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<document id="49AC8E9A3693948B217F8880F6AE9696" ID-CLB-Dataset="32105" ID-DOI="10.11646/zootaxa.4320.1.1" ID-GBIF-Dataset="6e46db2a-d938-4778-8ce1-f992f95049fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="891347" ID-ZooBank="11D5D943-C39D-4871-Aa71-8640C4026607" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505404278194" checkinUser="plazi" docAuthor="Palestrini, Claudia" docDate="2017" docId="0393DF7AFFDFFFF6FF51FC25FBDB87AE" docLanguage="en" docName="zooaxa.4320.1.1.pdf.imf" docOrigin="Zootaxa 4320 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epidrepanus Roggero, Barbero, &amp; Palestrini 2015" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FFAAA702FFD8FFF1FFC6FFF7FF968104" masterDocTitle="hips of Epidrepanus within the subtribe Drepanocerina (Coleoptera: Scarabaeidae: Scarabaeinae: Oniticellini), with the description of two new species" masterLastPageNumber="24" masterPageNumber="1" pageNumber="8" updateTime="1698468432061" updateUser="ExternalLinkService">
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<mods:title id="D70D80D87C605526303AB38D4C889AB3">hips of Epidrepanus within the subtribe Drepanocerina (Coleoptera: Scarabaeidae: Scarabaeinae: Oniticellini), with the description of two new species</mods:title>
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<mods:namePart id="D2AAB29581F6EFB64EF25B26F72084A0">Palestrini, Claudia</mods:namePart>
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<treatment id="0393DF7AFFDFFFF6FF51FC25FBDB87AE" ID-DOI="http://doi.org/10.5281/zenodo.5999290" ID-GBIF-Taxon="134089548" ID-Zenodo-Dep="5999290" LSID="urn:lsid:plazi:treatment:0393DF7AFFDFFFF6FF51FC25FBDB87AE" httpUri="http://treatment.plazi.org/id/0393DF7AFFDFFFF6FF51FC25FBDB87AE" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="C3203DE7FFDFFFF6FF51FC25FD018536" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B856E6CFFDFFFF6FF51FC25FCF682E8" blockId="7.[151,864,978,1004]" box="[151,864,978,1004]" pageId="7" pageNumber="8">
<heading id="D0CDD900FFDFFFF6FF51FC25FCF682E8" bold="true" box="[151,864,978,1004]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B94EB27EFFDFFFF6FF51FC25FCF682E8" bold="true" box="[151,864,978,1004]" pageId="7" pageNumber="8">
Genus
<taxonomicName id="4C3A15EFFFDFFFF6FF28FC25FCF682E8" ID-CoL="4D3J" authority="Roggero, Barbero, &amp; Palestrini, 2015" authorityName="Roggero, Barbero, &amp; Palestrini" authorityYear="2015" box="[238,864,978,1004]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94EB27EFFDFFFF6FF28FC25FE1F82E8" bold="true" box="[238,393,978,1004]" italics="true" pageId="7" pageNumber="8">Epidrepanus</emphasis>
<bibRefCitation id="EFAB139DFFDFFFF6FE56FC25FCF682E8" author="Roggero" box="[400,864,978,1004]" pageId="7" pageNumber="8" refString="Roggero, A., Barbero, E. &amp; Palestrini, C. (2015) Phylogenetic and biogeographical review of the Drepanocerina (Coleoptera: Scarabaeidae: Oniticellini). Arthropod Systematics and Phylogeny, 73, 153 - 174." type="journal article" year="2015">Roggero, Barbero, &amp; Palestrini, 2015</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B856E6CFFDFFFF6FF51FBEEFD018536" blockId="7.[151,1118,1049,1240]" box="[151,663,1049,1074]" pageId="7" pageNumber="8">
The genus
<taxonomicName id="4C3A15EFFFDFFFF6FED4FBEEFE368536" authorityName="Roggero, Barbero, &amp; Palestrini" authorityYear="2015" box="[274,416,1049,1074]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94EB27EFFDFFFF6FED4FBEEFE368536" box="[274,416,1049,1074]" italics="true" pageId="7" pageNumber="8">Epidrepanus</emphasis>
</taxonomicName>
includes five species:
</paragraph>
</subSubSection>
<subSubSection id="C3203DE7FFDFFFF6FF51FBCDFBDB87AE" pageId="7" pageNumber="8" type="description">
<paragraph id="8B856E6CFFDFFFF6FF51FBCDFDFB8557" blockId="7.[151,1118,1049,1240]" box="[151,621,1082,1107]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A15EFFFDFFFF6FF51FBCDFDFB8557" authority="(Gerstaecker, 1871)" authorityName="Gerstaecker" authorityYear="1871" baseAuthorityName="Gerstaecker" baseAuthorityYear="1871" box="[151,621,1082,1107]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="caelatus">
<emphasis id="B94EB27EFFDFFFF6FF51FBCDFE1F8557" box="[151,393,1082,1107]" italics="true" pageId="7" pageNumber="8">Epidrepanus caelatus</emphasis>
(Gerstaecker, 1871)
</taxonomicName>
</paragraph>
<paragraph id="8B856E6CFFDFFFF6FF51FBABFDE585B3" blockId="7.[151,1118,1049,1240]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A15EFFFDFFFF6FF51FBABFCA38571" authority="Roggero, Stanbrook, Josso, Barbero" authorityName="Roggero, Stanbrook, Josso, Barbero" box="[151,821,1116,1141]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kenyensis">
<emphasis id="B94EB27EFFDFFFF6FF51FBABFE018571" box="[151,407,1116,1141]" italics="true" pageId="7" pageNumber="8">Epidrepanus kenyensis</emphasis>
Roggero, Stanbrook, Josso, Barbero
</taxonomicName>
, &amp; Palestrini,
<emphasis id="B94EB27EFFDFFFF6FC12FBABFBC88571" bold="true" box="[980,1118,1116,1141]" pageId="7" pageNumber="8">new species</emphasis>
<taxonomicName id="4C3A15EFFFDFFFF6FF51FB8AFC908592" authority="Roggero, Stanbrook, Josso, Barbero" authorityName="Roggero, Stanbrook, Josso, Barbero" box="[151,774,1149,1174]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nyika">
<emphasis id="B94EB27EFFDFFFF6FF51FB8AFEFF8592" box="[151,361,1149,1174]" italics="true" pageId="7" pageNumber="8">Epidrepanus nyika</emphasis>
Roggero, Stanbrook, Josso, Barbero
</taxonomicName>
, &amp; Palestrini,
<emphasis id="B94EB27EFFDFFFF6FC60FB8AFBA68592" bold="true" box="[934,1072,1149,1174]" pageId="7" pageNumber="8">new species</emphasis>
<taxonomicName id="4C3A15EFFFDFFFF6FF51FB69FDE585B3" authority="(Balthasar, 1963)" authorityName="Balthasar" authorityYear="1963" baseAuthorityName="Balthasar" baseAuthorityYear="1963" box="[151,627,1182,1207]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pulvinarius">
<emphasis id="B94EB27EFFDFFFF6FF51FB69FE3A85B3" box="[151,428,1182,1207]" italics="true" pageId="7" pageNumber="8">Epidrepanus pulvinarius</emphasis>
(Balthasar, 1963)
</taxonomicName>
</paragraph>
<paragraph id="8B856E6CFFDFFFF6FF51FB48FDCC85DC" blockId="7.[151,1118,1049,1240]" box="[151,602,1215,1240]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A15EFFFDFFFF6FF51FB48FDCC85DC" authority="(Janssens, 1953)" authorityName="Janssens" authorityYear="1953" baseAuthorityName="Janssens" baseAuthorityYear="1953" box="[151,602,1215,1240]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="schimperi">
<emphasis id="B94EB27EFFDFFFF6FF51FB48FE0C85DC" box="[151,410,1215,1240]" italics="true" pageId="7" pageNumber="8">Epidrepanus schimperi</emphasis>
(Janssens, 1953)
</taxonomicName>
</paragraph>
<paragraph id="8B856E6CFFDFFFF6FF51FAF1FBDB87AE" blockId="7.[151,1437,1285,1706]" pageId="7" pageNumber="8">
A detailed comparative diagnosis of external and internal morphological traits for
<taxonomicName id="4C3A15EFFFDFFFF6FBF0FAF2FB52841A" authorityName="Roggero, Barbero, &amp; Palestrini" authorityYear="2015" box="[1078,1220,1285,1310]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94EB27EFFDFFFF6FBF0FAF2FB52841A" box="[1078,1220,1285,1310]" italics="true" pageId="7" pageNumber="8">Epidrepanus</emphasis>
</taxonomicName>
is given in Table 3 (
<emphasis id="B94EB27EFFDFFFF6FF66FADDFF558445" box="[160,195,1322,1345]" italics="true" pageId="7" pageNumber="8">i.e.</emphasis>
, identification keys of the
<taxonomicName id="4C3A15EFFFDFFFF6FE2BFADEFDED8446" authorityName="Roggero, Barbero, &amp; Palestrini" authorityYear="2015" box="[493,635,1321,1346]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94EB27EFFDFFFF6FE2BFADEFDED8446" box="[493,635,1321,1346]" italics="true" pageId="7" pageNumber="8">Epidrepanus</emphasis>
</taxonomicName>
species). While some structures (
<emphasis id="B94EB27EFFDFFFF6FC32FADDFB8B8445" box="[1012,1053,1322,1345]" italics="true" pageId="7" pageNumber="8">e.g.</emphasis>
, the carina of the head) can give a rather accurate, although not always conclusive, specific recognition, some others (the epipharynx and hindwing) are extremely well differentiated at the generic level in Drepanocerina, but their morphological variations at a specific level can be better appraised using geometric morphometric methods. Within the
<taxonomicName id="4C3A15EFFFDFFFF6FB43FA62FA8584AA" authorityName="Roggero, Barbero, &amp; Palestrini" authorityYear="2015" box="[1157,1299,1429,1454]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94EB27EFFDFFFF6FB43FA62FA8584AA" box="[1157,1299,1429,1454]" italics="true" pageId="7" pageNumber="8">Epidrepanus</emphasis>
</taxonomicName>
species, the structures can sometimes show variability between the sexes, and must be used with caution in the analyses. For example, although the head carina is a useful character, it is subject to sexual dimorphism in both
<taxonomicName id="4C3A15EFFFDFFFF6FB15FA29FAD884F2" baseAuthorityName="Gerstaecker" baseAuthorityYear="1871" box="[1235,1358,1501,1526]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="caelatus">
<emphasis id="B94EB27EFFDFFFF6FB15FA29FAD884F2" box="[1235,1358,1501,1526]" italics="true" pageId="7" pageNumber="8">E. caelatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A15EFFFDFFFF6FA42FA29FE81871E" baseAuthorityName="Balthasar" baseAuthorityYear="1963" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pulvinarius">
<emphasis id="B94EB27EFFDFFFF6FA42FA29FE81871E" italics="true" pageId="7" pageNumber="8">E. pulvinarius</emphasis>
</taxonomicName>
, but not in
<taxonomicName id="4C3A15EFFFDFFFF6FE5DF9F5FDB1871E" box="[411,551,1537,1562]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kenyensis">
<emphasis id="B94EB27EFFDFFFF6FE5DF9F5FDB1871E" box="[411,551,1537,1562]" italics="true" pageId="7" pageNumber="8">E. kenyensis</emphasis>
</taxonomicName>
. No information could be gained from
<taxonomicName id="4C3A15EFFFDFFFF6FC21F9F5FBE3871E" baseAuthorityName="Janssens" baseAuthorityYear="1953" box="[999,1141,1537,1562]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="schimperi">
<emphasis id="B94EB27EFFDFFFF6FC21F9F5FBE3871E" box="[999,1141,1537,1562]" italics="true" pageId="7" pageNumber="8">E. schimperi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A15EFFFDFFFF6FB68F9F5FA9C871E" box="[1198,1290,1537,1562]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nyika">
<emphasis id="B94EB27EFFDFFFF6FB68F9F5FA9C871E" box="[1198,1290,1537,1562]" italics="true" pageId="7" pageNumber="8">E. nyika</emphasis>
</taxonomicName>
, due to each species only being known from one sex (Table 3). The head carina was hence discarded from the present analysis. The genus is characterized by a widespread distribution in the Afrotropical region (
<figureCitation id="130172E9FFDFFFF6FBF2F9BEFB118766" box="[1076,1159,1609,1634]" captionStart="FIGURE 25" captionStartId="6.[151,250,1968,1990]" captionTargetBox="[190,1306,1163,1945]" captionTargetId="figure@6.[162,1426,1163,1956]" captionTargetPageId="6" captionText="FIGURE 25. Map of the distribution of Epidrepanus caelatus (green dots), E. nyika new species (purple rhomb), E. kenyensis new species (blue rhomb), E. pulvinarius (orange triangles) and E. schimperi (red star)." httpUri="https://zenodo.org/record/891359/files/figure.png" pageId="7" pageNumber="8">Fig. 25</figureCitation>
), and is typically present in savannah habitat.
<taxonomicName id="4C3A15EFFFDFFFF6FEB8F99AFDC68782" box="[382,592,1645,1670]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nyika">
<emphasis id="B94EB27EFFDFFFF6FEB8F99AFDC68782" box="[382,592,1645,1670]" italics="true" pageId="7" pageNumber="8">Epidrepanus nyika</emphasis>
</taxonomicName>
inhabits the Nyika Plateau (
<quantity id="4CC2C389FFDFFFF6FC4BF99AFBB88782" box="[909,1070,1645,1670]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="7" pageNumber="8" unit="m" value="2350.0" valueMax="2600.0" valueMin="2100.0">2100–2600 m</quantity>
), and
<taxonomicName id="4C3A15EFFFDFFFF6FBB4F999FB6A8782" box="[1138,1276,1645,1670]" class="Insecta" family="Scarabaeidae" genus="Epidrepanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kenyensis">
<emphasis id="B94EB27EFFDFFFF6FBB4F999FB6A8782" box="[1138,1276,1645,1670]" italics="true" pageId="7" pageNumber="8">E. kenyensis</emphasis>
</taxonomicName>
was collected from a high-elevation, open-canopy area in the Aberdare Mountains (
<quantity id="4CC2C389FFDFFFF6FC66F966FBD787AE" box="[928,1089,1681,1706]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">2000–2500 m</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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