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<document id="3FFB09D64FB2C3B62F58F8AC36E6EA1C" ID-ISBN="978-2-85653-614-8" ID-ISSN="1243-4442" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630456027313" checkinUser="felipe" docAuthor="Valdés, Ángel" docDate="2008" docId="03D087B2FF50BE98FF0177B5F265FAEC" docLanguage="en" docName="09-Valdés.pdf" docOrigin="Mémoires du Muséum national d'Histoire naturelle 196" docStyle="DocumentStyle:18F5342D935943D6DB33581BB9A2E502.4:MemMusNatHistNatu.2010-.journal_article" docStyleId="18F5342D935943D6DB33581BB9A2E502" docStyleName="MemMusNatHistNatu.2010-.journal_article" docStyleVersion="4" docTitle="Retusidae Thiele 1925" docType="treatment" docVersion="2" lastPageNumber="769" masterDocId="FFE9FFCAFFE4BE2EFFF3766FF729FFE3" masterDocTitle="Deep-sea “ cephalaspidean ” heterobranchs (Gastropoda) from the tropical southwest Pacific" masterLastPageNumber="792" masterPageNumber="587" pageNumber="767" updateTime="1699277432387" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8268EC288B312534FBD6E8BB99EC189D">Deep-sea “ cephalaspidean ” heterobranchs (Gastropoda) from the tropical southwest Pacific</mods:title>
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<mods:namePart id="755842EF617D0D3A7810A311601FB04A">Valdés, Ángel</mods:namePart>
<mods:affiliation id="9FFCEBC3F5326D8F5A6E3926060BAFB1">John T. Huber</mods:affiliation>
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<mods:title id="2D4736286D3D2A489BF83E44A58CB9CE">Mémoires du Muséum national d'Histoire naturelle</mods:title>
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<mods:date id="5D6FD6D8790289BA51FF70EE69848425">2008</mods:date>
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<mods:identifier id="70EF85F79DC3D6F04B3259A5F423F33E" type="ISSN">1243-4442</mods:identifier>
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<treatment id="03D087B2FF50BE98FF0177B5F265FAEC" LSID="urn:lsid:plazi:treatment:03D087B2FF50BE98FF0177B5F265FAEC" httpUri="http://treatment.plazi.org/id/03D087B2FF50BE98FF0177B5F265FAEC" lastPageId="182" lastPageNumber="769" pageId="180" pageNumber="767">
<subSubSection id="C363652FFF50BE9AFF0177B5F578FE11" box="[242,593,474,498]" pageId="180" pageNumber="767" type="nomenclature">
<paragraph id="8BC636A4FF50BE9AFF0177B5F578FE11" blockId="180.[242,593,474,498]" box="[242,593,474,498]" pageId="180" pageNumber="767">
<heading id="D08E81C8FF50BE9AFF0177B5F578FE11" box="[242,593,474,498]" fontSize="36" level="2" pageId="180" pageNumber="767" reason="4">
Family
<taxonomicName id="4C794D27FF50BE9AFEB377B5F6EFFE11" ID-CoL="7NKTS" authorityName="Thiele" authorityYear="1925" box="[320,454,474,498]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="180" pageNumber="767" phylum="Mollusca" rank="family">RETUSIDAE</taxonomicName>
<emphasis id="B90DEAB6FF50BE9AFE3F77B4F578FE11" box="[460,593,475,498]" italics="true" pageId="180" pageNumber="767">incertae sedis</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE9AFF277470F65BFCE4" pageId="180" pageNumber="767" type="discussion">
<paragraph id="8BC636A4FF50BE9AFF277470F5BEFD9E" blockId="180.[212,1443,543,637]" pageId="180" pageNumber="767">
REMARKS. — In this section I have included two deep-water species, only known from their shells, for which I was unable to determine a generic placement. The shell morphology of these species resembles that of
<taxonomicName id="4C794D27FF50BE9AFB1A742DF278FDB9" authorityName="Thiele" authorityYear="1925" box="[1257,1361,578,602]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="180" pageNumber="767" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
so they have been placed tentatively in this family.
</paragraph>
<paragraph id="8BC636A4FF50BE9AFF0174A2F65BFCE4" blockId="180.[212,403,717,775]" pageId="180" pageNumber="767">
<taxonomicName id="4C794D27FF50BE9AFF0174A2F673FD06" authorityName="Thiele" authorityYear="1925" box="[242,346,717,741]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="180" pageNumber="767" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
sp. 1 Figs 91A, B, 92
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE9AFF27755EF399FCA4" box="[212,1200,817,839]" pageId="180" pageNumber="767" type="materials_examined">
<paragraph id="8BC636A4FF50BE9AFF27755EF399FCA4" blockId="180.[212,1200,817,839]" box="[212,1200,817,839]" pageId="180" pageNumber="767">
<collectionCode id="ED68AE61FF50BE9AFF27755EF669FCA5" box="[212,320,817,838]" pageId="180" pageNumber="767">MATERIAL</collectionCode>
EXAMINED. —
<emphasis id="B90DEAB6FF50BE9AFE2B755EF576FCA4" bold="true" box="[472,607,817,839]" pageId="180" pageNumber="767">Loyalty Ridge</emphasis>
. BIOGEOCAL: stn KG 294,
<geoCoordinate id="EE4D5063FF50BE9AFC9E755EF495FCA5" box="[877,956,817,838]" degrees="20" direction="south" minutes="39" orientation="latitude" pageId="180" pageNumber="767" precision="925" value="-20.65">20°39S</geoCoordinate>
,
<geoCoordinate id="EE4D5063FF50BE9AFC35755EF30BFCA4" box="[966,1058,817,839]" degrees="167" direction="east" minutes="12" orientation="longitude" pageId="180" pageNumber="767" precision="925" value="167.2">167°12E</geoCoordinate>
,
<quantity id="4C819B41FF50BE9AFBD8755EF35CFCA4" box="[1067,1141,817,839]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="180" pageNumber="767" unit="m" value="1420.0">1420 m</quantity>
, 1 dd.
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE9AFF277515F355FC72" box="[212,1148,890,914]" pageId="180" pageNumber="767" type="distribution">
<paragraph id="8BC636A4FF50BE9AFF277515F355FC72" blockId="180.[212,1148,890,914]" box="[212,1148,890,914]" pageId="180" pageNumber="767">
<materialsCitation id="3B113CF9FF50BE9AFF277515F351FC72" box="[212,1144,890,914]" collectionCode="DISTRIBUTION" country="New Caledonia" elevation="1420" location="Single" pageId="180" pageNumber="767" specimenCount="1">
<collectionCode id="ED68AE61FF50BE9AFF277515F6AFFC71" box="[212,390,890,914]" pageId="180" pageNumber="767">DISTRIBUTION</collectionCode>
. —
<location id="8EA6607FFF50BE9AFE567515F6CFFC71" LSID="urn:lsid:plazi:treatment:03D087B2FF50BE98FF0177B5F265FAEC:8EA6607FFF50BE9AFE567515F6CFFC71" box="[421,486,890,914]" country="New Caledonia" name="Single" pageId="180" pageNumber="767">Single</location>
specimen collected from
<collectingCountry id="F36E7634FF50BE9AFD047515F4B3FC71" box="[759,922,890,914]" name="New Caledonia" pageId="180" pageNumber="767">New Caledonia</collectingCountry>
(Fig. 92), in
<quantity id="4C819B41FF50BE9AFBD77515F351FC72" box="[1060,1144,890,914]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="180" pageNumber="767" unit="m" value="1420.0">
<elevation id="0054D197FF50BE9AFBD77515F351FC72" box="[1060,1144,890,914]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="180" pageNumber="767" unit="m" value="1420.0">1420 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE9AFF2775D0F4B1FB2A" pageId="180" pageNumber="767" type="description">
<paragraph id="8BC636A4FF50BE9AFF2775D0F20EFB44" blockId="180.[212,1444,959,1226]" pageId="180" pageNumber="767">
DESCRIPTION. —
<emphasis id="B90DEAB6FF50BE9AFE6A75AEF56EFC3B" box="[409,583,961,984]" italics="true" pageId="180" pageNumber="767">Shell morphology</emphasis>
. Length 4.5 mm, width 2.5 mm (single specimen examined). Shell fragile, oval to elongate, with convex sides (Fig. 91A). Only 1 whorl visible, forming the entire shell. Apex rounded, not umbilicate (Fig. 91B), with the aperture lip rising from the right side and forming a short wing that is connected to the columellar margin. Anterior end of the shell rounded. Aperture as long as the shell, wider anteriorly and narrowing abruptly at about 1/4 of its length. Columellar margin thickened. Columella smooth, with no folds. Umbilicus narrow, partially covered by an extension of the columella. Sculpture of a number of faint growth lines. Colour uniformly dirty white with a number of brown-reddish spots, more densely arranged near the posterior end of the shell.
</paragraph>
<paragraph id="8BC636A4FF50BE9AFF0172DDF4B1FB2A" blockId="180.[212,1444,959,1226]" box="[242,920,1202,1226]" pageId="180" pageNumber="767">
<emphasis id="B90DEAB6FF50BE9AFF0172DDF665FB2A" box="[242,332,1202,1225]" italics="true" pageId="180" pageNumber="767">Anatomy</emphasis>
. Unknown. The single shell collected lacked soft parts.
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE9AFF277298F651FA3C" pageId="180" pageNumber="767" type="discussion">
<paragraph id="8BC636A4FF50BE9AFF277298F4A6FAD1" blockId="180.[212,1444,1271,1365]" pageId="180" pageNumber="767">REMARKS. — The shell of this species is different from other deep-water taxa examined in this study and I have been unable to find species with a similar morphology in the literature.</paragraph>
<paragraph id="8BC636A4FF50BE9AFF017352F265FAB6" blockId="180.[212,1444,1271,1365]" box="[242,1356,1341,1365]" pageId="180" pageNumber="767">A generic placement for this species is not possible until complete specimens become available for study.</paragraph>
<paragraph id="8BC636A4FF50BE9AFF0173CAF651FA3C" blockId="180.[212,403,1445,1503]" pageId="180" pageNumber="767">
<taxonomicName id="4C794D27FF50BE9AFF0173CAF673FA5E" authorityName="Thiele" authorityYear="1925" box="[242,346,1445,1469]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="180" pageNumber="767" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
sp. 2 Figs 91C, D, 92
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE9AFF277066F358F9FC" box="[212,1137,1545,1568]" pageId="180" pageNumber="767" type="materials_examined">
<paragraph id="8BC636A4FF50BE9AFF277066F358F9FC" blockId="180.[212,1137,1545,1568]" box="[212,1137,1545,1568]" pageId="180" pageNumber="767">
<collectionCode id="ED68AE61FF50BE9AFF277066F669F9FD" box="[212,320,1545,1566]" pageId="180" pageNumber="767">MATERIAL</collectionCode>
EXAMINED. —
<materialsCitation id="3B113CF9FF50BE9AFE2B7066F344F9FC" box="[472,1133,1545,1568]" country="Philippines" elevation="2570" latitude="4.633333" location="Philippines" longLatPrecision="1306" longitude="119.816666" pageId="180" pageNumber="767" specimenCount="1">
<emphasis id="B90DEAB6FF50BE9AFE2B7066F562F9FC" bold="true" box="[472,587,1545,1567]" pageId="180" pageNumber="767">
<collectingCountry id="F36E7634FF50BE9AFE2B7066F562F9FC" box="[472,587,1545,1567]" name="Philippines" pageId="180" pageNumber="767">Philippines</collectingCountry>
</emphasis>
. ESTASE 2: stn CP 6,
<geoCoordinate id="EE4D5063FF50BE9AFCE87066F459F9FD" box="[795,880,1545,1566]" degrees="04" direction="north" minutes="38" orientation="latitude" pageId="180" pageNumber="767" precision="925" value="4.633333">04°38N</geoCoordinate>
,
<geoCoordinate id="EE4D5063FF50BE9AFC8A7066F4FDF9FD" box="[889,980,1545,1566]" degrees="119" direction="east" minutes="49" orientation="longitude" pageId="180" pageNumber="767" precision="925" value="119.816666">119°49E</geoCoordinate>
,
<quantity id="4C819B41FF50BE9AFC2D7065F300F9FC" box="[990,1065,1546,1568]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="180" pageNumber="767" unit="m" value="2570.0">
<elevation id="0054D197FF50BE9AFC2D7065F300F9FC" box="[990,1065,1546,1568]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="180" pageNumber="767" unit="m" value="2570.0">2570 m</elevation>
</quantity>
, 15 dd
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE9AFF27703DF3BFF98A" box="[212,1174,1618,1642]" pageId="180" pageNumber="767" type="distribution">
<paragraph id="8BC636A4FF50BE9AFF27703DF3BFF98A" blockId="180.[212,1174,1618,1642]" box="[212,1174,1618,1642]" pageId="180" pageNumber="767">
<materialsCitation id="3B113CF9FF50BE9AFF27703DF3BBF98A" box="[212,1170,1618,1642]" collectionCode="DISTRIBUTION" country="Philippines" elevation="2570" location="Material" pageId="180" pageNumber="767" specimenCount="1">
<collectionCode id="ED68AE61FF50BE9AFF27703DF6AFF989" box="[212,390,1618,1642]" pageId="180" pageNumber="767">DISTRIBUTION</collectionCode>
. —
<location id="8EA6607FFF50BE9AFE56703DF6D5F989" LSID="urn:lsid:plazi:treatment:03D087B2FF50BE98FF0177B5F265FAEC:8EA6607FFF50BE9AFE56703DF6D5F989" box="[421,508,1618,1642]" country="Philippines" name="Material" pageId="180" pageNumber="767">Material</location>
examined collected from the
<collectingCountry id="F36E7634FF50BE9AFCCA703DF498F989" box="[825,945,1618,1642]" name="Philippines" pageId="180" pageNumber="767">Philippines</collectingCountry>
(Fig. 92), in
<quantity id="4C819B41FF50BE9AFBCF703DF3BBF98A" box="[1084,1170,1618,1642]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="180" pageNumber="767" unit="m" value="2570.0">
<elevation id="0054D197FF50BE9AFBCF703DF3BBF98A" box="[1084,1170,1618,1642]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="180" pageNumber="767" unit="m" value="2570.0">2570 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF50BE98FF2770F8F47BFB67" lastPageId="182" lastPageNumber="769" pageId="180" pageNumber="767" type="description">
<paragraph id="8BC636A4FF50BE9AFF2770F8F28AF8D9" blockId="180.[212,1444,1687,1850]" pageId="180" pageNumber="767">
DESCRIPTION. —
<emphasis id="B90DEAB6FF50BE9AFE6870F6F560F953" box="[411,585,1689,1712]" italics="true" pageId="180" pageNumber="767">Shell morphology</emphasis>
. Length
<quantity id="4C819B41FF50BE9AFD5B70F8F5C0F94C" box="[680,745,1687,1711]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="180" pageNumber="767" unit="mm" value="4.0">4 mm</quantity>
, width
<quantity id="4C819B41FF50BE9AFCCA70F8F450F94C" box="[825,889,1687,1711]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="180" pageNumber="767" unit="mm" value="3.0">3 mm</quantity>
(largest specimen examined). Shell solid, oval, with convex sides (Fig. 91C). Only 1 whorl visible, forming nearly the entire shell, except for a visible portion of the top of the spire. Apex rounded, deeply umbilicate (Fig. 91D), with the aperture lip rising from the right side and forming a short wing that is connected to the columellar margin. Anterior end of the shell rounded. Aperture as long as the shell, wider anteriorly and narrowing abruptly at about 1/3 of its length. Columellar margin conspicuously
</paragraph>
<paragraph id="8BC636A4FF51BE9BFECB709DF64AF8DB" blockId="181.[312,1422,1778,1848]" pageId="181" pageNumber="768">
<emphasis id="B90DEAB6FF51BE9BFECB709DF65CF8E0" bold="true" box="[312,373,1778,1795]" pageId="181" pageNumber="768">FIG. 91</emphasis>
. Shell morphology of species of
<taxonomicName id="4C794D27FF51BE9BFD8D709DF5E2F8E0" authorityName="Thiele" authorityYear="1925" box="[638,715,1778,1795]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="181" pageNumber="768" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
<emphasis id="B90DEAB6FF51BE9BFD22709DF415F8E0" box="[721,828,1778,1795]" italics="true" pageId="181" pageNumber="768">incertae sedis</emphasis>
.
<emphasis id="B90DEAB6FF51BE9BFCBB709DF47CF8E0" bold="true" box="[840,853,1778,1795]" pageId="181" pageNumber="768">A</emphasis>
,
<taxonomicName id="4C794D27FF51BE9BFC92709DF487F8E0" authorityName="Thiele" authorityYear="1925" box="[865,942,1778,1795]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="181" pageNumber="768" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
sp. 1, (4.5 mm), ventral view,
<collectingCountry id="F36E7634FF51BE9BFB55709DF20AF8E0" box="[1190,1315,1778,1795]" name="New Caledonia" pageId="181" pageNumber="768">New Caledonia</collectingCountry>
, BIOGEOCAL stn KG 294;
<emphasis id="B90DEAB6FF51BE9BFE6D7162F680F8FD" bold="true" box="[414,425,1805,1822]" pageId="181" pageNumber="768">B</emphasis>
, same shell, posterior view.
<emphasis id="B90DEAB6FF51BE9BFD7F7162F5BEF8FD" bold="true" box="[652,663,1805,1822]" pageId="181" pageNumber="768">C</emphasis>
,
<taxonomicName id="4C794D27FF51BE9BFD517162F5C6F8FD" authorityName="Thiele" authorityYear="1925" box="[674,751,1805,1822]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="181" pageNumber="768" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
sp. 2, (
<quantity id="4C819B41FF51BE9BFCD87162F477F8FD" box="[811,862,1805,1822]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="181" pageNumber="768" unit="mm" value="4.0">4 mm</quantity>
), ventral view,
<collectingCountry id="F36E7634FF51BE9BFC267162F305F8FD" box="[981,1068,1805,1822]" name="Philippines" pageId="181" pageNumber="768">Philippines</collectingCountry>
, ESTASE 2 stn CP 6;
<emphasis id="B90DEAB6FF51BE9BFB227162F3F7F8FD" bold="true" box="[1233,1246,1805,1822]" pageId="181" pageNumber="768">D</emphasis>
, same shell, posterior view.
</paragraph>
<paragraph id="8BC636A4FF52BE98FF277248F697FB81" blockId="182.[212,1444,1063,1157]" pageId="182" pageNumber="769">thickened. Columella smooth, with no folds. Umbilicus closed. Sculpture of a number of faint growth lines. Colour uniformly dirty white.</paragraph>
<paragraph id="8BC636A4FF52BE98FF017202F47BFB67" blockId="182.[212,1444,1063,1157]" box="[242,850,1133,1157]" pageId="182" pageNumber="769">
<emphasis id="B90DEAB6FF52BE98FF017202F665FB67" box="[242,332,1133,1156]" italics="true" pageId="182" pageNumber="769">Anatomy</emphasis>
. Unknown. All shells collected lacked soft parts.
</paragraph>
</subSubSection>
<subSubSection id="C363652FFF52BE98FF2772DDF265FAEC" pageId="182" pageNumber="769" type="discussion">
<paragraph id="8BC636A4FF52BE98FF2772DDF6C9FB0E" blockId="182.[212,1444,1202,1295]" pageId="182" pageNumber="769">
REMARKS. — The shells of
<taxonomicName id="4C794D27FF52BE98FE0F72DDF54DFB29" authorityName="Thiele" authorityYear="1925" box="[508,612,1202,1226]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="182" pageNumber="769" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
sp. 2 resemble that of
<taxonomicName id="4C794D27FF52BE98FCAD72DDF4EFFB29" authorityName="Thiele" authorityYear="1925" box="[862,966,1202,1226]" class="Gastropoda" family="Retusidae" kingdom="Animalia" order="Cephalaspidea" pageId="182" pageNumber="769" phylum="Mollusca" rank="family">Retusidae</taxonomicName>
sp. 1, but the former are shorter, wider and have an apical umbilicus.
</paragraph>
<paragraph id="8BC636A4FF52BE98FF017298F265FAEC" blockId="182.[212,1444,1202,1295]" box="[242,1356,1271,1295]" pageId="182" pageNumber="769">A generic placement for this species is not possible until complete specimens become available for study.</paragraph>
</subSubSection>
</treatment>
</document>