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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.100798" ID-Pensoft-Pub="1313-2970-1168-295" ID-Pensoft-UUID="F19054F21F3D516FA0292C896F566182" ID-ZooBank="745FFCD5E3E84DC1B76CC82AD1AC5C64" ModsDocID="1313-2970-1168-295" checkinTime="1688439520051" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Zhao, Lingchen &amp; Zhang, Feng" docDate="2023" docId="ACB5C00C722F5B12AED0FAD4DA2DE82E" docLanguage="en" docName="ZooKeys 1168: 295-327" docOrigin="ZooKeys 1168" docPubDate="2023-07-03" docSource="http://dx.doi.org/10.3897/zookeys.1168.100798" docTitle="Catatemnus huae Hou &amp; Zhao &amp; Zhang 2023, sp. nov." docType="treatment" docUuid="3046D213-0100-4492-8883-CE96EE2393C9" docUuidSource="ZooBank" docVersion="1" id="F19054F21F3D516FA0292C896F566182" lastPageNumber="295" masterDocId="F19054F21F3D516FA0292C896F566182" masterDocTitle="First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)" masterLastPageNumber="327" masterPageNumber="295" pageNumber="295" updateTime="1688439520051" updateUser="pensoft">
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<mods:title id="47EB5510C4C8F5671ACAAAC92DF8C8B0">First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)</mods:title>
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<mods:namePart id="88BA8E185D10AFCF33E1678C7A9BEE1C">Hou, Yanmeng</mods:namePart>
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<mods:affiliation id="650A35E05D6435270F137220B828E317">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="DF5ADA45254761E2980AFA06493B3BA1">Zhao, Lingchen</mods:namePart>
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<mods:affiliation id="DBB61233635D320624EDD139D2841EEB">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China &amp; Changzhi Tunliu District No. 6 Middle School, Changzhi, Shanxi 046100, China</mods:affiliation>
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<mods:namePart id="7EC4AABE8F86E0DE966E0488BFC11179">Zhang, Feng</mods:namePart>
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<mods:affiliation id="60496DC0EEFE45F52CB9928D19D31B27">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<paragraph id="FB88D1A7D1C01BAEBA8942E01ADE6BB6" pageId="0" pageNumber="295">
<taxonomicName id="92CC50C3E050678F97013ED408FB2C8C" LSID="https://zoobank.org/3046D213-0100-4492-8883-CE96EE2393C9" authority="Hou &amp; Zhao &amp; Zhang, 2023" authorityName="Hou &amp; Zhao &amp; Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus huae" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="huae" status="sp. nov.">Catatemnus huae</taxonomicName>
<taxonomicNameLabel id="DF7964464F276C9794C532B2C21CCBFF" pageId="0" pageNumber="295">sp. nov.</taxonomicNameLabel>
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<paragraph id="6C155240B4339D9E72FC060F794BF103" pageId="0" pageNumber="295">
<figureCitation id="E1297B73BC3F7867D74E53D475D13315" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Catatemnus huae sp. nov. A holotype male, habitus (minus left pedipalp, left legs I and IV, dorsal view) B paratype female, habitus (minus left pedipalp, left legs I and III, right leg IV, dorsal view). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1168.100798.figure1" httpUri="https://binary.pensoft.net/fig/872299" pageId="0" pageNumber="295">Figs 1</figureCitation>
<figureCitation id="C5B15F79572A7BEBA7B0B868E57C54C2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">, 2</figureCitation>
<figureCitation id="B9BC07B1E9F40695F205BA75B964C3BD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">, 3 (胡氏猫伪蝎)</figureCitation>
</paragraph>
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<subSubSection id="D962D404C58064376B959303ED962388" pageId="0" pageNumber="295" type="type material">
<paragraph id="25CBA428C36338D7C1797D094486834C" pageId="0" pageNumber="295">Type material.</paragraph>
<paragraph id="BCC0B3AF904A87BF387E6FB9B66EA6A9" pageId="0" pageNumber="295">
<emphasis id="9B830EC79443DB0C1244A30FAC79EA78" bold="true" italics="true" pageId="0" pageNumber="295">Holotype</emphasis>
: China • ♂; Hainan Province, Lingshui County, Xincun Town, Tonghai Village;
<geoCoordinate id="C88FBC03876A2E35D334E73AD075856B" degrees="18" direction="north" minutes="26.084" orientation="latitude" precision="1" value="18.434732">18°26.084'N</geoCoordinate>
,
<geoCoordinate id="5F79FA41C8062CAF23ADC28A33C1A167" degrees="110" direction="east" minutes="0.495" orientation="longitude" precision="1" value="110.00825">110°0.495'E</geoCoordinate>
; 10 m a.s.l.; 19 May. 2011; Junfang Hu leg.; Ps.-MHBU-HA11051901.
<emphasis id="95E078A112EE0874262D58EA5DC7EAE5" bold="true" italics="true" pageId="0" pageNumber="295">Paratype</emphasis>
: • 1♀; same data as the holotype; Ps.-MSWU-HA11051902.
</paragraph>
</subSubSection>
<subSubSection id="6352E6C649C6CBC95B64A57A3E241585" pageId="0" pageNumber="295" type="diagnosis">
<paragraph id="465574199094A4EE62D9A8F4B866B3EE" pageId="0" pageNumber="295">Diagnosis (♂♀).</paragraph>
<paragraph id="658FD133B40B987CC4C839E5D7D7B48F" pageId="0" pageNumber="295">
This new species is characterized by carapace smooth but with a faint medial furrow; with two faint eyespots situated near anterior margin of carapace. Palpal stout, femur (♂) 2.48
<normalizedToken id="D1743DDA888CA1C0F624B877DFF053E8" originalValue="×">x</normalizedToken>
, (♀) 2.52
<normalizedToken id="689AD6DA675D26F1705023124D781EA8" originalValue="×">x</normalizedToken>
, chela with pedicel (♂) 3.05
<normalizedToken id="97BC9EA127EE06D69D3B2762AE832025" originalValue="×">x</normalizedToken>
, (♀) 2.93
<normalizedToken id="C63D92F6E9DD82FA20DF2B39DA4BF1B2" originalValue="×">x</normalizedToken>
, without pedicel (♂) 2.81
<normalizedToken id="D16CFB0E33BEFEBB9157642B5DD0CBB6" originalValue="×">x</normalizedToken>
, (♀) 2.74
<normalizedToken id="F0C7DC1DD6FD46B02AE6BB2CF40006AE" originalValue="×">x</normalizedToken>
longer than broad; the retrolateral surface of trochanter, the prolateral surface of femur, patella and hand granular; male genitalia: the hooked branch (br) relatively small.
</paragraph>
</subSubSection>
<subSubSection id="F361FA1732880C9EC0F9BF21A1960113" pageId="0" pageNumber="295" type="description">
<paragraph id="D0DDDA449AFBC05444BA034C203E17C8" pageId="0" pageNumber="295">Description.</paragraph>
<paragraph id="E152265945A97C5429FFCED362D9F2FA" pageId="0" pageNumber="295">
<emphasis id="084E358EED5F024E674238395D7DE9F7" bold="true" pageId="0" pageNumber="295">Adult male</emphasis>
(Figs
<figureCitation id="D2AA12D26629007549E3552276C89544" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Catatemnus huae sp. nov. A holotype male, habitus (minus left pedipalp, left legs I and IV, dorsal view) B paratype female, habitus (minus left pedipalp, left legs I and III, right leg IV, dorsal view). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1168.100798.figure1" httpUri="https://binary.pensoft.net/fig/872299" pageId="0" pageNumber="295">1A</figureCitation>
,
<figureCitation id="6FA76C898CC9F0677D2919BE0F92B4AD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2A-E, G-I</figureCitation>
,
<figureCitation id="8B482142F06ED1CB102F7F8CDC7E7337" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3A-H, J-L</figureCitation>
).
<emphasis id="CF53E6EEA675A23FE54D7F8C7B452A24" bold="true" italics="true" pageId="0" pageNumber="295">Color</emphasis>
: anterior half of carapace yellowish brown, paler in posterior half; pedipalps reddish brown, paler in female; tergites brown, paler in legs; remainder yellow.
<emphasis id="D3558BD048D1918A9D40D3F81800E230" bold="true" italics="true" pageId="0" pageNumber="295">Carapace</emphasis>
(Figs
<figureCitation id="6E3D1063A6A5BD5D722864FE618B7748" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2A</figureCitation>
,
<figureCitation id="FC3F626FE8A16C8CC0BB527C49971409" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3A</figureCitation>
): anterior half slightly darker than posterior half; surface smooth, but with a faint medial furrow; with two faint eyespots situated near anterior margin of carapace; anterior margin with six setae, posterior margin with seven setae, 47 in total, each seta acicular and very slightly curved; with a pair of lyrifissures on anterior margin and posterior margin, respectively.
<emphasis id="2928C975B8E471496AE42F2FCEA93099" bold="true" italics="true" pageId="0" pageNumber="295">Chelicera</emphasis>
(Fig.
<figureCitation id="A9BF14A4301B4D4944748797ED9FA63D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3B, C, E</figureCitation>
): much smaller than carapace length; surface smooth; four setae (sbs absent; bs shorter than others) and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand; movable finger with one galeal seta; bs and es dentate apically, is and ls long and acute. Fixed finger with six large retrorse teeth and three small apical teeth, movable finger with a long broadly rounded subapical lobe; galea present, slender, apically with five branchlets (Fig.
<figureCitation id="65CAA82CAAA68B85EB6B879B63BF033B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3C</figureCitation>
). Serrula interior connected to fixed finger for entire length, proximally modified to form velum, serrula exterior with 26 lamellae, the basal one longest; lamina exterior present. Rallum composed of four blades, the basal two blades shorter than others, the distal one dentated anteriorly, remainder smooth (Fig.
<figureCitation id="F737AE3CE6866B8E19A54996E7940C75" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3E</figureCitation>
).
<emphasis id="9EB4647F4B3C9E1B2676C5C82FBF5BC3" bold="true" italics="true" pageId="0" pageNumber="295">Pedipalp</emphasis>
(Figs
<figureCitation id="6BD243AB4288CF0F7CD9ED1AD5B8D79C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2B-D</figureCitation>
,
<figureCitation id="5C98FB9B436C16001C112E46444C858E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3F, H</figureCitation>
): stout, trochanter 1.85
<normalizedToken id="A3B87AF2A520B27BAD3F0F3B05E7F95A" originalValue="×">x</normalizedToken>
, femur 2.48
<normalizedToken id="7487916476526DC7F6F6E07C710F3484" originalValue="×">x</normalizedToken>
, patella 2.14
<normalizedToken id="606B2DA73F9736B0FC227074B47DF227" originalValue="×">x</normalizedToken>
, chela with pedicel (without pedicel) 3.05
<normalizedToken id="00F228C9BD0AC8C385858F4B8ECDCAEA" originalValue="×">x</normalizedToken>
(2.81
<normalizedToken id="272EC0E38305FD5CBDC8A1400A7B6A23" originalValue="×">x</normalizedToken>
), hand with pedicel (without pedicel) 1.79
<normalizedToken id="75552AC91F1C631AF2A3E2B6EF4E05E8" originalValue="×">x</normalizedToken>
(1.55
<normalizedToken id="A7D5DE465045AF961801948F926CD5E5" originalValue="×">x</normalizedToken>
) longer than broad; movable chelal finger 0.71
<normalizedToken id="AAFBF7264395F7AE8B6E85AC08A2A0E6" originalValue="×">x</normalizedToken>
(0.82
<normalizedToken id="4EAD449F316F0C084E6561D0D0CEBE5E" originalValue="×">x</normalizedToken>
) longer than hand with pedicel (without pedicel). Setae generally long and acuminate. Retrolateral surface of trochanter, the prolateral surface of femur, patella, and hand granular; trochanter with a rounded ventral tubercle and a well-developed conical dorsal tubercle. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria: eb and esb situated at base of fixed finger on retrolateral face, esb slightly distal to eb; ib and isb situated at base of fixed finger on prolateral face, isb slightly distal to ib; est in the middle of fixed finger; et near sub-distal of fixed finger; est closer to et than to esb; it slightly distal to est and proximal to et; ist situated proximal to est and distal to isb; it closer to ist than to fingertip; distance between est and esb farther to that of ist and isb, distance between it and fingertip nearly shorter to distance between ist and isb; b and sb situated at base of movable finger on retrolateral face; t in the middle of movable finger and at same level as est; sb closer to b than to st; st closer to t than to sb (Fig.
<figureCitation id="7CAF849BF34178F737BBADEAFD7609F1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3H</figureCitation>
). Venom apparatus only present in fixed chelal finger, venom ducts curved and short, terminating in inflated nodus ramosus between et and est and closer to est. Both chelal fingers with a row of acute teeth, spaced contiguously along the margin, slightly rounded proximally: fixed chelal finger with 28 teeth; movable chelal finger with 35 teeth (nearly as large as teeth on fixed chelal finger); without accessory teeth (Fig.
<figureCitation id="BE0D5C510E9833064AB51F5912F91A2D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3H</figureCitation>
). Sense spots absent; femur without long tactile setae. Movable chelal finger slightly curved in lateral view (Figs
<figureCitation id="A39D8C438312A9DFB4147C30B3C42A6C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2C</figureCitation>
,
<figureCitation id="529C5A045B0220570602DE27FB841AEA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3H</figureCitation>
).
<emphasis id="C9B0248F81FD422B766FED05F53E9131" bold="true" italics="true" pageId="0" pageNumber="295">Opisthosoma</emphasis>
: generally typical, all setae long and acuminate, pleural membrane longitudinally striate, without setae. Tergites I and XI undivided and other tergites incompletely divided, tergal chaetotaxy I-XI: 9: 9: 4-4: 7-7: 9-8: 7-8: 7-8: 8-7: 8-6: 7-7+ (2T): 13 + (4T), arranged in two rows on tergites IV-XI. Sternites XI undivided and other sternites divided, sternal chaetotaxy IV-XI: 6-6: 8-7:8-8: 8-8: 6-7: 7-7: 7-7 (4T): 13 + (4T). Anus (tergite XII and sternite XII) without raised rim. Anterior genital operculum with seven setae on each side, posterior margin with four setae on each side and with one or two lyrifissures. Male genitalia (Figs
<figureCitation id="0B34938657CFC8C9EB64BB9CBD72696A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2E</figureCitation>
,
<figureCitation id="6EF8F4306CEBE9B9FFF115DAD9B90BBF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3J, L</figureCitation>
): the distal part (l) of lateral apodemes with a distinctive inner ridge curved into semicircle; the hooked branch (br) bowed distally and terminated in a plate-like tip; the proximal part with a distinct dark sclerotized bar (c), shorter and incurved; the longitudinal fold of medial diverticula (d) with a projection midway along its length, forming a lobe bulge; the ejaculatory canal atrium (e) not well-developed, curved distally; the lateral rods (f) long and diverging proximally; the tip of dorsal apodeme (g) completely joined; the ventral diverticulum (h) bilobed; genital atrium without genital setae.
<emphasis id="25ACFFA048C4F1B68DE73AAC7B7D6772" bold="true" italics="true" pageId="0" pageNumber="295">Legs</emphasis>
(Figs
<figureCitation id="A056D64E224A13DB6C5037500CF88F88" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2G-I</figureCitation>
,
<figureCitation id="A6FB7F697081CB144B4B50A5DAD2AD71" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3G, K</figureCitation>
): typical, fairly smooth, slightly stout; junction between femora and patellae I and II oblique. Femoropatella of leg IV 3.13
<normalizedToken id="65E487E1AD0DB0966E51494126809659" originalValue="×">x</normalizedToken>
longer than deep; tibia 3.93
<normalizedToken id="63D4F1D876BDA749994C77D84F4BBFA0" originalValue="×">x</normalizedToken>
longer than deep; with basal tactile setae on tarsal segment: tarsus 4.50
<normalizedToken id="14FA2767A73FAF3CEEFC2D6D5F4A58FA" originalValue="×">x</normalizedToken>
longer than deep (TS = 0.09-0.11); subterminal tarsal setae arcuate and acute. Arolium slightly shorter than claws, not divided; claws smooth.
</paragraph>
<caption id="DF7CE4E67815C547E17A3B252E0EF836" doi="10.3897/zookeys.1168.100798.figure1" httpUri="https://binary.pensoft.net/fig/872299" pageId="0" pageNumber="295" start="Figure 1" startId="F1">
<paragraph id="582929BCE0EA0B3B45279259A0565A1B" pageId="0" pageNumber="295">
<emphasis id="58693AC1E0524475B4BB15171F9DC2AA" bold="true" pageId="0" pageNumber="295">Figure 1.</emphasis>
<taxonomicName id="28AAA4076688EE9E2C744694048A0E3B" authorityName="Hou &amp; Zhao &amp; Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus huae" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="huae">
<emphasis id="93C230D4015E967857C1885C1CC5468C" italics="true" pageId="0" pageNumber="295">Catatemnus huae</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="B7651FB7B4D7CB6FE9983248CB38EEB9" bold="true" pageId="0" pageNumber="295">A</emphasis>
holotype male, habitus (minus left pedipalp, left legs I and IV, dorsal view)
<emphasis id="5827FE653C3D7D9D7BE3F121D75925CB" bold="true" pageId="0" pageNumber="295">B</emphasis>
paratype female, habitus (minus left pedipalp, left legs I and III, right leg IV, dorsal view). Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="53EA1F19A01EEA024EB6D65213E4B395" doi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295" start="Figure 2" startId="F2">
<paragraph id="DE38F729240C3B948854A30D993D819B" pageId="0" pageNumber="295">
<emphasis id="432E94CA587E005BD100F9178423F3C0" bold="true" pageId="0" pageNumber="295">Figure 2.</emphasis>
<taxonomicName id="F51F466D7FC5F03BB6FD30B6A28497D9" authorityName="Hou &amp; Zhao &amp; Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus huae" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="huae">
<emphasis id="3B798601D7EE966AA2F0C0276D6689E6" italics="true" pageId="0" pageNumber="295">Catatemnus huae</emphasis>
</taxonomicName>
sp. nov., holotype male (
<emphasis id="76D280DB188DE346090EA05DF0985A02" bold="true" pageId="0" pageNumber="295">A-E, G-I</emphasis>
), paratype female (
<emphasis id="DC1756D2FE25BC99782827A999F757BE" bold="true" pageId="0" pageNumber="295">F</emphasis>
)
<emphasis id="AEE5548EDFACD1CCCEE5670867048AAA" bold="true" pageId="0" pageNumber="295">A</emphasis>
carapace (dorsal view)
<emphasis id="F072824095F620D95FAE1975739F5AD8" bold="true" pageId="0" pageNumber="295">B</emphasis>
left chela (lateral view)
<emphasis id="5A2BF64EA4D67F187F35C42FA8FC0029" bold="true" pageId="0" pageNumber="295">C</emphasis>
left chelal fingers (lateral view)
<emphasis id="E70B99D7B6E87FE14E5FECB890AE8FE9" bold="true" pageId="0" pageNumber="295">D</emphasis>
left pedipalp (dorsal view)
<emphasis id="DCB1A066FFC05A3205E75B9443E7ECEC" bold="true" pageId="0" pageNumber="295">E</emphasis>
male genital area (ventral view)
<emphasis id="E63042D3D4BD458242BBB27F496839C6" bold="true" pageId="0" pageNumber="295">F</emphasis>
female genital area (ventral view)
<emphasis id="5EB9849ED312EF0BE27B07EF5EA8EA35" bold="true" pageId="0" pageNumber="295">G</emphasis>
tarsus of left leg IV (lateral view)
<emphasis id="BEA4AA4C77B0BA32D792CD275B2875DF" bold="true" pageId="0" pageNumber="295">H</emphasis>
left leg I (lateral view)
<emphasis id="397B8E9C242477E5FCCFFF26970CFBEE" bold="true" pageId="0" pageNumber="295">I</emphasis>
left leg IV (lateral view). Scale bars: 0.20 mm (
<emphasis id="FFFD338A29A82202588A8637805EDC3B" bold="true" pageId="0" pageNumber="295">B, D, H, I</emphasis>
); 0.10 mm (
<emphasis id="455BC37D99DDF1256FA1B2F9549B6809" bold="true" pageId="0" pageNumber="295">A, C, E-G</emphasis>
).
</paragraph>
</caption>
<caption id="72B3F39FB3B5EC6341DD7320B1CC6E7E" doi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295" start="Figure 3" startId="F3">
<paragraph id="1AFFB61D4B07D47840E9FB1B46725184" pageId="0" pageNumber="295">
<emphasis id="1B01A09A00242A2B3D9F05E8628DB82A" bold="true" pageId="0" pageNumber="295">Figure 3.</emphasis>
<taxonomicName id="BF58951AA8615D430B99070006426D98" authorityName="Hou &amp; Zhao &amp; Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus huae" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="huae">
<emphasis id="5610B39C09E19FE247A715DF5CA02ECD" italics="true" pageId="0" pageNumber="295">Catatemnus huae</emphasis>
</taxonomicName>
sp. nov., holotype male (
<emphasis id="4065BB2DD09F6AD0A55D859584320125" bold="true" pageId="0" pageNumber="295">A-C, E-H, J-L</emphasis>
), paratype female (
<emphasis id="36C20CC5CD21BAA2A2B455F65C21CD2C" bold="true" pageId="0" pageNumber="295">D, I</emphasis>
)
<emphasis id="D9388AF8C2A38182F1F70B55FF7603FB" bold="true" pageId="0" pageNumber="295">A</emphasis>
carapace (dorsal view)
<emphasis id="A54161B77A8AF0D3C6260CA1AC036D0D" bold="true" pageId="0" pageNumber="295">B</emphasis>
left chelicera (dorsal view)
<emphasis id="1CECAF74442C642543F63396DA59FC6C" bold="true" pageId="0" pageNumber="295">C</emphasis>
male galea
<emphasis id="D1D6C118BE94D0FC41F91539F7842622" bold="true" pageId="0" pageNumber="295">D</emphasis>
female galea
<emphasis id="0E436375115D5537274F95D6A62CA42D" bold="true" pageId="0" pageNumber="295">E</emphasis>
rallum
<emphasis id="135BDB2B51D4C5181D0B203C968147A0" bold="true" pageId="0" pageNumber="295">F</emphasis>
left pedipalp (dorsal view)
<emphasis id="0FAA6B4B77F6635B769A6315CE930AEE" bold="true" pageId="0" pageNumber="295">G</emphasis>
left leg I (lateral view)
<emphasis id="6566524F74FC1EEAADAA2A8B0EF7A0A1" bold="true" pageId="0" pageNumber="295">H</emphasis>
left chelal fingers (lateral view), with details of trichobothrial pattern
<emphasis id="941DCC24B37ECB128E83C79BD9DA26ED" bold="true" pageId="0" pageNumber="295">I</emphasis>
female genital area (ventral view)
<emphasis id="1DCDF7598773C84339C679C984EB7DE0" bold="true" pageId="0" pageNumber="295">J</emphasis>
male genital area (ventral view)
<emphasis id="0B8A72B2DA857CCB70F67B21A8AFFB24" bold="true" pageId="0" pageNumber="295">K</emphasis>
left leg IV (lateral view)
<emphasis id="4A2E27B6AF8BCE143B5CA1A13236DB93" bold="true" pageId="0" pageNumber="295">L</emphasis>
male genital organ. Scale bars: 0.20 mm (
<emphasis id="F3BEFDF8EE148445510D0483563EE00D" bold="true" pageId="0" pageNumber="295">F, G, K</emphasis>
); 0.10 mm (
<emphasis id="BE7B222F45632363E231B70DA735F855" bold="true" pageId="0" pageNumber="295">A-E, H-J, L</emphasis>
). Abbreviations:
<emphasis id="36D02354DC3C4E4640D075D299B3BBEC" bold="true" pageId="0" pageNumber="295">b</emphasis>
= basal;
<emphasis id="1D89676FF1663905ABC61688D558B61F" bold="true" pageId="0" pageNumber="295">sb</emphasis>
= sub-basal;
<emphasis id="401260B786C909A7F9A6A5E9CA1F37C2" bold="true" pageId="0" pageNumber="295">st</emphasis>
= sub-terminal;
<emphasis id="64A17B35AF76473A2A98623A249D7660" bold="true" pageId="0" pageNumber="295">t</emphasis>
= terminal;
<emphasis id="7089F2141F1CA0AFF8992C70E5BF3DC2" bold="true" pageId="0" pageNumber="295">ib</emphasis>
= interior basal;
<emphasis id="C31D9B3D27AD660C559B9D1314EA5E1F" bold="true" pageId="0" pageNumber="295">isb</emphasis>
= interior sub-basal;
<emphasis id="DBE53F02682607CD675AA5513DAC4E61" bold="true" pageId="0" pageNumber="295">ist</emphasis>
= interior sub-terminal;
<emphasis id="C70040DD0DD4D197F90587DE2C4B2F36" bold="true" pageId="0" pageNumber="295">it</emphasis>
= interior terminal;
<emphasis id="76FF3595AF87DE968B0000A3A6EA78FC" bold="true" pageId="0" pageNumber="295">eb</emphasis>
= exterior basal;
<emphasis id="279582517873F7112152A545331523A0" bold="true" pageId="0" pageNumber="295">esb</emphasis>
= exterior sub-basal;
<emphasis id="20DB47628ED4A026B19BE40D51DE0510" bold="true" pageId="0" pageNumber="295">est</emphasis>
= exterior sub-terminal;
<emphasis id="B715A8C7608127B2999589CAFEEF77D9" bold="true" pageId="0" pageNumber="295">et</emphasis>
= exterior terminal;
<emphasis id="2F2CFDC5C5FEDFCED77F6296A9291903" bold="true" pageId="0" pageNumber="295">a</emphasis>
= lateral apodeme;
<emphasis id="444C900FBAC80E6FD6015CB64DBD9157" bold="true" pageId="0" pageNumber="295">br</emphasis>
= hooked branch;
<emphasis id="675E5A4EBFE04F147D558FEED8DFEB89" bold="true" pageId="0" pageNumber="295">c</emphasis>
= sclerotized bar;
<emphasis id="7547A50EDBDA4584E3C8AEFB716CEE10" bold="true" pageId="0" pageNumber="295">d</emphasis>
= longitudinal fold of medial diverticulum;
<emphasis id="8D27786891EB8A0D67F3B427DF2F4C8F" bold="true" pageId="0" pageNumber="295">e</emphasis>
= ejaculatory canal atrium;
<emphasis id="E40397DB17BAA8D437290D06CDA74A1C" bold="true" pageId="0" pageNumber="295">f</emphasis>
= lateral rods;
<emphasis id="EF92253787FDE70642F924E7476492CE" bold="true" pageId="0" pageNumber="295">g</emphasis>
= dorsal apodeme;
<emphasis id="061D1693CE1B422070D5F089686FA863" bold="true" pageId="0" pageNumber="295">h</emphasis>
= ventral diverticulum;
<emphasis id="9D9B62A08D59C6835F2C090AF04F3ADE" bold="true" pageId="0" pageNumber="295">l</emphasis>
= lateral lip of lateral apodeme.
</paragraph>
</caption>
<paragraph id="A89025E25452EC11738AF73C0E962B67" pageId="0" pageNumber="295">
<emphasis id="2B450A9303C2209305988BBC7F15C4E8" bold="true" pageId="0" pageNumber="295">Adult Female</emphasis>
(Figs
<figureCitation id="086BEC426DDD091162FB838ACD1BEB17" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Catatemnus huae sp. nov. A holotype male, habitus (minus left pedipalp, left legs I and IV, dorsal view) B paratype female, habitus (minus left pedipalp, left legs I and III, right leg IV, dorsal view). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1168.100798.figure1" httpUri="https://binary.pensoft.net/fig/872299" pageId="0" pageNumber="295">1B</figureCitation>
,
<figureCitation id="30380779D9BC4C73187C43FD5F5E9DEE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2F</figureCitation>
,
<figureCitation id="D19404E6449BEAFE64F62C3AC07234C6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3D, I</figureCitation>
): mostly same as the male, but a little larger in body size; anterior half of carapace yellowish brown, paler in posterior half; pedipalps reddish brown; tergites yellowish brown, paler in legs; remainder yellow.
<emphasis id="E8E4D6E5F0F13F90FAEDD5A971ED219C" bold="true" italics="true" pageId="0" pageNumber="295">Carapace</emphasis>
: anterior margin with two setae, posterior margin with eight setae, 46 in total.
<emphasis id="A3244B7025B5C0CEF84D10E7CA9601BA" bold="true" italics="true" pageId="0" pageNumber="295">Chelicera</emphasis>
: hand with four setae, bs and es dentate apically, is and ls long and acute; fixed finger with six large retrorse teeth and three small apical teeth, movable finger with two long broadly rounded subapical lobes; galea with six branchlets (Fig.
<figureCitation id="DF69C2CA88B64D30A475150F1E9DA029" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3D</figureCitation>
); serrula exterior with 21 lamellae; rallum with four blades.
<emphasis id="6C81FCDF503B957F4F9F04CDA4F7E032" bold="true" italics="true" pageId="0" pageNumber="295">Pedipalp</emphasis>
: stout, the retrolateral surface of trochanter, the prolateral surface of femur, patella and base of chelal fingers granular; trochanter with two distinct conical dorsal tubercles; trochanter 1.88
<normalizedToken id="E687F2A59916AAFFF50E9CBF23D51FCE" originalValue="×">x</normalizedToken>
, femur 2.52
<normalizedToken id="4AC74EF40AC519CF372E61491D7B139B" originalValue="×">x</normalizedToken>
, patella 2.20
<normalizedToken id="2D8AACDF933BF0E1828ADFB3B4A32B0C" originalValue="×">x</normalizedToken>
, chela (with pedicel) 2.93
<normalizedToken id="17B7222BA7F8AB0FDD35FFB71E7ABC87" originalValue="×">x</normalizedToken>
, chela (without pedicel) 2.74
<normalizedToken id="957DB990229B168851F0E1BEBA13C195" originalValue="×">x</normalizedToken>
, hand (without pedicel) 1.58
<normalizedToken id="AED53CE069F40161693E8D8107CE097B" originalValue="×">x</normalizedToken>
longer than broad, movable chelal finger 0.67
<normalizedToken id="C644A3BD20C56710B0C81A7ABF76C4D9" originalValue="×">x</normalizedToken>
(0.75
<normalizedToken id="FE15C81282C12A4302607DEDF3B59133" originalValue="×">x</normalizedToken>
) longer than hand with pedicel (without pedicel).
<emphasis id="53987786AC7DD26A5B74C6BB31043851" bold="true" italics="true" pageId="0" pageNumber="295">Opisthosoma</emphasis>
: tergites I, II, and XI undivided, tergites III and IV incompletely divided and other tergites completely divided; tergal chaetotaxy (I-XI): 8: 11: 6-6: 8-7: 7-8: 9-8: 9-9: 8-8: 8-8: 14+ (2T): 10+ (4T); sternites XI undivided and other sternites divided, sternal chaetotaxy (IV-XI): 6-6: 8-8:8-9: 7-5: 8-6: 8-8: 8-9 (4T): 9 + (4T); genital structure (Figs
<figureCitation id="A5456B8A8A7AC2862BA5B8803854A290" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Catatemnus huae sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chela (lateral view) C left chelal fingers (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (B, D, H, I); 0.10 mm (A, C, E-G)." figureDoi="10.3897/zookeys.1168.100798.figure2" httpUri="https://binary.pensoft.net/fig/872300" pageId="0" pageNumber="295">2F</figureCitation>
,
<figureCitation id="1675E22E5A3CBD8D24F9E628C5A413B9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Catatemnus huae sp. nov., holotype male (A-C, E-H, J-L), paratype female (D, I) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F left pedipalp (dorsal view) G left leg I (lateral view) H left chelal fingers (lateral view), with details of trichobothrial pattern I female genital area (ventral view) J male genital area (ventral view) K left leg IV (lateral view) L male genital organ. Scale bars: 0.20 mm (F, G, K); 0.10 mm (A-E, H-J, L). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure3" httpUri="https://binary.pensoft.net/fig/872301" pageId="0" pageNumber="295">3I</figureCitation>
) simple, spermathecae provided with separated median cribriform plates.
</paragraph>
<paragraph id="CB06039E6040CA4C17E0CEEC7C68C55B" pageId="0" pageNumber="295">
<emphasis id="CC8D1F6AA258A3874237A26F3917E278" bold="true" pageId="0" pageNumber="295">Dimensions</emphasis>
(length/breadth or, in the case of the legs, length/depth in mm). Male (female in parentheses): body length 3.19 (3.21). Carapace 0.87/0.66 (0.87/0.69). Pedipalp: trochanter 0.48/0.26 (0.49/0.26), femur 0.72/0.29 (0.78/0.31), patella 0.77/0.36 (0.77/0.35), chela (with pedicel) 1.28/0.42 (1.35/0.46), chela (without pedicel) 1.18 (1.26), hand (with pedicel) 0.75 (0.82), hand (without pedicel) 0.65 (0.73), movable finger length 0.53 (0.55). Leg I: trochanter 0.17/0.14 (0.16/0.15), femur 0.28/0.18 (0.27/0.19), patella 0.40/0.16 (0.40/0.16), tibia 0.40/0.12 (0.42/0.12), tarsus 0.38/0.08 (0.41/0.09). Leg IV: trochanter 0.30/0.16 (0.30/0.17), femoropatella 0.75/0.24 (0.79/0.25), tibia 0.59/0.15 (0.62/0.16), tarsus 0.45/0.10 (0.48/0.11).
</paragraph>
</subSubSection>
<subSubSection id="30DC5F725D886EA9B577875FB1F81A31" pageId="0" pageNumber="295" type="remarks">
<paragraph id="90771B0AF3ABF6A56B3A15B2B7D432D3" pageId="0" pageNumber="295">Remarks.</paragraph>
<paragraph id="B82B1843450C57592EAE03102357D22C" pageId="0" pageNumber="295">
Prior to this study, 13 species of
<taxonomicName id="E19A4A227E79C7043D48BE42F1BA39A5" authorityName="Beier" authorityYear="1932" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="F571662FA9F50A53C3C74ED4A13E5F9C" italics="true" pageId="0" pageNumber="295">Catatemnus</emphasis>
</taxonomicName>
have been recorded around the world, of which five come from Asia, but none have been recorded in China. The medial furrow on the carapace of
<taxonomicName id="7AAA97AA9B2F23225D5E29D81FB8B7CB" authorityName="Hou &amp; Zhao &amp; Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus huae" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="huae">
<emphasis id="7700B6EBD20CA63A2154C871766387EE" italics="true" pageId="0" pageNumber="295">Catatemnus huae</emphasis>
</taxonomicName>
sp. nov. was a little faint (but still visible) probably due to the pale body color.
</paragraph>
<paragraph id="7E75F35E9C7900B8C04127AA8E0CC7C1" pageId="0" pageNumber="295">
<taxonomicName id="5F6E8A4AF9F7F900C7D426C6E8CD3932" authorityName="Hou &amp; Zhao &amp; Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus huae" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="huae">
<emphasis id="142DAE875D269873F642F483E77C8084" italics="true" pageId="0" pageNumber="295">Catatemnus huae</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName id="CD5B3EF8ABD1AF1877055D31498A8A45" class="Arachnida" kingdom="Animalia" lsidName="C. birmanicus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="9F2B6EC0ECF26FBD9CE6E5F86F857A1A" italics="true" pageId="0" pageNumber="295">C. birmanicus</emphasis>
</taxonomicName>
(Thorell, 1889) by the stouter pedipalps (e.g., palpal femur (♀) 2.52
<normalizedToken id="587120DD6687B7945447FDC7A3B550B1" originalValue="×">x</normalizedToken>
vs 2.36
<normalizedToken id="231B0B23319E270BBC6F42C39624A785" originalValue="×">x</normalizedToken>
longer than broad, length 0.78 mm vs 0.92 mm) and the traits of pedipalps (distinctly granular vs minutely granular); from
<taxonomicName id="5F80246BF02A420C2CC4260635802412" class="Arachnida" kingdom="Animalia" lsidName="C. concavus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="concavus">
<emphasis id="510282B3B6E45688B6FB4F27DE2D546C" italics="true" pageId="0" pageNumber="295">C. concavus</emphasis>
</taxonomicName>
(With, 1906) by the stouter pedipalps (♀) (e.g., palpal femur 2.52
<normalizedToken id="E3AA9801E9AA9378F8E22765DAE6EDAE" originalValue="×">x</normalizedToken>
vs 2.29
<normalizedToken id="E589EE3BF47068023F93ACCDDB29C11E" originalValue="×">x</normalizedToken>
longer than broad, length 0.78 mm vs 0.90 mm), the trait of galea (galea well-developed and with six branchlets vs galea undeveloped and partly lost) and the color of the tergites (pale yellow vs darker brown); from
<taxonomicName id="1C909B0A2B45C851543F9BB3CD0EC1DF" class="Arachnida" kingdom="Animalia" lsidName="C. monitor" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="monitor">
<emphasis id="4B30E521B8C78487A2815B71CD9558A3" italics="true" pageId="0" pageNumber="295">C. monitor</emphasis>
</taxonomicName>
(With, 1906) by stouter pedipalps (e.g., palpal femur (♂) 2.48
<normalizedToken id="BF1D0CBB0653558D5029F18A71B82972" originalValue="×">x</normalizedToken>
vs 2.25
<normalizedToken id="9FD122E4355B7ACAC2A4A1C01FAB6BC0" originalValue="×">x</normalizedToken>
, (♀) 2.52
<normalizedToken id="FCF0C2AD25090B0555DE1B7013D34225" originalValue="×">x</normalizedToken>
vs 2.27
<normalizedToken id="3F89E7D8C572692091466CA465972823" originalValue="×">x</normalizedToken>
longer than broad, length (♂) 0.72 mm vs 1.01 mm, (♀) 0.78 mm vs 0.95 mm), the trait of eyespots (with two faint eyespots vs no eyespots visible) and the color of the dorsal surface of the body (pale yellow vs dark brown); from
<taxonomicName id="1BB083749B5ACDB3D432BBB6E9FD5D6C" class="Arachnida" kingdom="Animalia" lsidName="C. nicobarensis" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="10441841ED23B2B047D96E54E90877EA" italics="true" pageId="0" pageNumber="295">C. nicobarensis</emphasis>
</taxonomicName>
(With, 1906) by the stouter pedipalps (♂) (e.g., palpal femur 2.48
<normalizedToken id="69BF2ED32E633D3A655C64CF9609D1F7" originalValue="×">x</normalizedToken>
vs 2.14
<normalizedToken id="46499D01B59B84F096CE2DA9246DB265" originalValue="×">x</normalizedToken>
longer than broad, length 0.72 mm vs 0.97 mm) and the traits of the palpal trochanter (anterior surface granular vs smooth or almost smooth); from
<taxonomicName id="66AF79D2131988ABA235CC88C01E1FE9" class="Arachnida" kingdom="Animalia" lsidName="C. thorelli" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="thorelli">
<emphasis id="59CFA9C27F80A75C1CDA9E17E3543F29" italics="true" pageId="0" pageNumber="295">C. thorelli</emphasis>
</taxonomicName>
(Balzan, 1892) by the trait of eyespots (with two faint eyespots vs with two distinct eyespots) and the traits of pedipalps (surface of all segments (except chelal fingers) granular vs surface of palps smooth) (
<bibRefCitation id="CF5883DC49A5D75B18DDCB7046392FBE" DOI="https://doi.org/10.5962/bhl.title.10400" author="Thorell, T" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" pagination="521 - 729" refId="B9" refString="Thorell, T, 1889. Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885-1887. Annali del Museo Civico di Storia Naturale di Genova 7 (2): 521 - 729, DOI: https://doi.org/10.5962/bhl.title.10400" title="Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885 - 1887." url="https://doi.org/10.5962/bhl.title.10400" volume="7" year="1889">Thorell 1889</bibRefCitation>
;
<bibRefCitation id="DD357E1C1D2AC4B96718008576EC57F5" author="Balzan, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" pagination="497 - 512" refId="B1" refString="Balzan, L, 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societe Entomologique de France 60: 497 - 512" title="Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones)." volume="60" year="1891">Balzan 1891</bibRefCitation>
;
<bibRefCitation id="78DF6660E4B31DC10074AC97440017FD" author="With, CJ" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" refId="B10" refString="With, CJ, 1906. The Danish expedition to Siam 1899-1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7)3: 1-214." title="The Danish expedition to Siam 1899 - 1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7) 3: 1 - 214." year="1906">With 1906</bibRefCitation>
;
<bibRefCitation id="CC5D8212F7317B51BE72A19BFC575318" DOI="https://doi.org/10.1515/9783111385402" author="Beier, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" refId="B2" refString="Beier, M, 1932. Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1-294. https://doi.org/10.1515/9783111385402" title="Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1 - 294." url="https://doi.org/10.1515/9783111385402" year="1932">Beier 1932</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1E4F8B5519DF9C0F19D4D6329ABEFD36" pageId="0" pageNumber="295" type="distribution">
<paragraph id="98F8896308B101BFB429D5D73C33F496" pageId="0" pageNumber="295">Distribution.</paragraph>
<paragraph id="05B9F82A1E480699E7820E9FE1535987" pageId="0" pageNumber="295">Known only from the type locality.</paragraph>
</subSubSection>
<subSubSection id="6A89C49DF32994256EE610B4CCBE4D80" pageId="0" pageNumber="295" type="etymology">
<paragraph id="F2FB0A8F423C7E883A5C6EF56A9E300A" pageId="0" pageNumber="295">Etymology.</paragraph>
<paragraph id="52E7901F5ECF485B4991B35F4FAD0531" pageId="0" pageNumber="295">The new species is named after Junfang Hu, who collected the specimen for this study.</paragraph>
</subSubSection>
</treatment>
</document>