>
<subSubSection id="C3636568EA47CF50FF71F8E1FC9BCA7D" lastPageId="11" lastPageNumber="392" pageId="10" pageNumber="391" type="etymology">
<paragraph id="8BC636E3EA47CF51FF71F8E1FA7DCC61" blockId="10.[151,1437,1571,2028]" pageId="10" pageNumber="391">
<emphasis id="B90DEAF1EA47CF51FF71F8E1FEA6CC05" bold="true" box="[199,326,1824,1848]" pageId="10" pageNumber="391">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C794D60EA47CF51FDF2F8DEFD5BCC0A" box="[580,699,1823,1847]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="metropolis">
<emphasis id="B90DEAF1EA47CF51FDF2F8DEFD5BCC0A" box="[580,699,1823,1847]" italics="true" pageId="10" pageNumber="391">metropolis</emphasis>
</taxonomicName>
is a noun in apposition and refers to the fact that this endemic species is found in a highly urbanized area near the heart of the largest metropolitan center of Peninsular
<collectingCountry id="F36E7673EA47CF51FA98F882FA78CC61" box="[1326,1432,1859,1884]" name="Malaysia" pageId="10" pageNumber="391">Malaysia</collectingCountry>
.
</paragraph>
<paragraph id="8BC636E3EA47CF50FF71F8A6FC9BCA7D" blockId="10.[151,1437,1571,2028]" lastBlockId="11.[151,1436,151,320]" lastPageId="11" lastPageNumber="392" pageId="10" pageNumber="391">
<emphasis id="B90DEAF1EA47CF51FF71F8A6FE8DCCBD" bold="true" box="[199,365,1895,1920]" pageId="10" pageNumber="391">Comparisons.</emphasis>
<taxonomicName id="4C794D60EA47CF51FECFF8A6FD79CC42" box="[377,665,1895,1919]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA47CF51FECFF8A6FD79CC42" box="[377,665,1895,1919]" italics="true" pageId="10" pageNumber="391">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA47CF51FD13F8A9FCE4CCBD" box="[677,772,1896,1920]" pageId="10" pageNumber="391" rank="species">
<emphasis id="B90DEAF1EA47CF51FD13F8A9FCE4CCBD" bold="true" box="[677,772,1896,1920]" pageId="10" pageNumber="391">sp. nov.</emphasis>
</taxonomicNameLabel>
can be differentiated from all other members of the
<taxonomicName id="4C794D60EA47CF51FA34F8A9FE83CC9E" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="semenanjungensis">
<emphasis id="B90DEAF1EA47CF51FA34F8A9FE83CC9E" italics="true" pageId="10" pageNumber="391">C. semenanjungensis</emphasis>
</taxonomicName>
complex by having four or five narrow, light-colored, dorsal bands as opposed to having 4–6 wide dorsal bands or being blotched.
<taxonomicName id="4C794D60EA47CF51FD88F86EFCBBCCFA" box="[574,859,1967,1991]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="metropolis" status="sp. nov.">
<emphasis id="B90DEAF1EA47CF51FD88F86EFCBBCCFA" box="[574,859,1967,1991]" italics="true" pageId="10" pageNumber="391">Cyrtodactylus metropolis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E578AEA47CF51FCD4F871FC54CCFA" box="[866,948,1968,1991]" pageId="10" pageNumber="391" rank="species">sp. nov</taxonomicNameLabel>
is possibly further separated from all other species of the
<taxonomicName id="4C794D60EA47CF51FE80F815FDC2CCD6" box="[310,546,2004,2027]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="semenanjungensis">
<emphasis id="B90DEAF1EA47CF51FE80F815FDC2CCD6" box="[310,546,2004,2027]" italics="true" pageId="10" pageNumber="391">C. semenanjungensis</emphasis>
</taxonomicName>
complex by nine light-colored caudal bands on a regenerated tail as opposed to having 13–16. They can be additionally differentiated further by the having the unique combination of nine or 10 supralabials; seven or eight infralabials; 32–34 paravertebral tubercles; 37–44 ventral scales; no enlarged, median subcaudal scales; abrupt contact between the small and large postfemoral scales; 20 or 21 subdigital lamellae on the fourth toe; no deep, precloacal groove; no femoral pores; enlarged and continuous precloacal and femoral scales; and narrow, proximal light-colored caudal bands (
<tableCitation id="C6FB0358EA46CF50FCA1FEE6FC8DCA7D" box="[791,877,295,320]" captionStart="TABLE 5" captionStartId="7.[151,239,362,385]" captionText="TABLE 5. Diagnostic characters (bold) separating species of the Cyrtodactylus semenanjungensis complex from C. metropolis sp. nov." pageId="11" pageNumber="392">Table 5</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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