614 lines
78 KiB
XML
614 lines
78 KiB
XML
<document id="7C22790F7FF7AD8A9C976CE412B9F174" ID-DOI="10.3897/zookeys.1224.140549" ID-ZooBank="8F8AAC17-7284-46F8-A2B5-1BA03FDB54E1" ID-publisher-id="140549" URI-arpha="1734317E-C27C-553F-9F2E-405A35777A94" URI-zoobank="http://zoobank.org/8F8AAC17-7284-46F8-A2B5-1BA03FDB54E1" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1738088624095" checkinUser="pensoft" docAuthor="Jeratthitikul, Ekgachai, Sutcharit, Chirasak & Prasankok, Pongpun" docDate="2025" docId="9BDBEDA98C975EA793681649DEF9E8B2" docLanguage="en" docName="ZooKeys 1224: 187-209" docOrigin="ZooKeys 1224" docSource="https://zookeys.pensoft.net/article/140549/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Songkhlanaia moreleti Jeratthitikul & Sutcharit & Prasankok 2025, comb. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="209" masterDocId="1734317EC27C553F9F2E405A35777A94" masterDocTitle="An integrative approach to a revision of the freshwater mussel genus Songkhlanaia (Bivalvia, Unionidae), with the description of a new species" masterLastPageNumber="209" masterPageNumber="187" pageNumber="187" updateTime="1738094874349" updateUser="admin">
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<mods:titleInfo id="8FBC2BFC554D1A8D9A27F20109EC842C">
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<mods:title id="CB083E35398E97141DF37884773EEF55">An integrative approach to a revision of the freshwater mussel genus Songkhlanaia (Bivalvia, Unionidae), with the description of a new species</mods:title>
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<mods:name id="C78B0C43F6C7FAD9FEDA6C3B9EE77703" type="personal">
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<mods:namePart id="542EFFA17E7B7CBFCD0597EAFF5A0931">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:nameIdentifier id="EF6FE411B9E1617E62F9C1050D108AD3" type="ORCID">0000-0002-3477-9548</mods:nameIdentifier>
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<mods:affiliation id="576A4F8A9F1DA95BA639D1A31779F5DF">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="C39F361EC98272098BB3ECBBEF41D592">Sutcharit, Chirasak</mods:namePart>
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<mods:nameIdentifier id="9A8BD3098FDF4FEF5C214CD999DABC84" type="ORCID">0000-0001-7670-9540</mods:nameIdentifier>
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<mods:affiliation id="5712FA5D53D4DE31FD49E1B4984FB3A8">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="0D7480D778F43FD9D7FBF23604A94F12">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="97992CD8638971A0E39D3CBC87334B6A">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:typeOfResource id="B95645828390CFC3643745C58D9C300D">text</mods:typeOfResource>
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<mods:title id="9970B4018BAFB8C4F9D7B01B8CB711CD">ZooKeys</mods:title>
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<mods:part id="3E03EBF22E4350F145F1E360F7E8BE07">
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<mods:date id="46A15F6428C87096CC7E8A2F0EE647F6">2025</mods:date>
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<mods:number id="41166423E98369FFBBED54B23A29FF2C">2025-01-28</mods:number>
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<mods:number id="595EC6708027C624E2E7BB5FD8E6004E">1224</mods:number>
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<mods:start id="599AB9858B5C8DB7FF67F2CA64985795">187</mods:start>
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<mods:identifier id="79FF66F3506D40BBF3442BFDD06CDACB" type="DOI">10.3897/zookeys.1224.140549</mods:identifier>
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<mods:identifier id="60F000AE3CD39DE2696119C51B7E776F" type="ZooBank">8F8AAC17-7284-46F8-A2B5-1BA03FDB54E1</mods:identifier>
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<treatment id="9BDBEDA98C975EA793681649DEF9E8B2" ID-arpha="9BDBEDA9-8C97-5EA7-9368-1649DEF9E8B2" LSID="urn:lsid:plazi:treatment:9BDBEDA98C975EA793681649DEF9E8B2" httpUri="http://treatment.plazi.org/id/9BDBEDA98C975EA793681649DEF9E8B2" scope_class="Bivalvia" scope_family="Unionidae" scope_subFamily="Gonideinae" scope_subTribe="Pseudodontina" scope_tribe="Pseudodontini">
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<subSubSection id="7B475E0DD8BF07A77F8037C9A805B3A8" type="nomenclature">
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<paragraph id="6F4F6B31FF70498F9E457904DC17654A">
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<taxonomicName id="A074FE146615E2F0628AE8838BE424BE" authority="(Crosse & Fischer, 1876)" authorityName="Jeratthitikul & Sutcharit & Prasankok" authorityYear="2025" baseAuthorityName="Crosse & Fischer" baseAuthorityYear="1876" class="Bivalvia" family="Unionidae" genus="Songkhlanaia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Unionoida" phylum="Mollusca" rank="species" species="moreleti" status="comb. nov.">
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||
<emphasis id="0622FC86A7660B6DB7BE78FF44D23D1F" italics="true">Songkhlanaia moreleti</emphasis>
|
||
(
|
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<bibRefCitation id="214A844DF3995888A60554830BBA418F" author="Crosse H & Fischer P" journalOrPublisher="Journal de Conchyliologie" pagination="313-342" refId="B11" refString="Crosse H, Fischer P (1876) Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873. Journal de Conchyliologie 24: 313 - 342." title="Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873." volume="24" year="1876">Crosse & Fischer, 1876</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
<taxonomicNameLabel id="1EA2325B6C43E052729A415B3A7F41E4" rank="species">comb. nov.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F5165FA76B3DA9D6365D608B851CF5A9" type="description">
|
||
<paragraph id="6A37205C052ED3DCC47FE2EAC9DE6D93">
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||
<figureCitation id="3B1F260681161E7532B0C84D84C44E2E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shells of Songkhlanaia species A, B topotype of S. tamodienica from Tamot Stream, Songkhla Lake Basin, Thailand A adult specimen MUMNH - UNI 2956 and B young specimen MUMNH - UNI 2971 C – F S. moreleti C syntype MNHN - IM- 2000–34623 from Mekong Basin, Cambodia, with original labels D syntype of Pseudodon thomsoni Morlet, 1884, MNHN - IM- 2000-1800 from Mekong Basin, Cambodia, with original labels E adult specimen MUMNH - UNI 0301, and F young specimen MUMNH - UNI 3428 from Phrom Hot Stream, Tonle Sap Basin, Thailand. Photographs C, D M. Caballer (2019, MNHN Project: RECOLNAT No. ANR- 11 - INBS- 0004)." figureDoi="10.3897/zookeys.1224.140549.figure2" httpUri="https://binary.pensoft.net/fig/1236494">Figs 2 C – F</figureCitation>
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,
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<figureCitation id="5BC257E056D205FDFBFCDCB7FE1D7DC0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudocardinal teeth, left valve on the left-hand side, and right valve on the right-hand side A S. tamodienica, topotype MUMNH - UNI 2956 from Tamot Stream, Songkhla Lake Basin, Thailand B S. moreleti, specimen MUMNH - UNI 0301 from Phrom Hot Stream, Tonle Sap Basin, Thailand C S. songkhramensis sp. nov., holotype MUMNH - UNI 3024 from Prahang River, Songkhram Basin, Thailand. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1224.140549.figure5" httpUri="https://binary.pensoft.net/fig/1236497">5 B</figureCitation>
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||
,
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<tableCitation id="7C3225A56426CB50F6C4C91B0DCD5591" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of conchological characteristics among Songkhlanaia species." httpUri="http://table.plazi.org/id/4CC699547008E3F0220C9F7A03AAAE7A" tableUuid="4CC699547008E3F0220C9F7A03AAAE7A">Table 1</tableCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A35CA2E21AAFC79E03CEF16ED218A7BE" type="reference_group">
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<paragraph id="C3E4C513BFC8D7D52B1DE3B67BE4EC52">
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<treatmentCitationGroup id="E64958373C77BC3FFF64EC08CCB02317">
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||
<treatmentCitation id="5D65BC9F99B1B7CF0EF64C08B45C7B64" author="Deshayes G-P & Jullien J" page="120" year="1876">
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||
<taxonomicName id="A4F9CE317EF315592DE3D0D96176FE2B" authority="Deshayes & Jullien, 1876: 117 - 120" authorityName="Deshayes & Jullien" authorityPageNumber="117 - 120" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Monocondylea" isUncertain="true" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="tumida">
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||
<emphasis id="C67D2E8BE10E4EE02349C711272739C5" italics="true">Monocondylea</emphasis>
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||
<comment id="6CB8B3400351E9F0913DD51278CFD051">
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||
[sic]
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||
<emphasis id="21C6E19F5945E073EC27650C2273C293" italics="true">tumida</emphasis>
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||
<bibRefCitation id="081D9C704EBCE95B6DB319006A6502B9" DOI="10.5962/bhl.title.10273" author="Deshayes G-P & Jullien J" journalOrPublisher="Bulletin des Nouvelles Archives du Muséum d’Histoire Naturelle" pagination="115-163" refId="B13" refString="Deshayes G-P, Jullien J (1876) Memoire sur les mollusques nouveaux du Cambodge: Envoyes au museum par M. le docteur Jullien. Bulletin des Nouvelles Archives du Museum d'Histoire Naturelle 10: 115 - 163. https: // doi. org / 10.5962 / bhl. title. 10273" title="Mémoire sur les mollusques nouveaux du Cambodge: Envoyés au muséum par M. le docteur Jullien." volume="10" year="1876">Deshayes & Jullien, 1876: 117–120</bibRefCitation>
|
||
, pl. 5, figs 1–3 [in part; non
|
||
<taxonomicName id="3BA3991EEEA8BD012A45A13E0EBA66E5" ID-CoL="7S6Q6" authority="Morelet, 1866" authorityName="Morelet" authorityYear="1866" class="Bivalvia" family="Unionidae" genus="Monocondylus" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="tumidus">
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||
<emphasis id="D1F425B036CDAD40FF217D252E8B8E54" italics="true">Monocondylus tumidus</emphasis>
|
||
<bibRefCitation id="E11C6A540CA038C4E9402B2A0714D2E0" author="Morelet A" journalOrPublisher="Journal de Conchyliologie" pagination="62-64" refId="B45" refString="Morelet A (1866) Description d'especes appartenant a la faune malacologique de l'Indo-Chine. Journal de Conchyliologie 14: 62 - 64." title="Description d’espèces appartenant à la faune malacologique de l’Indo-Chine." volume="14" year="1866">Morelet, 1866</bibRefCitation>
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||
</taxonomicName>
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||
].
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="F29BA1DCD61F2028C95CBFA38366789D">
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||
<treatmentCitationGroup id="1C57C14808C6BCD8C70ED6EB92C493DB">
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||
<treatmentCitation id="24BA40535E1119CDD0B5E10BE439D4F8" author="Crosse H & Fischer P" page="330" year="1876">
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||
<taxonomicName id="8418E3FA354214CDCC5EDC57F60E1C5E" ID-CoL="7STY8" authority="Crosse & Fischer, 1876: 330" authorityName="Crosse & Fischer" authorityPageNumber="330" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
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||
<emphasis id="99FE4CFE0525E4C3FFDF3520F3BE903D" italics="true">Pseudodon moreleti</emphasis>
|
||
<comment id="B440E4EDBFFAD275F328D2F13E457BB6">
|
||
<bibRefCitation id="62A445EC9226A7D8E557610CEB2AC633" author="Crosse H & Fischer P" journalOrPublisher="Journal de Conchyliologie" pagination="313-342" refId="B11" refString="Crosse H, Fischer P (1876) Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873. Journal de Conchyliologie 24: 313 - 342." title="Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873." volume="24" year="1876">Crosse & Fischer, 1876: 330</bibRefCitation>
|
||
. Type Locality: “ les marécages qui avoisinent les rives du Mékong; dans un lac, à Préai-Bac, arroyo de Peam-Chelang; ďeau de la province de Compong-Soai ” [= Peam Chileang and Kampong Svay,
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||
<collectingCountry id="CA9AE054ABE0A7B88C53C4364304BA4A" name="Cambodia">Cambodia</collectingCountry>
|
||
].
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||
<treatmentCitation id="493B24F5A1BD46BC3E872771A08F029E" author="Fischer P" page="221" year="1891">
|
||
<bibRefCitation id="6E60F2E7C8F76D8430C3BF44B685E1D0" DOI="10.5962/bhl.title.14809" author="Fischer P" firstAuthor="Fischer" journalOrPublisher="Bulletin de la Société d’Histoire Naturelle d’Autun" page="221" pagination="87-276" refId="B15" refString="Fischer P (1891) Catalogue et distribution géographique des mollusques terrestres, fluviatiles & marins d’une partie de l’Indo-chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Bulletin de la Société d’Histoire Naturelle d’Autun 4: 87–276. https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution géographique des mollusques terrestres, fluviatiles & marins d’une partie de l’Indo-chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." volume="4" year="1891">Fischer 1891: 221</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
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||
<treatmentCitation id="74FE8CF957828B1DF550850CFFAA03BF" author="Fischer H & Dautzenberg P" page="436" year="1904">
|
||
<bibRefCitation id="3325E18E634C54329F8311478878216D" author="Fischer H & Dautzenberg P" firstAuthor="Fischer" journalOrPublisher="Mission Pavie, Etudes diverses" page="436" pagination="390-442" refId="B16" refString="Fischer H, Dautzenberg P (1904) Catalogue des mollusques terrestres et fluviatiles de l’Indo-Chine orientale cites jusqu’à ce jour. Mission Pavie, Etudes diverses 3: 390–442." title="Catalogue des mollusques terrestres et fluviatiles de l’Indo-Chine orientale cites jusqu’à ce jour." volume="3" year="1904">Fischer and Dautzenberg 1904: 436</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="56D619538D0A7AAFE0ED6697BF0D6690" author="Dautzenberg P & Fischer H" page="452" year="1905">
|
||
<bibRefCitation id="04FA499EAE9D4D7488925A4E6BEB70EF" author="Dautzenberg P & Fischer H" firstAuthor="Dautzenberg" journalOrPublisher="Journal de Conchyliologie" page="452" pagination="343-471" refId="B12" refString="Dautzenberg P, Fischer H (1905) Liste des mollusques récoltés par M. H. Mansuy en Indo-chine et au Yunnan et description d’especes nouvelles. Journal de Conchyliologie 53: 343–471." title="Liste des mollusques récoltés par M. H. Mansuy en Indo-chine et au Yunnan et description d’especes nouvelles." volume="53" year="1905">Dautzenberg and Fischer 1905: 452</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 453.
|
||
</comment>
|
||
</taxonomicName>
|
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</treatmentCitation>
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</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="781F015D0AEFBF3495090B4B63D54AE8">
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<treatmentCitationGroup id="65E83A67DDDD140C5445C7EFE87F00DD">
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<taxonomicName id="FB695E19A80F104C5E7BA4B4E56D5BB3" ID-CoL="7STY8" authorityName="Crosse & Fischer" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti" subGenus="Pseudodon">
|
||
<emphasis id="8AA94E023674F287C2018F6CB8A13B25" italics="true">Pseudodon</emphasis>
|
||
(
|
||
<emphasis id="99653199ADE166E9BC766B3F72D9BE5C" italics="true">Pseudodon</emphasis>
|
||
)
|
||
<emphasis id="AA05E3A36240DB2A4B513A26DDD2B4F0" italics="true">moreleti</emphasis>
|
||
</taxonomicName>
|
||
<comment id="5B5BCEB4DA4DCF944C076C32DC8CBA28">
|
||
—
|
||
<treatmentCitation id="BC756AEF4CCAFE8436AAE59CECB74227" author="Simpson CT" page="838" year="1900">
|
||
<bibRefCitation id="62CC5F13D1D05D53919035BF44014F4E" DOI="10.5479/si.00963801.22-1205.501" author="Simpson CT" firstAuthor="Simpson" journalOrPublisher="Proceedings of the United States National Museum" page="838" pagination="501-1044" refId="B58" refString="Simpson CT (1900) Synopsis of the naiades, or pearly fresh-water mussels. Proceedings of the United States National Museum 22: 501–1044. https://doi.org/10.5479/si.00963801.22-1205.501" title="Synopsis of the naiades, or pearly fresh-water mussels." volume="22" year="1900">Simpson 1900: 838</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="BA380B73C603C60AC8B24E8B56845EA9" author="Simpson CT" page="1094" year="1914">
|
||
<bibRefCitation id="3C897AEF3D46AA0BDE374E5FA40017BF" DOI="10.5962/bhl.title.10910" author="Simpson CT" firstAuthor="Simpson" journalOrPublisher="Bryant Walker, Detroit, Michigan" page="1094" refId="B59" refString="Simpson CT (1914) A descriptive catalogue of the naiades, or pearly fresh-water mussels. Parts I – III. Bryant Walker, Detroit, Michigan, 1540 pp. https://doi.org/10.5962/bhl.title.10910" title="A descriptive catalogue of the naiades, or pearly fresh-water mussels. Parts I - III." year="1914">Simpson 1914: 1094</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 1095.
|
||
<treatmentCitation id="78DD592C4A553C15ABEB4344016F44F1" author="Haas F" page="137" year="1924">
|
||
<bibRefCitation id="8108863794A578E7AEFF43FCCA6B19FF" author="Haas F" firstAuthor="Haas" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" page="137" pagination="129-203" refId="B24" refString="Haas F (1924) Beiträge zu einer Monographie der asiatischen Unioniden. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 38: 129–203." title="Beiträge zu einer Monographie der asiatischen Unioniden." volume="38" year="1924">Haas 1924: 137</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 138.
|
||
<treatmentCitation id="4D40EBC5D9B4259DC06D1E3DC6E39316" author="Haas F" page="130" year="1969">
|
||
<bibRefCitation id="F5EE5CAE4F29338CC7A85A507FA422B7" author="Haas F" editor="Martens R & Hennig W" firstAuthor="Haas" journalOrPublisher="Walter de Gruyter, Berlin" page="130" pagination="1-663" refId="B25" refString="Haas F (1969) Superfamilia Unionacea. In: Martens R, Hennig W (Eds) Das Tierrich, Lieferung Vol 88. Walter de Gruyter, Berlin, 1–663." title="Superfamilia Unionacea." volumeTitle="Das Tierrich, Lieferung Vol 88." year="1969">Haas 1969: 130</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="B332987F7EACA02B32D1EB110078259F">
|
||
<treatmentCitationGroup id="E161E3F17D191E1FE267304A3221D54D">
|
||
<taxonomicName id="8626F3FB0140FE2977F04C049D7466F9" ID-CoL="7T9WV" baseAuthorityName="Crosse & P. Fischer" baseAuthorityYear="1876" class="Bivalvia" family="Unionidae" genus="Sundadontina" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="53F9F3FDA1FE033583F90E63F0B2D8A2" italics="true">Sundadontina moreleti</emphasis>
|
||
</taxonomicName>
|
||
<comment id="FADEA8AFAA994E744B6B09760535942A">
|
||
—
|
||
<treatmentCitation id="A35D0CA8BB338DCF74129801B5C4E098" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Lopes-Lima M & Bogan AE & Lunn Z & Chan N & Win T & Aksenova OV & Tomilova AA & Tanmuangpak K & Tumpeesuwan S & Kondakov AV" page="10" year="2020">
|
||
<bibRefCitation id="E7CFD9EB1B4E7373FE96F69458F525D7" DOI="10.1038/s41598-020-63612-5" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Lopes-Lima M & Bogan AE & Lunn Z & Chan N & Win T & Aksenova OV & Tomilova AA & Tanmuangpak K & Tumpeesuwan S & Kondakov AV" etAl="et al." firstAuthor="Bolotov" page="10" refId="B6" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Lopes-Lima M, Bogan AE, Lunn Z, Chan N, Win T, Aksenova OV, Tomilova AA, Tanmuangpak K, Tumpeesuwan S, Kondakov AV (2020) New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10: 6616. https://doi.org/10.1038/s41598-020-63612-5" year="2020">Bolotov et al. 2020: 10</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 4 e.
|
||
<treatmentCitation id="2F8B8CD054D425992C657E9B1EFB5A54" author="Graf DL & Cummings KS" page="22" year="2021">
|
||
<bibRefCitation id="943EB123C5E567A23032123063F571D3" DOI="10.1093/mollus/eyaa034" author="Graf DL & Cummings KS" firstAuthor="Graf" page="22" refId="B20" refString="Graf DL, Cummings KS (2021) A ‘ big data’ approach to global freshwater mussel diversity (Bivalvia: Unionoida), with an updated checklist of genera and species. Journal of Molluscan Studies 87: eyaa 034. https://doi.org/10.1093/mollus/eyaa034" year="2021">Graf and Cummings 2021: 22</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="72912ED0A294A12BC0B98B82273EA6E3" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Kondakov AV & Lyubas AA & Soboleva AA & Chan N & Lunn Z & Win T & Inkhavilay K" page="13" year="2023">
|
||
<bibRefCitation id="A27290A2D88D0111096A728635E2AC37" DOI="10.3390/w15173117" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Kondakov AV & Lyubas AA & Soboleva AA & Chan N & Lunn Z & Win T & Inkhavilay K" etAl="et al." firstAuthor="Bolotov" page="13" refId="B8" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Kondakov AV, Lyubas AA, Soboleva AA, Chan N, Lunn Z, Win T, Inkhavilay K (2023) Integrative taxonomic reappraisal and evolutionary biogeography of the most diverse freshwater mussel clade from Southeast Asia (Pseudodontini). Water 15: 3117. https://doi.org/10.3390/w15173117" year="2023">Bolotov et al. 2023: 13</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="169195A94083E9E277B5A0DA2ACEE756">
|
||
<treatmentCitationGroup id="1C506FC197B704FD824C2B72E3ADFB0C">
|
||
<treatmentCitation id="F94E9E61802CD902FFBA01027EECA7B3" author="Morlet L" page="401" year="1884">
|
||
<taxonomicName id="3A433BD5D061FEDF962440B5E4D14C90" ID-CoL="7STYL" authority="Morlet, 1884: 401" authorityName="Morlet" authorityPageNumber="401" authorityYear="1884" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni">
|
||
<emphasis id="84C8A1D3889B60A7485CC460C18A0EFA" italics="true">Pseudodon thomsoni</emphasis>
|
||
<comment id="0AD7455392B1480D25028942D8496E7C">
|
||
<bibRefCitation id="4DB82DCC6475C77BAFBAA03847F049F7" author="Morlet L" journalOrPublisher="Journal de Conchyliologie" pagination="386-403" refId="B46" refString="Morlet L (1884) Description d'espece nouvelles de coquilles, recueillies par M. Pavie, au Cambodge. Journal de Conchyliologie 32: 386 - 403." title="Description d’espèce nouvelles de coquilles, recueillies par M. Pavie, au Cambodge." volume="32" year="1884">Morlet, 1884: 401</bibRefCitation>
|
||
, 402, pl. 13, figs 2, 2 a. Type Locality: “
|
||
<collectingCountry id="D101EF1888CE81F46C96B614DDFCCBD7" name="Cambodia">Cambodge</collectingCountry>
|
||
” [=
|
||
<collectingCountry id="09186043D298BF33259BC0CB45F6210C" name="Cambodia">Cambodia</collectingCountry>
|
||
].
|
||
<treatmentCitation id="0E0F17733F26C284CF8CD161C9BE2B3A" author="Morlet L" page="166" year="1889">
|
||
<bibRefCitation id="2DF5141C1AED9495AF26ADBAD38E5A8D" author="Morlet L" firstAuthor="Morlet" journalOrPublisher="Journal de Conchyliologie" page="166" pagination="121-199" refId="B47" refString="Morlet L (1889) Catalogue des coquilles recueillies, par M. Pavie, dans le Cambodge et le Royaume de Siam, et description ďespèces nouvelles. Journal de Conchyliologie 37: 121–199." title="Catalogue des coquilles recueillies, par M. Pavie, dans le Cambodge et le Royaume de Siam, et description ďespèces nouvelles." volume="37" year="1889">Morlet 1889: 166</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="9F338B6E8B758D6390DD906FDFE99FA0" author="Fischer P" page="221" year="1891">
|
||
<bibRefCitation id="771508AC5558DFA33F85938291D1C4A6" DOI="10.5962/bhl.title.14809" author="Fischer P" firstAuthor="Fischer" journalOrPublisher="Bulletin de la Société d’Histoire Naturelle d’Autun" page="221" pagination="87-276" refId="B15" refString="Fischer P (1891) Catalogue et distribution géographique des mollusques terrestres, fluviatiles & marins d’une partie de l’Indo-chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Bulletin de la Société d’Histoire Naturelle d’Autun 4: 87–276. https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution géographique des mollusques terrestres, fluviatiles & marins d’une partie de l’Indo-chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." volume="4" year="1891">Fischer 1891: 221</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="35702D7C13D7F7593182DC23938ECD5D" author="Fischer-Piette E" page="154" year="1950">
|
||
<bibRefCitation id="FA29275A1D458715A9978F5760A79FA0" author="Fischer-Piette E" firstAuthor="Fischer-Piette" journalOrPublisher="Journal de Conchyliologie" page="154" pagination="149-180" refId="B17" refString="Fischer-Piette E (1950) Liste des types décrits dans le Journal de Conchyliologie et conservés dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149–180." title="Liste des types décrits dans le Journal de Conchyliologie et conservés dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette 1950: 154</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="7E1968580A7D42A9D67A5BE97507D946">
|
||
<bibRefCitation id="DD22E55CB78FF5C61AD77F9E46E6D298" DOI="10.1007/s10750-017-3104-8" author="Zieritz" etAl="et al." firstAuthor="Zieritz" journalOrPublisher="Hydrobiologia" pagination="29-44" refId="B63" refString="Zieritz A, Bogan AE, Froufe E, Klishko O, Kondo T, Kovitvadhi U, Kovitvadhi S, Lee JH, Lopes-Lima M, Pfeiffer JM, Sousa R, Van Do T, Vikhrev I, Zanatta DT (2018) Diversity, biogeography and conservation of freshwater mussels (Bivalvia: Unionida) in East and Southeast Asia. Hydrobiologia 810: 29–44. https://doi.org/10.1007/s10750-017-3104-8" title="Diversity, biogeography and conservation of freshwater mussels (Bivalvia: Unionida) in East and Southeast Asia." volume="810" year="2018">Zieritz et al. 2018</bibRefCitation>
|
||
</treatmentCitation>
|
||
: supplementary material 1.
|
||
<treatmentCitation id="666E3A6CF5323BC37F24EE3FA597D863" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Kondakov AV & Lyubas AA & Soboleva AA & Chan N & Lunn Z & Win T & Inkhavilay K" page="11" year="2023">
|
||
<bibRefCitation id="B87C9EE9D17F3E6C088EE42979C59F6D" DOI="10.3390/w15173117" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Kondakov AV & Lyubas AA & Soboleva AA & Chan N & Lunn Z & Win T & Inkhavilay K" etAl="et al." firstAuthor="Bolotov" page="11" refId="B8" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Kondakov AV, Lyubas AA, Soboleva AA, Chan N, Lunn Z, Win T, Inkhavilay K (2023) Integrative taxonomic reappraisal and evolutionary biogeography of the most diverse freshwater mussel clade from Southeast Asia (Pseudodontini). Water 15: 3117. https://doi.org/10.3390/w15173117" year="2023">Bolotov et al. 2023: 11</bibRefCitation>
|
||
</treatmentCitation>
|
||
. Syn. nov.
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D0D687FBB5E0DBA1D5C907771C182A5D">
|
||
<treatmentCitationGroup id="7FFF49774C30C55CA866D80D37D414E4">
|
||
<taxonomicName id="2EF6438D767F8D53A353E87A5C2D03C6" baseAuthorityName="Morlet" baseAuthorityYear="1884" class="Bivalvia" family="Unionidae" genus="Margaritana" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni">
|
||
<emphasis id="FAA7C0E93F6FD2B0CF6BF5E40144B15F" italics="true">Margaritana thomsoni</emphasis>
|
||
</taxonomicName>
|
||
<comment id="EB53CCDF400155BA9988648EB2EEA1F4">
|
||
—
|
||
<treatmentCitation id="13B42788B1907964DEA0D153BC41198D" author="Paetel F" page="174" year="1890">
|
||
<bibRefCitation id="8B6E8C806A85A8405289BD0B6340919C" author="Paetel F" firstAuthor="Paetel" page="174" refId="B51" refString="Paetel F (1890) Catalog der Conchylien-Sammlung. Vol. 3, 256 pp." year="1890">Paetel 1890: 174</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="E482BFC5B6FEDB2A4E5AC6A6B8714783">
|
||
<treatmentCitationGroup id="1A3175609A877BB6F482EDDA53BB7F1E">
|
||
<taxonomicName id="5DD508EB6E6362ED4B5E0E1B8F18EBFF" ID-CoL="7STYL" authorityName="Morlet" authorityYear="1884" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni" subGenus="Pseudodon">
|
||
<emphasis id="5953264025055DFFABEFBB920F18C8E7" italics="true">Pseudodon</emphasis>
|
||
(
|
||
<emphasis id="8F539B2F04D05FFFD6C476E02DB29746" italics="true">Pseudodon</emphasis>
|
||
)
|
||
<emphasis id="B85D205E8E9B6FE979E5169B1518C31F" italics="true">thomsoni</emphasis>
|
||
</taxonomicName>
|
||
<comment id="CA554CA0605005FB81B04C0D0C003101">
|
||
—
|
||
<treatmentCitation id="741E3944ADACB8FCBED4B949B26CCC0A" author="Simpson CT" page="838" year="1900">
|
||
<bibRefCitation id="C57FCFCA0398C44BF0B475DA8542B31A" DOI="10.5479/si.00963801.22-1205.501" author="Simpson CT" firstAuthor="Simpson" journalOrPublisher="Proceedings of the United States National Museum" page="838" pagination="501-1044" refId="B58" refString="Simpson CT (1900) Synopsis of the naiades, or pearly fresh-water mussels. Proceedings of the United States National Museum 22: 501–1044. https://doi.org/10.5479/si.00963801.22-1205.501" title="Synopsis of the naiades, or pearly fresh-water mussels." volume="22" year="1900">Simpson 1900: 838</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="62393DC12462052DB823C795E3C99A1F" author="Simpson CT" page="1092" year="1914">
|
||
<bibRefCitation id="7ED5912F6618055A0D06C4EC61CFCA72" DOI="10.5962/bhl.title.10910" author="Simpson CT" firstAuthor="Simpson" journalOrPublisher="Bryant Walker, Detroit, Michigan" page="1092" refId="B59" refString="Simpson CT (1914) A descriptive catalogue of the naiades, or pearly fresh-water mussels. Parts I – III. Bryant Walker, Detroit, Michigan, 1540 pp. https://doi.org/10.5962/bhl.title.10910" title="A descriptive catalogue of the naiades, or pearly fresh-water mussels. Parts I - III." year="1914">Simpson 1914: 1092</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 1093.
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="709FCAB20B7E27E67E838849101421CE">
|
||
<treatmentCitationGroup id="B4F26CF99FFA56789DF9F615A4ACE050">
|
||
<taxonomicName id="7B2A0B475599EE150DB11D01E487AC2D" ID-CoL="7STYL" authorityName="Morlet" authorityYear="1884" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni" subGenus="Bineurus">
|
||
<emphasis id="3A722C767FF7E898578AB2A7055FE636" italics="true">Pseudodon</emphasis>
|
||
(
|
||
<emphasis id="0B65D5C9BB2D8E061C2948C9DEE63343" italics="true">Bineurus</emphasis>
|
||
)
|
||
<emphasis id="D762BEB5662D134411BDADD1C1E8D5EB" italics="true">thomsoni</emphasis>
|
||
</taxonomicName>
|
||
<comment id="79FEFE2B2D44065E8E37FF3683158B8B">
|
||
—
|
||
<treatmentCitation id="82067D7A38181E05228656C75AB0B68B" author="Haas F" page="336 - 337" year="1920">
|
||
<bibRefCitation id="A8E057F4ED05F7642609F4F2D8433D44" DOI="10.5962/bhl.title.120018" author="Haas F" firstAuthor="Haas" page="336 - 337" refId="B23" refString="Haas F (1920) Die Unioniden. In: Küster HC (Ed) Systematisches Conchylien-Cabinet von Martini und Chemnitz 9. pt. 2, h. 53. 305–344, pls. 369–373. https://doi.org/10.5962/bhl.title.120018" year="1920">Haas 1920: 336–337</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="7371AC9F488BD337A77E8209EBE3120B" author="Haas F" page="144" year="1924">
|
||
<bibRefCitation id="FDE31F42A5A474F67D562411579F18C9" author="Haas F" firstAuthor="Haas" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" page="144" pagination="129-203" refId="B24" refString="Haas F (1924) Beiträge zu einer Monographie der asiatischen Unioniden. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 38: 129–203." title="Beiträge zu einer Monographie der asiatischen Unioniden." volume="38" year="1924">Haas 1924: 144</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="05A2C99650DA737862BF0516C0AF4D33" author="Graf DL & Cummings KS" page="311" year="2007">
|
||
<bibRefCitation id="89CA748729C19D1D065878F8930FE3C3" DOI="10.1093/mollus/eym029" author="Graf DL & Cummings KS" firstAuthor="Graf" journalOrPublisher="Journal of Molluscan Studies" page="311" pagination="291-314" refId="B19" refString="Graf DL, Cummings KS (2007) Review of the systematics and global diversity of freshwater mussel species (Bivalvia: Unionoida). Journal of Molluscan Studies 73: 291–314. https://doi.org/10.1093/mollus/eym029" title="Review of the systematics and global diversity of freshwater mussel species (Bivalvia: Unionoida)." volume="73" year="2007">Graf and Cummings 2007: 311</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="411F94C1C768E375EB70B158AD5211C3">
|
||
<treatmentCitationGroup id="63B1AF4396DFFB71B830401B207B0B29">
|
||
<comment id="4D0ABFFE9F257050B991D0B5770690A9">
|
||
?
|
||
<taxonomicName id="04E79717194E7D0E267CDC679B65CF85" ID-CoL="7STYL" authorityName="Morlet" authorityYear="1884" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni">
|
||
<emphasis id="18B8A91713D00A48AEE2ECD612D29B4E" italics="true">Pseudodon thomsoni</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="5852C5A99AD1661A9CC8D6AEC80378E1" author="Konopleva ES & Bolotov IN & Pfeiffer JM & Vikhrev IV & Kondakov AV & Gofarov MY & Tomilova AA & Tanmuangpak K & Tumpeesuwan S" page="11" year="2021">
|
||
<bibRefCitation id="AD970CD73219913FA0E70FA69D83EC3E" DOI="10.1038/s41598-021-87633-w" author="Konopleva ES & Bolotov IN & Pfeiffer JM & Vikhrev IV & Kondakov AV & Gofarov MY & Tomilova AA & Tanmuangpak K & Tumpeesuwan S" etAl="et al." firstAuthor="Konopleva" page="11" refId="B37" refString="Konopleva ES, Bolotov IN, Pfeiffer JM, Vikhrev IV, Kondakov AV, Gofarov MY, Tomilova AA, Tanmuangpak K, Tumpeesuwan S (2021) New freshwater mussels from two Southeast Asian genera Bineurus and Thaiconcha (Pseudodontini, Gonideinae, Unionidae). Scientific Reports 11: 8244. https://doi.org/10.1038/s41598-021-87633-w" year="2021">Konopleva et al. 2021: 11</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 3 h.
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EQ1AG" type="type material">
|
||
<paragraph id="DE757AA1F50BDC23961B6EA51F9CCCA9">
|
||
<heading id="9F35330CE00CE1800A35D4D759813A21" reason="title">Type material.</heading>
|
||
</paragraph>
|
||
<paragraph id="CD3E54C2A67A1954E277E52E7016186C">
|
||
<emphasis id="542775FC1CF0C9EB239B767B958DFA2F" bold="true" italics="true">
|
||
<typeStatus id="C465706B445263B3F4FECF409D9815CA" type="syntype">Syntypes</typeStatus>
|
||
</emphasis>
|
||
<named-content id="80910B09393E6A35C592CAF0C907091D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national ďHistoire naturelle, Paris">
|
||
<collectionCode id="0B1081217344A6F804D3664E07D3FB6E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</named-content>
|
||
- IM- 2000–34623 (
|
||
<specimenCount id="31D05640CA5D2A0BC5AC1DC44297A7F9" count="2" type="generic">2 shells</specimenCount>
|
||
).
|
||
<emphasis id="AF486B2B4F17F2EF1BF4CE9974FB8204" bold="true" italics="true">
|
||
<typeStatus id="A9FCBF6BE2E7B8FDD42D62D8614E463D" type="syntype">Syntype</typeStatus>
|
||
</emphasis>
|
||
<named-content id="A21FA0DEA19D41A6A15084933468BA9A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national ďHistoire naturelle, Paris">
|
||
<collectionCode id="8FA5C8CA28C480399265FACEAC5E60BD" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</named-content>
|
||
- IM- 2000-1800 (
|
||
<specimenCount id="989BB5686281DC1D83AEABADDD8A576B" count="1" type="generic">1 shell</specimenCount>
|
||
) of
|
||
<taxonomicName id="80AF9BC788AEB12FA904BD09CED82177" authority="Morlet, 1884" authorityName="Morlet" authorityYear="1884" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni">
|
||
<emphasis id="C24CAD1AA8EEC9C4A86A40833B676C62" italics="true">Pseudodon thomsoni</emphasis>
|
||
<bibRefCitation id="E27FF0C27C125926FBC88F4286536186" author="Morlet L" journalOrPublisher="Journal de Conchyliologie" pagination="386-403" refId="B46" refString="Morlet L (1884) Description d'espece nouvelles de coquilles, recueillies par M. Pavie, au Cambodge. Journal de Conchyliologie 32: 386 - 403." title="Description d’espèce nouvelles de coquilles, recueillies par M. Pavie, au Cambodge." volume="32" year="1884">Morlet, 1884</bibRefCitation>
|
||
</taxonomicName>
|
||
, (inadvertently stated as
|
||
<typeStatus id="4138C9A7329024D1CA2E88D41CA67569" type="holotype">holotype</typeStatus>
|
||
by
|
||
<bibRefCitation id="E9EEFFAAF239AF90625AF0CCA9A8F054" author="Fischer-Piette E" journalOrPublisher="Journal de Conchyliologie" pagination="149-180" refId="B17" refString="Fischer-Piette E (1950) Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149 - 180." title="Liste des types décrits dans le Journal de Conchyliologie et conservés dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette, 1950</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ET2AG" type="material">
|
||
<paragraph id="D5C1449EFABB107728AE049A847CDF99">
|
||
<heading id="52048D232B4750A47944AABEE559E246" reason="title">Other material examined.</heading>
|
||
</paragraph>
|
||
<paragraph id="EA7305D9700A9D4B907B4B39CC4C1F5A">
|
||
<materialsCitation id="9AA83FBFF2AA6F81B6D60828C010D7E9" collectingDate="2015-01-31" collectionCode="MUMNH" collectorName="E. Jeratthitikul" country="Thailand" county="Aranyaprathet District" latitude="13.6694" location="Phrom Hot Stream" longLatPrecision="7" longitude="102.521" municipality="Aranyaprathet Subdistrict" specimenCode="UNI 0544" specimenCount="1" stateProvince="Sa Kaeo Province">
|
||
<collectingCountry id="0DB2A177E22C45721E5B115ABD12F0AD" name="Thailand">Thailand</collectingCountry>
|
||
–
|
||
<emphasis id="9A8B3E6DFE3AAC3497AA1E57F28248BC" bold="true">
|
||
<collectingRegion id="510B6C7B8DC7240F5D522D0328410A03" country="Thailand" name="Sa Kaeo">Sa Kaeo Province</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="D28C5DA1B5199DED7A5F874EE96AEE87" count="1" type="generic">1 shell</specimenCount>
|
||
;
|
||
<collectingCounty id="B5AFBBBD31F6EAD107B43E2891AA6D47">Aranyaprathet District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="86C5A9715EC99B3DA1ADC273A2CD3F0F">Aranyaprathet Subdistrict</collectingMunicipality>
|
||
,
|
||
<collectingMunicipality id="82874AA38E2B3A59EF4F69498D33EB13">Tonle Sap Basin</collectingMunicipality>
|
||
,
|
||
<location id="6954004360B3330FFC47E38B381C5A49" LSID="urn:lsid:plazi:treatment:9BDBEDA98C975EA793681649DEF9E8B2:6954004360B3330FFC47E38B381C5A49" country="Thailand" county="Aranyaprathet District" latitude="13.6694" longLatPrecision="7" longitude="102.521" municipality="Aranyaprathet Subdistrict" name="Phrom Hot Stream" stateProvince="Sa Kaeo Province">Phrom Hot Stream</location>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E52AG" specific-use="{"type":"Point","coordinates":[102.521000,13.669400]}">
|
||
<geoCoordinate id="AFBC6FD0127526F3D3C77E4549E77F8B" degrees="13.6694" direction="north" orientation="latitude" precision="5" value="13.6694">13.6694 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CD6E842EB196B6E75B32891951E0A7CF" degrees="102.5210" direction="east" orientation="longitude" precision="5" value="102.521">102.5210 ° E</geoCoordinate>
|
||
</named-content>
|
||
;
|
||
<date id="A341E4463AA2080FF493D472B4C77EAA" value="2015-01-31">
|
||
<collectingDate id="D7268F1BD451F4A96A5E39FEE37F943F" value="2015-01-31">31 Jan. 2015</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="3B2B039541B5B68EDBA8D233083B8810">E. Jeratthitikul</collectorName>
|
||
leg.;
|
||
<named-content id="BA807A6FEA5407DAA39660C014AD5986" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">
|
||
<collectionCode id="5676C3079A3191DB6B38C9DF65D08719">MUMNH</collectionCode>
|
||
</named-content>
|
||
-
|
||
<specimenCode id="6B931E3582D801175CDB8F0815055898" collectionCode="UNI" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" type="">UNI 0544</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6D6DDCCFCEF60B446740A4A1BC32A930" collectionCode="MUMNH" country="Thailand" county="Aranyaprathet District" latitude="13.6718" location="Phrom Hot Stream" longLatPrecision="7" longitude="102.5166" municipality="Aranyaprathet Subdistrict" specimenCode="UNI 3428" specimenCount="1" stateProvince="Sa Kaeo">
|
||
<specimenCount id="1D105E86C3ED4CDEC5FA164C145773E0" count="1" type="generic">1 shell</specimenCount>
|
||
;
|
||
<collectingCounty id="D6442A266C3E4AEC8F3018D709EFC665">Aranyaprathet District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="0EFA7C34923A1686D36A6A3BC084C39A">Aranyaprathet Subdistrict</collectingMunicipality>
|
||
,
|
||
<collectingMunicipality id="B5A5CDCA155D279D51425CF7B97E67BC">Tonle Sap Basin</collectingMunicipality>
|
||
,
|
||
<location id="EFAFC4F520B194D26D56787E40BC8C65" LSID="urn:lsid:plazi:treatment:9BDBEDA98C975EA793681649DEF9E8B2:EFAFC4F520B194D26D56787E40BC8C65" country="Thailand" county="Aranyaprathet District" latitude="13.6718" longLatPrecision="7" longitude="102.5166" municipality="Aranyaprathet Subdistrict" name="Phrom Hot Stream" stateProvince="Sa Kaeo">Phrom Hot Stream</location>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EL3AG" specific-use="{"type":"Point","coordinates":[102.516600,13.671800]}">
|
||
<geoCoordinate id="57F2FFE6F29C7450633A0AEDB499762E" degrees="13.6718" direction="north" orientation="latitude" precision="5" value="13.6718">13.6718 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A8793125915E39AEA5C3AAD8B4FB9693" degrees="102.5166" direction="east" orientation="longitude" precision="5" value="102.5166">102.5166 ° E</geoCoordinate>
|
||
</named-content>
|
||
;
|
||
<named-content id="0D16228B5CD7CF0D707D518DD38C23D1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">
|
||
<collectionCode id="9FEF958809CE11EAEDA64DEEAEC9FE4C">MUMNH</collectionCode>
|
||
</named-content>
|
||
-
|
||
<specimenCode id="F8A7263C8E73358F0C3C1400539708E0" collectionCode="UNI" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" type="">UNI 3428</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="DEE45A1B7AB933DB9B8F3E63F6193C85" collectingDate="2015-05-05" collectionCode="MUMNH" collectorName="P. Prasankok" country="Thailand" county="Watthana Nakhon District" latitude="13.749" location="Phrom Hot Stream" longLatPrecision="7" longitude="102.4271" municipality="Phak Kha Subdistrict" specimenCode="UNI 0300, 0301" specimenCount="2" stateProvince="Sa Kaeo">
|
||
<specimenCount id="3FE3A23BCEB49FB39A79F7AA82792011" count="2" type="generic">2 shells</specimenCount>
|
||
;
|
||
<collectingCounty id="F848F809D0B5A68614CE64883F7EC378">Watthana Nakhon District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="99234BA9CFE7BCC6B4F7641614FC2302">Phak Kha Subdistrict</collectingMunicipality>
|
||
,
|
||
<collectingMunicipality id="14F7230868D57373E62198DDAE9AB0A9">Tonle Sap Basin</collectingMunicipality>
|
||
,
|
||
<location id="0EC666122EEE4585652CD9CCDD1E00A1" LSID="urn:lsid:plazi:treatment:9BDBEDA98C975EA793681649DEF9E8B2:0EC666122EEE4585652CD9CCDD1E00A1" country="Thailand" county="Watthana Nakhon District" latitude="13.749" longLatPrecision="7" longitude="102.4271" municipality="Phak Kha Subdistrict" name="Phrom Hot Stream" stateProvince="Sa Kaeo">Phrom Hot Stream</location>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EY3AG" specific-use="{"type":"Point","coordinates":[102.427100,13.749000]}">
|
||
<geoCoordinate id="F9241948B00FEB7E75242684A18C68D3" degrees="13.7490" direction="north" orientation="latitude" precision="5" value="13.749">13.7490 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1CC344399EEC3AA4081638B25C2A12D1" degrees="102.4271" direction="east" orientation="longitude" precision="5" value="102.4271">102.4271 ° E</geoCoordinate>
|
||
</named-content>
|
||
;
|
||
<date id="CB3915C38E2EB9302E111F4851740DDB" value="2015-05-05">
|
||
<collectingDate id="95979E449441B84F1C98AA73B22F5565" value="2015-05-05">5 May 2015</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="DE31B3392EE70196B642C08253C61D08">P. Prasankok</collectorName>
|
||
leg.;
|
||
<named-content id="BD91C0D0D4C6D537D7AE9DB202852863" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">
|
||
<collectionCode id="2E89905CCA27C4D93BE9B91158B4F43C">MUMNH</collectionCode>
|
||
</named-content>
|
||
-
|
||
<specimenCode id="C5EA0470411B74725D28F182F9B03F06" collectionCode="UNI" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" type="">UNI 0300</specimenCode>
|
||
to
|
||
<specimenCode id="95BD503436B3F25EC21ADD869124FE23" collectionCode="MUMNH" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" type="">0301</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EC4AG" type="diagnosis">
|
||
<paragraph id="2AB9866515F77D69EED42DC98C39EF41">
|
||
<heading id="FAE363C417B46152CE8F6F2E5056F8FF" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="4D1441AE61A294D155C436BCD6C237B2">Shell large, subrhomboidal to ovate, moderately thickened, rather inflated. Anteriorly constricted, dorsal slightly curved, slightly elevated posteriorly. Posterior slope with one or two faint folds. Umbo rounded, moderately elevated. Shell surface with irregular growth lines, roughened on posterior slope and border of shell. Right valve with one smooth tubercular pseudocardinal tooth, left valve with hill-like or triangulate pseudocardinal tooth, subcompressed. V-shaped furrow on posterior end of hinge structure prominent, wide. Anterior adductor muscle scar impressed, ovate, separated from anterior protractor muscle scar. Umbo cavity moderately deep.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EH4AG" type="Differential diagnosis">
|
||
<paragraph id="ED9C6DA278BB6B1DB12A5B231ABAA452">
|
||
<heading id="3A038CB579C18EF5C0A580FBA67BE5B3" reason="title">Differential diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="2F2E52806F368F7A5659DEB1855EB848">
|
||
This species is much larger and more inflated than the type species. Irregular growth lines on the shell surface are rougher, especially on the posterior slope and along the shell border. This species is also distinct from congenerics due to 14 fixed nucleotide substitutions in the
|
||
<abbrev id="ABBRID0EN4AG" xlink_title="mitochondrial cytochrome c oxidase subunit I">COI</abbrev>
|
||
gene fragment (Table
|
||
<tableCitation id="5182B41BC81A399FC125C075D2F57151" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean genetic distances (uncorrected p-distance: % ± SD) based on 660 - bp COI fragment sequences among Songkhlanaia species (below diagonal), and within each species (in bold)." httpUri="http://table.plazi.org/id/D9D5132FC2BA2A9EAA683B9B5BCF4CF9" tableUuid="D9D5132FC2BA2A9EAA683B9B5BCF4CF9">2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EV4AG" type="distribution">
|
||
<paragraph id="9240EF24CB6C2B543C8C7F87840661E9">
|
||
<heading id="C28FB34DCE93065719F5575B18F6A863" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="E3E39EE9161B32237B12B6B6D5ACD793">
|
||
Tonle Sap Basin in
|
||
<collectingCountry id="21CE0F47C764336A43AF10348D05B00E" name="Thailand">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="E0B82F22F06973457BCDA145B407FA0E" name="Cambodia">Cambodia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="BFC2E746E30F11B1A6BA51A109E65330" author="Crosse and Fischer" firstAuthor="Crosse" journalOrPublisher="Journal de Conchyliologie" pagination="313-342" refId="B11" refString="Crosse H, Fischer P (1876) Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873. Journal de Conchyliologie 24: 313–342." title="Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873." volume="24" year="1876">Crosse and Fischer 1876</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="FF34396826F2B254A34F2DEE97088ABE" DOI="10.5962/bhl.title.10273" author="Deshayes and Jullien" firstAuthor="Deshayes" journalOrPublisher="Bulletin des Nouvelles Archives du Muséum d’Histoire Naturelle" pagination="115-163" refId="B13" refString="Deshayes G-P, Jullien J (1876) Mémoire sur les mollusques nouveaux du Cambodge: Envoyés au muséum par M. le docteur Jullien. Bulletin des Nouvelles Archives du Muséum d’Histoire Naturelle 10: 115–163. https://doi.org/10.5962/bhl.title.10273" title="Mémoire sur les mollusques nouveaux du Cambodge: Envoyés au muséum par M. le docteur Jullien." volume="10" year="1876">Deshayes and Jullien 1876</bibRefCitation>
|
||
; this study), and the Lower Mekong Basin in
|
||
<collectingCountry id="FD61E01C64D9A170E3194CE040E84E95" name="Cambodia">Cambodia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="D633EC5F2228DFBA1C4DCBCAF14EA66A" author="Morlet" firstAuthor="Morlet" journalOrPublisher="Journal de Conchyliologie" pagination="386-403" refId="B46" refString="Morlet L (1884) Description d’espèce nouvelles de coquilles, recueillies par M. Pavie, au Cambodge. Journal de Conchyliologie 32: 386–403." title="Description d’espèce nouvelles de coquilles, recueillies par M. Pavie, au Cambodge." volume="32" year="1884">Morlet 1884</bibRefCitation>
|
||
) (Fig.
|
||
<figureCitation id="82B51FF0A939BB87165CCDFAD1330ED9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B map showing the geographical distribution of all known Songkhlanaia species (circles) and their type locality (stars). Map was generated using QGIS v. 3.36. 0 with the outline of major river basins from the Freshwater Ecoregions of the World (Abell et al. 2008), river and lake topology from the HydroSHEDS database (https://www.hydrosheds.org), and the map raster data from the NASA EARTHDATA (https://www.earthdata.nasa.gov/)." figureDoi="10.3897/zookeys.1224.140549.figure3" httpUri="https://binary.pensoft.net/fig/1236495">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EL5AG" type="comments">
|
||
<paragraph id="C7AA80D4D341E6BCBB4DE2A6C777CD26">
|
||
<heading id="8E1D1C20963C59C1DD2019285D117EA3" reason="title">Comments.</heading>
|
||
</paragraph>
|
||
<paragraph id="7143DB5C31E58EB6080218AE01DDFF30">
|
||
Originally,
|
||
<taxonomicName id="857A5EABCBF0E9AABF0FCD527D8B8F2C" authorityName="Jeratthitikul & Sutcharit & Prasankok" authorityYear="2025" baseAuthorityName="Crosse & Fischer" baseAuthorityYear="1876" class="Bivalvia" family="Unionidae" genus="Songkhlanaia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="EFF8771F18CFA4C8EA08AB3D10699D29" italics="true">Songkhlanaia moreleti</emphasis>
|
||
</taxonomicName>
|
||
was described based on a partially misidentified specimen from
|
||
<collectingCountry id="F7CB025F28E88F379FFA7AF82A8469E0" name="Cambodia">Cambodia</collectingCountry>
|
||
as
|
||
<taxonomicName id="49BDF4A17B55DBF7EDF3E3A747084EFD" authority="Morelet, 1866" authorityName="Morelet" authorityYear="1866" class="Bivalvia" family="Unionidae" genus="Monocondylus" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="tumidus">
|
||
<emphasis id="14E6AFF5EE55FD749171D47E7E0FE0E6" italics="true">Monocondylus tumidus</emphasis>
|
||
<bibRefCitation id="A61CE0464747732535CB058E735DB322" author="Morelet A" journalOrPublisher="Journal de Conchyliologie" pagination="62-64" refId="B45" refString="Morelet A (1866) Description d'especes appartenant a la faune malacologique de l'Indo-Chine. Journal de Conchyliologie 14: 62 - 64." title="Description d’espèces appartenant à la faune malacologique de l’Indo-Chine." volume="14" year="1866">Morelet, 1866</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="7DF6B5F5DB27CF4854779233C2220D93" DOI="10.5962/bhl.title.10273" author="Deshayes G-P & Jullien J" firstAuthor="Deshayes" journalOrPublisher="Bulletin des Nouvelles Archives du Muséum d’Histoire Naturelle" pagination="115-163" refId="B13" refString="Deshayes G-P, Jullien J (1876) Mémoire sur les mollusques nouveaux du Cambodge: Envoyés au muséum par M. le docteur Jullien. Bulletin des Nouvelles Archives du Muséum d’Histoire Naturelle 10: 115–163. https://doi.org/10.5962/bhl.title.10273" title="Mémoire sur les mollusques nouveaux du Cambodge: Envoyés au muséum par M. le docteur Jullien." volume="10" year="1876">Deshayes and Jullien (1876)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="40562C244281F665E524D365D16194FB" author="Crosse H & Fischer P" firstAuthor="Crosse" journalOrPublisher="Journal de Conchyliologie" pagination="313-342" refId="B11" refString="Crosse H, Fischer P (1876) Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873. Journal de Conchyliologie 24: 313–342." title="Mollusques fluviatiles, recueillis au Cambodge, par la mission scientifique francaise de 1873." volume="24" year="1876">Crosse and Fischer (1876)</bibRefCitation>
|
||
reexamined the specimens and provided it with a new name,
|
||
<taxonomicName id="2914DA2BF8F4E5A3E576E0F5795750D6" authorityName="Crosse & Fischer" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="EC8D20EA7B9FCA6890A48EACA7D1D3D4" italics="true">Pseudodon moreleti</emphasis>
|
||
</taxonomicName>
|
||
. This nominal species had been recognized as valid by subsequent studies for more than a hundred years (e. g.,
|
||
<bibRefCitation id="2600B4756C350FC66F5CC4980864BCB5" DOI="10.5962/bhl.title.14809" author="Fischer" firstAuthor="Fischer" journalOrPublisher="Bulletin de la Société d’Histoire Naturelle d’Autun" pagination="87-276" refId="B15" refString="Fischer P (1891) Catalogue et distribution géographique des mollusques terrestres, fluviatiles & marins d’une partie de l’Indo-chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Bulletin de la Société d’Histoire Naturelle d’Autun 4: 87–276. https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution géographique des mollusques terrestres, fluviatiles & marins d’une partie de l’Indo-chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." volume="4" year="1891">Fischer 1891</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="CE6F0AC8667152C92A0C65D4DD6C9EBA" author="Fischer and Dautzenberg" firstAuthor="Fischer" journalOrPublisher="Mission Pavie, Etudes diverses" pagination="390-442" refId="B16" refString="Fischer H, Dautzenberg P (1904) Catalogue des mollusques terrestres et fluviatiles de l’Indo-Chine orientale cites jusqu’à ce jour. Mission Pavie, Etudes diverses 3: 390–442." title="Catalogue des mollusques terrestres et fluviatiles de l’Indo-Chine orientale cites jusqu’à ce jour." volume="3" year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9BEB5AF9FDDD950D0C6889747262BF4A" author="Dautzenberg and Fischer" firstAuthor="Dautzenberg" journalOrPublisher="Journal de Conchyliologie" pagination="343-471" refId="B12" refString="Dautzenberg P, Fischer H (1905) Liste des mollusques récoltés par M. H. Mansuy en Indo-chine et au Yunnan et description d’especes nouvelles. Journal de Conchyliologie 53: 343–471." title="Liste des mollusques récoltés par M. H. Mansuy en Indo-chine et au Yunnan et description d’especes nouvelles." volume="53" year="1905">Dautzenberg and Fischer 1905</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D26EA630158DF4A8E158B780270B1773" DOI="10.5479/si.00963801.22-1205.501" author="Simpson" firstAuthor="Simpson" journalOrPublisher="Proceedings of the United States National Museum" pagination="501-1044" refId="B58" refString="Simpson CT (1900) Synopsis of the naiades, or pearly fresh-water mussels. Proceedings of the United States National Museum 22: 501–1044. https://doi.org/10.5479/si.00963801.22-1205.501" title="Synopsis of the naiades, or pearly fresh-water mussels." volume="22" year="1900">Simpson 1900</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="18D23B9C186FC4A99D8E71880D9A778A" DOI="10.5962/bhl.title.10910" author="Simpson" firstAuthor="Simpson" journalOrPublisher="Bryant Walker, Detroit, Michigan" refId="B59" refString="Simpson CT (1914) A descriptive catalogue of the naiades, or pearly fresh-water mussels. Parts I – III. Bryant Walker, Detroit, Michigan, 1540 pp. https://doi.org/10.5962/bhl.title.10910" title="A descriptive catalogue of the naiades, or pearly fresh-water mussels. Parts I - III." year="1914">1914</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DA2DCFF44C233DE205BEC960134D820F" author="Haas" firstAuthor="Haas" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pagination="129-203" refId="B24" refString="Haas F (1924) Beiträge zu einer Monographie der asiatischen Unioniden. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 38: 129–203." title="Beiträge zu einer Monographie der asiatischen Unioniden." volume="38" year="1924">Haas 1924</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A896633D9E78D7C9D2ABE9A9170F34FB" author="Haas" editor="Martens R & Hennig W" firstAuthor="Haas" journalOrPublisher="Walter de Gruyter, Berlin" pagination="1-663" refId="B25" refString="Haas F (1969) Superfamilia Unionacea. In: Martens R, Hennig W (Eds) Das Tierrich, Lieferung Vol 88. Walter de Gruyter, Berlin, 1–663." title="Superfamilia Unionacea." volumeTitle="Das Tierrich, Lieferung Vol 88." year="1969">1969</bibRefCitation>
|
||
). Later,
|
||
<bibRefCitation id="50D6E980DFB184290BDD39A3D8AE3D46" author="Brandt RAM" firstAuthor="Brandt" journalOrPublisher="Archiv für Molluskenkunde" page="271" pagination="1-423" refId="B9" refString="Brandt RAM (1974) The non-marine aquatic Mollusca of Thailand. Archiv für Molluskenkunde 105: 1–423." title="The non-marine aquatic Mollusca of Thailand." volume="105" year="1974">Brandt (1974: 271)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="03F040D1F5D2C886EB7076B702CFAE21" DOI="10.1093/mollus/eym029" author="Graf DL & Cummings KS" firstAuthor="Graf" journalOrPublisher="Journal of Molluscan Studies" page="311" pagination="291-314" refId="B19" refString="Graf DL, Cummings KS (2007) Review of the systematics and global diversity of freshwater mussel species (Bivalvia: Unionoida). Journal of Molluscan Studies 73: 291–314. https://doi.org/10.1093/mollus/eym029" title="Review of the systematics and global diversity of freshwater mussel species (Bivalvia: Unionoida)." volume="73" year="2007">Graf and Cummings (2007: 311)</bibRefCitation>
|
||
listed this taxon as a junior synonym of either species or subspecies of
|
||
<taxonomicName id="F8F843F0257AA7DA687E675FD6F60041" authority="Morelet, 1866" authorityName="Morelet" authorityYear="1866" class="Bivalvia" family="Unionidae" genus="Monocondylus" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="tumidus">
|
||
‘
|
||
<emphasis id="35F0FC3E228476AF59DCB64D5C9FE8FB" italics="true">tumidus</emphasis>
|
||
<bibRefCitation id="DEC62F8FD76F2BB203A63860D97D4E6B" author="Morelet A" journalOrPublisher="Journal de Conchyliologie" pagination="62-64" refId="B45" refString="Morelet A (1866) Description d'especes appartenant a la faune malacologique de l'Indo-Chine. Journal de Conchyliologie 14: 62 - 64." title="Description d’espèces appartenant à la faune malacologique de l’Indo-Chine." volume="14" year="1866">Morelet, 1866</bibRefCitation>
|
||
</taxonomicName>
|
||
’. Recently,
|
||
<bibRefCitation id="5A0BD10357051463424D3CC97B950FB2" DOI="10.1038/s41598-020-63612-5" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Lopes-Lima M & Bogan AE & Lunn Z & Chan N & Win T & Aksenova OV & Tomilova AA & Tanmuangpak K & Tumpeesuwan S & Kondakov AV" etAl="et al." firstAuthor="Bolotov" refId="B6" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Lopes-Lima M, Bogan AE, Lunn Z, Chan N, Win T, Aksenova OV, Tomilova AA, Tanmuangpak K, Tumpeesuwan S, Kondakov AV (2020) New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10: 6616. https://doi.org/10.1038/s41598-020-63612-5" year="2020">Bolotov et al. (2020)</bibRefCitation>
|
||
raised this species as valid and placed it in their new genus,
|
||
<taxonomicName id="41F6A93370680332713807BA4E65D251" class="Bivalvia" family="Unionidae" genus="Sundadontina" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
|
||
<emphasis id="40B1A7E94044752A2BB6B3A84EEF5341" italics="true">Sundadontina</emphasis>
|
||
</taxonomicName>
|
||
. However, this resurrection and generic placement seemed provisionally based on conchological characters alone. Fortunately, specimens collected from
|
||
<collectingRegion id="872E379D147ED7780DB93A77F32DC1CA" country="Thailand" name="Sa Kaeo">Sa Kaeo Province</collectingRegion>
|
||
in
|
||
<collectingCountry id="BC0864056F78E04F2CA7FA37CE28D563" name="Thailand">Thailand</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="081E043F2DDD9D9E9DA3EB575D283CC6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shells of Songkhlanaia species A, B topotype of S. tamodienica from Tamot Stream, Songkhla Lake Basin, Thailand A adult specimen MUMNH - UNI 2956 and B young specimen MUMNH - UNI 2971 C – F S. moreleti C syntype MNHN - IM- 2000–34623 from Mekong Basin, Cambodia, with original labels D syntype of Pseudodon thomsoni Morlet, 1884, MNHN - IM- 2000-1800 from Mekong Basin, Cambodia, with original labels E adult specimen MUMNH - UNI 0301, and F young specimen MUMNH - UNI 3428 from Phrom Hot Stream, Tonle Sap Basin, Thailand. Photographs C, D M. Caballer (2019, MNHN Project: RECOLNAT No. ANR- 11 - INBS- 0004)." figureDoi="10.3897/zookeys.1224.140549.figure2" httpUri="https://binary.pensoft.net/fig/1236494">2 E</figureCitation>
|
||
), the location of the headwaters of the Tonle Sap Basin and the type locality for this species, have been found to match well with the
|
||
<typeStatus id="80D79FD858D3F4338CB3773E85627044" type="syntype">syntypes</typeStatus>
|
||
(Fig.
|
||
<figureCitation id="D47F0AE5849F1B624042E53CF629E5B9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shells of Songkhlanaia species A, B topotype of S. tamodienica from Tamot Stream, Songkhla Lake Basin, Thailand A adult specimen MUMNH - UNI 2956 and B young specimen MUMNH - UNI 2971 C – F S. moreleti C syntype MNHN - IM- 2000–34623 from Mekong Basin, Cambodia, with original labels D syntype of Pseudodon thomsoni Morlet, 1884, MNHN - IM- 2000-1800 from Mekong Basin, Cambodia, with original labels E adult specimen MUMNH - UNI 0301, and F young specimen MUMNH - UNI 3428 from Phrom Hot Stream, Tonle Sap Basin, Thailand. Photographs C, D M. Caballer (2019, MNHN Project: RECOLNAT No. ANR- 11 - INBS- 0004)." figureDoi="10.3897/zookeys.1224.140549.figure2" httpUri="https://binary.pensoft.net/fig/1236494">2 C</figureCitation>
|
||
). Furthermore, these specimens cluster within the phylogenetic position of the
|
||
<taxonomicName id="147E318826DBF22BB8DAD2167D1FD501" class="Bivalvia" family="Unionidae" genus="Songkhlanaia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
|
||
<emphasis id="A570EE8BB06D129E1002FEDBB6BD3FBF" italics="true">Songkhlanaia</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="41CFED1B864B13690E62AF7A84F734BD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood (ML) phylogenetic tree of freshwater mussels within the subfamily Gonideinae based on a combined DNA sequence dataset of COI, 16 S rRNA, and 28 S rRNA genes (1,959 bp). Branch support values are indicated on nodes as bootstrap percentages from ML analysis and Bayesian posterior probabilities from BI analysis, and shown as ML / BI. The scale bar represents the estimated evolutionary distance between taxa. The clade of the genus Songkhlanaia is highlighted in blue, and type specimens are indicated by an asterisk (*). Type species of Pseudodontini genera are marked with two asterisks (**)." figureDoi="10.3897/zookeys.1224.140549.figure1" httpUri="https://binary.pensoft.net/fig/1236493">1</figureCitation>
|
||
). Therefore, we propose transferring this species to the more appropriate genus
|
||
<taxonomicName id="B6C5ED018210FD1F2FE3720455BAC71A" class="Bivalvia" family="Unionidae" genus="Songkhlanaia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
|
||
<emphasis id="1C819C6E78F41EC0396C219DBB52BDEB" italics="true">Songkhlanaia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D5CA51B022FB95A76759BCD300C63529">
|
||
<taxonomicName id="80F7020D74EDD6336DB7A951BB6FD762" authority="Morlet, 1884" authorityName="Morlet" authorityYear="1884" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni">
|
||
<emphasis id="A6DD39F0707929116C637336DA75E05C" italics="true">Pseudodon thomsoni</emphasis>
|
||
<bibRefCitation id="80A746819400C0463B233D8F9331401E" author="Morlet L" journalOrPublisher="Journal de Conchyliologie" pagination="386-403" refId="B46" refString="Morlet L (1884) Description d'espece nouvelles de coquilles, recueillies par M. Pavie, au Cambodge. Journal de Conchyliologie 32: 386 - 403." title="Description d’espèce nouvelles de coquilles, recueillies par M. Pavie, au Cambodge." volume="32" year="1884">Morlet, 1884</bibRefCitation>
|
||
</taxonomicName>
|
||
was described based on specimens collected from
|
||
<collectingCountry id="7A571975346AFEE418CB2D5265EA37E2" name="Cambodia">Cambodia</collectingCountry>
|
||
by Auguste Jean-Marie Pavie. Later,
|
||
<bibRefCitation id="5EB28A73B0F708E75642CC68192AB21E" author="Morlet L" firstAuthor="Morlet" journalOrPublisher="Journal de Conchyliologie" page="166" pagination="121-199" refId="B47" refString="Morlet L (1889) Catalogue des coquilles recueillies, par M. Pavie, dans le Cambodge et le Royaume de Siam, et description ďespèces nouvelles. Journal de Conchyliologie 37: 121–199." title="Catalogue des coquilles recueillies, par M. Pavie, dans le Cambodge et le Royaume de Siam, et description ďespèces nouvelles." volume="37" year="1889">Morlet (1889: 166)</bibRefCitation>
|
||
detailed and specified the type locality as " Etang de Pnom-Penh (
|
||
<collectingCountry id="1F9DCFC25EEF7A9156A5C26267C8B93D" name="Cambodia">Cambodge</collectingCountry>
|
||
) " [= pond in Phnom Penh,
|
||
<collectingCountry id="47A08B902BB1710FF5EF4E62861FCDE6" name="Cambodia">Cambodia</collectingCountry>
|
||
]. It was recognized as a distinct species for more than a century, until it was recently treated as a junior synonym of
|
||
<taxonomicName id="40184007AB26A3BFABE8172AA7E66E19" authority="(von Martens, 1860)" baseAuthorityName="von Martens" baseAuthorityYear="1860" class="Bivalvia" family="Unionidae" genus="Thaiconcha" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="callifera">
|
||
<emphasis id="27AF8568329BEB9D9FA2808370A0ADC1" italics="true">Thaiconcha callifera</emphasis>
|
||
(
|
||
<bibRefCitation id="CD2A1393A3F62C6802993E4541E4C4BF" author="von Martens EV" journalOrPublisher="Proceedings of the Zoological Society of London" pagination="6-18" refId="B61" refString="von Martens EV (1860) On the mollusca of Siam. Proceedings of the Zoological Society of London 1860: 6 - 18." title="On the mollusca of Siam." volume="1860" year="1860">von Martens, 1860</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by some authors (
|
||
<bibRefCitation id="48DF8FD8B7C940A17C474E3EAEB608B3" DOI="10.1038/s41598-020-63612-5" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Lopes-Lima M & Bogan AE & Lunn Z & Chan N & Win T & Aksenova OV & Tomilova AA & Tanmuangpak K & Tumpeesuwan S & Kondakov AV" etAl="et al." firstAuthor="Bolotov" page="10" refId="B6" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Lopes-Lima M, Bogan AE, Lunn Z, Chan N, Win T, Aksenova OV, Tomilova AA, Tanmuangpak K, Tumpeesuwan S, Kondakov AV (2020) New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10: 6616. https://doi.org/10.1038/s41598-020-63612-5" year="2020">Bolotov et al. 2020: 10</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DEA1C41F2F5733C9F34DA4C2CAFB969E" DOI="10.1093/mollus/eyaa034" author="Graf DL & Cummings KS" firstAuthor="Graf" page="22" refId="B20" refString="Graf DL, Cummings KS (2021) A ‘ big data’ approach to global freshwater mussel diversity (Bivalvia: Unionoida), with an updated checklist of genera and species. Journal of Molluscan Studies 87: eyaa 034. https://doi.org/10.1093/mollus/eyaa034" year="2021">Graf and Cummings 2021: 22</bibRefCitation>
|
||
). The following year, it was resurrected as a valid species by
|
||
<bibRefCitation id="0B57C8CDC1694FE2A3EF5B3F525A7FC9" DOI="10.1038/s41598-021-87633-w" author="Konopleva ES & Bolotov IN & Pfeiffer JM & Vikhrev IV & Kondakov AV & Gofarov MY & Tomilova AA & Tanmuangpak K & Tumpeesuwan S" etAl="et al." firstAuthor="Konopleva" refId="B37" refString="Konopleva ES, Bolotov IN, Pfeiffer JM, Vikhrev IV, Kondakov AV, Gofarov MY, Tomilova AA, Tanmuangpak K, Tumpeesuwan S (2021) New freshwater mussels from two Southeast Asian genera Bineurus and Thaiconcha (Pseudodontini, Gonideinae, Unionidae). Scientific Reports 11: 8244. https://doi.org/10.1038/s41598-021-87633-w" year="2021">Konopleva et al. (2021)</bibRefCitation>
|
||
and again by
|
||
<bibRefCitation id="8E7B9CB26CDE009EF84DD9F5BD5638C6" DOI="10.3390/w15173117" author="Bolotov IN & Konopleva ES & Vikhrev IV & Gofarov MY & Kondakov AV & Lyubas AA & Soboleva AA & Chan N & Lunn Z & Win T & Inkhavilay K" etAl="et al." firstAuthor="Bolotov" refId="B8" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Kondakov AV, Lyubas AA, Soboleva AA, Chan N, Lunn Z, Win T, Inkhavilay K (2023) Integrative taxonomic reappraisal and evolutionary biogeography of the most diverse freshwater mussel clade from Southeast Asia (Pseudodontini). Water 15: 3117. https://doi.org/10.3390/w15173117" year="2023">Bolotov et al. (2023)</bibRefCitation>
|
||
. However, the
|
||
<typeStatus id="97081229DEF04405083593E6B1CC5463" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="C961A768C56E62F356E7B27D96169AB8" authority="Morlet, 1884" authorityName="Morlet" authorityYear="1884" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="thomsoni">
|
||
<emphasis id="1C689D925504244659AF2085644FBD72" italics="true">Pseudodon thomsoni</emphasis>
|
||
<bibRefCitation id="C3218A5BAFCCA89FDF80EA709CB108BD" author="Morlet L" journalOrPublisher="Journal de Conchyliologie" pagination="386-403" refId="B46" refString="Morlet L (1884) Description d'espece nouvelles de coquilles, recueillies par M. Pavie, au Cambodge. Journal de Conchyliologie 32: 386 - 403." title="Description d’espèce nouvelles de coquilles, recueillies par M. Pavie, au Cambodge." volume="32" year="1884">Morlet, 1884</bibRefCitation>
|
||
</taxonomicName>
|
||
is relatively small (Fig.
|
||
<figureCitation id="CC9DACF71728F781AF85A5A8D123B709" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shells of Songkhlanaia species A, B topotype of S. tamodienica from Tamot Stream, Songkhla Lake Basin, Thailand A adult specimen MUMNH - UNI 2956 and B young specimen MUMNH - UNI 2971 C – F S. moreleti C syntype MNHN - IM- 2000–34623 from Mekong Basin, Cambodia, with original labels D syntype of Pseudodon thomsoni Morlet, 1884, MNHN - IM- 2000-1800 from Mekong Basin, Cambodia, with original labels E adult specimen MUMNH - UNI 0301, and F young specimen MUMNH - UNI 3428 from Phrom Hot Stream, Tonle Sap Basin, Thailand. Photographs C, D M. Caballer (2019, MNHN Project: RECOLNAT No. ANR- 11 - INBS- 0004)." figureDoi="10.3897/zookeys.1224.140549.figure2" httpUri="https://binary.pensoft.net/fig/1236494">2 D</figureCitation>
|
||
; shell length
|
||
<quantity id="EFD7A945E25C5527ACE2EFB5117BEB8D" metricMagnitude="-2" metricUnit="m" metricValue="5.3" unit="mm" value="53.0">53 mm</quantity>
|
||
), and its shell characteristics generally resemble those of young specimens of
|
||
<taxonomicName id="E83EF5E8F3FD07D0FA321A3C07CCD456" authorityName="Crosse & Fischer" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="A8E98FC67CCEFD16E823AA91F081B388" italics="true">S. moreleti</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="84B6ABC8F7452F2340C376B18928FC46" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shells of Songkhlanaia species A, B topotype of S. tamodienica from Tamot Stream, Songkhla Lake Basin, Thailand A adult specimen MUMNH - UNI 2956 and B young specimen MUMNH - UNI 2971 C – F S. moreleti C syntype MNHN - IM- 2000–34623 from Mekong Basin, Cambodia, with original labels D syntype of Pseudodon thomsoni Morlet, 1884, MNHN - IM- 2000-1800 from Mekong Basin, Cambodia, with original labels E adult specimen MUMNH - UNI 0301, and F young specimen MUMNH - UNI 3428 from Phrom Hot Stream, Tonle Sap Basin, Thailand. Photographs C, D M. Caballer (2019, MNHN Project: RECOLNAT No. ANR- 11 - INBS- 0004)." figureDoi="10.3897/zookeys.1224.140549.figure2" httpUri="https://binary.pensoft.net/fig/1236494">2 F</figureCitation>
|
||
), such as the long obovate shell that is constricted anteriorly, moderately elevated umbo, and a bean-shaped anterior protractor scar. Furthermore, the type locality in ‘ Phnom Penh, Cambodia’ is in the lower Mekong Basin (Fig.
|
||
<figureCitation id="F317D204019C823B6A7E15640DB18192" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B map showing the geographical distribution of all known Songkhlanaia species (circles) and their type locality (stars). Map was generated using QGIS v. 3.36. 0 with the outline of major river basins from the Freshwater Ecoregions of the World (Abell et al. 2008), river and lake topology from the HydroSHEDS database (https://www.hydrosheds.org), and the map raster data from the NASA EARTHDATA (https://www.earthdata.nasa.gov/)." figureDoi="10.3897/zookeys.1224.140549.figure3" httpUri="https://binary.pensoft.net/fig/1236495">3</figureCitation>
|
||
), the same basin as the type locality of
|
||
<taxonomicName id="68F282445911E3557308C3A0644A532B" authorityName="Crosse & Fischer" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="E125670AC07E189C900234397CAAEC8C" italics="true">S. moreleti</emphasis>
|
||
</taxonomicName>
|
||
. Based on this conchological and biogeographic evidence, we thus synonymise this species with
|
||
<taxonomicName id="61C10DD9BA61978A1F39CE2A11A15187" authorityName="Crosse & Fischer" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="635B919FF56D4B33C56F9A756F673471" italics="true">S. moreleti</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="349441D259ED980B299D9E457B81AA1A">
|
||
The molecular data examined in this study included individuals of
|
||
<taxonomicName id="840257578BBA47BC43002C63D3D1A5F6" authorityName="Crosse & Fischer" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="A7140701F01A917E05C895660DC00B22" italics="true">S. moreleti</emphasis>
|
||
</taxonomicName>
|
||
solely collected from the headwater areas of the Tonle Sap Basin in
|
||
<collectingCountry id="2AD8597B89AF45AD90DE38BEDBD5B5C1" name="Thailand">Thailand</collectingCountry>
|
||
. In fact, previous freshwater mollusk surveys of areas surrounding the Tonle Sap Lake in
|
||
<collectingCountry id="72804ACDF2CF342EC07049A1A5D37646" name="Cambodia">Cambodia</collectingCountry>
|
||
by
|
||
<bibRefCitation id="CED8F6B7DD74B48924463AFA5B5EF2F7" DOI="10.3897/zookeys.958.53865" author="Ng TH & Jeratthitikul E & Sutcharit C & Chhuoy S & Pin K & Pholyotha A & Siriwut W & Srisonchai R & Hogan ZS & Ngor PB" etAl="et al." firstAuthor="Ng" journalOrPublisher="ZooKeys" pagination="107-141" refId="B49" refString="Ng TH, Jeratthitikul E, Sutcharit C, Chhuoy S, Pin K, Pholyotha A, Siriwut W, Srisonchai R, Hogan ZS, Ngor PB (2020) Annotated checklist of freshwater molluscs from the largest freshwater lake in Southeast Asia. ZooKeys 958: 107–141. https://doi.org/10.3897/zookeys.958.53865" title="Annotated checklist of freshwater molluscs from the largest freshwater lake in Southeast Asia." volume="958" year="2020">Ng et al. (2020)</bibRefCitation>
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did not recover any specimens identified as
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<taxonomicName id="8ADFCD1ECF390C87FD808910C81E43EF" authorityName="Crosse & Fischer" authorityYear="1876" class="Bivalvia" family="Unionidae" genus="Pseudodon" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="moreleti">
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<emphasis id="944018CDEF0DD8DA238346918B3224B5" italics="true">S. moreleti</emphasis>
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</taxonomicName>
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from over 40 sampling localities. This possibly suggests a low abundance or local disappearance from the area. Further intensive surveys throughout the basin, including the headwater area and its tributaries, may encounter more specimens, which would be beneficial for assessing the genetic viability and conservation status of this species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |