204 lines
29 KiB
XML
204 lines
29 KiB
XML
<document id="05A9CC981CE947DF66187374485DCCF0" ID-DOI="10.26107/RBZ-2019-0050" ID-ISSN="2345-7600" ID-Zenodo-Dep="5343848" ID-ZooBank="45C7D1BA-73D7-4867-A44D-E375BB0790B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630197826329" checkinUser="diego" docAuthor="Páll-Gergely, Barna & Hunyadi, András" docDate="2019" docId="9B2CD72FD90EFF86D776B7313819FE81" docLanguage="en" docName="RBZ-2019-0050.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Entadella kilchomani Pall-Gergely 2019, new species" docType="treatment" docVersion="4" lastPageNumber="700" masterDocId="6715AF57D90AFF80D428B3573C43FF94" masterDocTitle="Two new species of Entadella Páll-Gergely & Hunyadi in Páll-Gergely et al., 2016 from Vietnam (Gastropoda: Heterobranchia: Camaenidae)" masterLastPageNumber="700" masterPageNumber="694" pageNumber="698" updateTime="1699183192238" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="49759BBFD0AEEA1B03060BCE1BAC27E1">
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<mods:title id="1FB857282E0A136FEBFB2A641F9C245E">Two new species of Entadella Páll-Gergely & Hunyadi in Páll-Gergely et al., 2016 from Vietnam (Gastropoda: Heterobranchia: Camaenidae)</mods:title>
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<mods:namePart id="FB7B08145BBBDB5E691099732D561CBF">Páll-Gergely, Barna</mods:namePart>
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<mods:roleTerm id="255BFB79213F1FB1D56C952121B3E8DF">Author</mods:roleTerm>
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<mods:namePart id="2D12C4AC13F0CB346A77739A59E92E43">Hunyadi, András</mods:namePart>
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<mods:typeOfResource id="BFF88BA220504129D5E246A360132C48">text</mods:typeOfResource>
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<mods:title id="A3F70174845A60AF9B52455ACF6B4A57">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="6B88CB0274A8A46B979EAE43855DFAB3">
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<mods:date id="F444337684DE79CCF09F4875DF3B4E73">2019</mods:date>
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<mods:number id="80F2DA437EA285C52169AD08EE0A7FBC">2019-12-04</mods:number>
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<mods:number id="E6C1388D575005A134347A8E084C9B4A">67</mods:number>
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<mods:classification id="38192A3A0160BFF6ED02642794C98A1F">journal article</mods:classification>
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<mods:identifier id="A6984F4E21D5953681F88FE0FC330E7D" type="DOI">10.26107/RBZ-2019-0050</mods:identifier>
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<mods:identifier id="F78DE9A00D939F37EC3B15159FAD0C5B" type="ISSN">2345-7600</mods:identifier>
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<treatment id="9B2CD72FD90EFF86D776B7313819FE81" ID-DOI="http://doi.org/10.5281/zenodo.5448895" ID-Zenodo-Dep="5448895" LSID="urn:lsid:plazi:treatment:9B2CD72FD90EFF86D776B7313819FE81" httpUri="http://treatment.plazi.org/id/9B2CD72FD90EFF86D776B7313819FE81" lastPageId="6" lastPageNumber="700" pageId="4" pageNumber="698">
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<subSubSection id="5B9F35B2D90EFF84D776B7313938FBEB" box="[862,1403,1126,1151]" pageId="4" pageNumber="698" type="nomenclature">
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<paragraph id="133A6639D90EFF84D776B7313938FBEB" blockId="4.[862,1403,1126,1183]" box="[862,1403,1126,1151]" pageId="4" pageNumber="698">
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<heading id="4872D155D90EFF84D776B7313938FBEB" bold="true" box="[862,1403,1126,1151]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="698" reason="2">
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<emphasis id="21F1BA2BD90EFF84D776B7313938FBEB" bold="true" box="[862,1403,1126,1151]" pageId="4" pageNumber="698">
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<taxonomicName id="D4851DBAD90EFF84D776B73138A4FBEB" ID-CoL="7R7H8" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[862,1255,1126,1151]" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="698" phylum="Mollusca" rank="species" species="kilchomani" status="sp. nov.">
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<emphasis id="21F1BA2BD90EFF84D776B731380EFBEA" bold="true" box="[862,1101,1126,1150]" italics="true" pageId="4" pageNumber="698">Entadella kilchomani</emphasis>
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Páll-Gergely
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</taxonomicName>
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,
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<taxonomicNameLabel id="3AC20750D90EFF84D0DDB7303938FBEB" box="[1269,1403,1127,1151]" pageId="4" pageNumber="698" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5B9F35B2D90EFF84D022B7D0388DFB0B" box="[1034,1230,1159,1183]" pageId="4" pageNumber="698" type="description">
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<paragraph id="133A6639D90EFF84D022B7D0388DFB0B" blockId="4.[862,1403,1126,1183]" box="[1034,1230,1159,1183]" pageId="4" pageNumber="698">
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(
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<figureCitation id="8BBE7ABCD90EFF84D03AB7D038CDFB0B" box="[1042,1166,1159,1183]" captionStart="Fig" captionStartId="1.[136,171,1950,1972]" captionTargetBox="[136,1476,218,1935]" captionTargetId="figure-7@1.[136,1476,218,1935]" captionTargetPageId="1" captionText="Fig. 1. Shells of Entadella Páll-Gergely & Hunyadi in Páll-Gergely et al., 2016, species. A, B, lectotype of Entadella athrix (Möllendorff, 1901) (SMF 8528); B, sculpture of the ventral side, not to scale; C–F, holotype of Entadella kilchomani Páll-Gergely, new species (NHMUK 20190591); D, protoconch; E, dorsal surface; F, ventral surface. Scale “a” represents 3 mm and refers to figs. D, E, and F; scale “b” represents 30 mm and refers to figs. A and C. Note that the close-up pictures were taken using different illumination; original colours are closer to the ones on the entire shell." figureDoi="http://doi.org/10.5281/zenodo.5343850" httpUri="https://zenodo.org/record/5343850/files/figure.png" pageId="4" pageNumber="698">Figs. 1C–F</figureCitation>
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,
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<figureCitation id="8BBE7ABCD90EFF84D0B3B7D038E9FB0B" box="[1179,1194,1159,1183]" captionStart="Fig" captionStartId="4.[114,149,992,1014]" captionTargetBox="[117,1450,190,978]" captionTargetId="figure-460@4.[101,1467,177,988]" captionTargetPageId="4" captionText="Fig. 4. Genitalia of Entadella kilchomani Páll-Gergely, new species. Abbreviations: bc, bursa copulatrix; e, epiphallus; f1, pointed part of penial caecum; f2, blunt part of penial caecum; go, genital opening; p, penis; rm, retractor muscle; so, spermoviduct; v, vagina; vd, vas deferens. Scale bar represents 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5343858" httpUri="https://zenodo.org/record/5343858/files/figure.png" pageId="4" pageNumber="698">4</figureCitation>
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,
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<figureCitation id="8BBE7ABCD90EFF84D090B7D03885FB0B" box="[1208,1222,1159,1183]" captionStart="Fig" captionStartId="5.[136,171,1911,1933]" captionTargetBox="[136,1474,311,1896]" captionTargetId="figure-73@5.[136,1474,311,1896]" captionTargetPageId="5" captionText="Fig. 5. Inner structure of genital organs of Entadella kilchomani Páll-Gergely, new species. A, penis, epiphallus and penial caecum (showed after opening the penial verge); B, intact penial verge; C, vagina. Abbreviations: bs, stalk of bursa copulatrix; e, epiphallus; f, flagellum; fp, flagellar papilla; f1, pointed part of flagellum; f2, blunt part of flagellum; p, penis; pv, penial verge; rm, retractor muscle; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5343862" httpUri="https://zenodo.org/record/5343862/files/figure.png" pageId="4" pageNumber="698">5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="5B9F35B2D90EFF84D705B7903963FAEB" pageId="4" pageNumber="698" type="materials_examined">
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<paragraph id="133A6639D90EFF84D705B7903963FAEB" blockId="4.[811,1452,1223,1407]" pageId="4" pageNumber="698">
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<emphasis id="21F1BA2BD90EFF84D705B7903F90FB4B" bold="true" box="[813,979,1223,1247]" pageId="4" pageNumber="698">Type material.</emphasis>
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<materialsCitation id="A3ED6C64D90EFF84D7F0B7903963FAEB" collectingDate="2013-09-18" collectionCode="V" country="Vietnam" county="Expedition" latitude="21.659483" location="Huu Lien Natural" longLatPrecision="1" longitude="106.36535" municipality="Site" pageId="4" pageNumber="698" specimenCode="NHMUK 20190591, NHMUK 20140589" specimenCount="2" typeStatus="holotype">
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<typeStatus id="CC3ED89BD90EFF84D7F0B7903878FB4B" box="[984,1083,1223,1247]" pageId="4" pageNumber="698" type="holotype">Holotype</typeStatus>
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(1 shell, D:
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<quantity id="D47DCBDCD90EFF84D093B7903958FB4A" box="[1211,1307,1223,1247]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" pageId="4" pageNumber="698" unit="mm" value="34.9">34.9 mm</quantity>
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, H:
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<quantity id="D47DCBDCD90EFF84D16DB79039E7FB4B" box="[1349,1444,1223,1247]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="4" pageNumber="698" unit="mm" value="19.3">19.3 mm</quantity>
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) (
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<specimenCode id="4323CE42D90EFF84D71AB7B03853FB6B" box="[818,1040,1255,1279]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="698" type="Museum">NHMUK 20190591</specimenCode>
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),
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<collectingCountry id="6B9226A9D90EFF84D00AB7B038C2FB6B" box="[1058,1153,1255,1279]" name="Vietnam" pageId="4" pageNumber="698">Vietnam</collectingCountry>
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,
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<location id="165A30E2D90EFF84D0A4B7B03909FB6B" LSID="urn:lsid:plazi:treatment:9B2CD72FD90EFF86D776B7313819FE81:165A30E2D90EFF84D0A4B7B03909FB6B" box="[1164,1354,1255,1279]" country="Vietnam" county="Expedition" latitude="21.659483" longLatPrecision="1" longitude="106.36535" municipality="Site" name="Huu Lien Natural" pageId="4" pageNumber="698">Huu Lien Natural</location>
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RF,
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<collectingMunicipality id="F35EFC43D90EFF84D1A9B7B039E9FB6B" box="[1409,1450,1255,1279]" pageId="4" pageNumber="698">Site</collectingMunicipality>
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||
32, coll. no. 408
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||
<collectionCode id="7594FEFCD90EFF84D7C9B6503FB6FA8B" box="[993,1013,1287,1311]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="698" type="Museum">V</collectionCode>
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||
,
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<geoCoordinate id="76B100FED90EFF84D028B65238C9FA8A" box="[1024,1162,1285,1310]" degrees="21" direction="north" minutes="39.569" orientation="latitude" pageId="4" pageNumber="698" precision="1" value="21.659483">21°39.569′N</geoCoordinate>
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,
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||
<geoCoordinate id="76B100FED90EFF84D0BFB6523968FA8A" box="[1175,1323,1285,1310]" degrees="106" direction="east" minutes="21.921" orientation="longitude" pageId="4" pageNumber="698" precision="1" value="106.36535">106°21.921′E</geoCoordinate>
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, coll. NHM 2013
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<collectingCountry id="6B9226A9D90EFF84D744B6703F8BFAAB" box="[876,968,1319,1343]" name="Vietnam" pageId="4" pageNumber="698">Vietnam</collectingCountry>
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<collectingCounty id="FA5B1EB5D90EFF84D7F9B670380AFAAB" box="[977,1097,1319,1343]" pageId="4" pageNumber="698">Expedition</collectingCounty>
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,
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<date id="673B40F9D90EFF84D07EB6703973FAAB" box="[1110,1328,1319,1343]" pageId="4" pageNumber="698" value="2013-09-18">
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||
<collectingDate id="777FB911D90EFF84D07EB6703973FAAB" box="[1110,1328,1319,1343]" pageId="4" pageNumber="698" value="2013-09-18">18 September 2013</collectingDate>
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</date>
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||
; 1 ethanolpreserved
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<typeStatus id="CC3ED89BD90EFF84D7B4B6103FBBFACA" box="[924,1016,1351,1374]" pageId="4" pageNumber="698" type="paratype">paratype</typeStatus>
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(broken shell, body, prepared genitalia) (
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<specimenCode id="4323CE42D90EFF84D71DB6303850FAEB" box="[821,1043,1383,1407]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="698" type="Museum">NHMUK 20140589</specimenCode>
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), same data as holotype.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5B9F35B2D90EFF86D705B6F03DF6FB21" lastPageId="6" lastPageNumber="700" pageId="4" pageNumber="698" type="description">
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<paragraph id="133A6639D90EFF86D705B6F03EB2FF61" blockId="4.[811,1453,1447,2111]" lastBlockId="6.[113,753,189,245]" lastPageId="6" lastPageNumber="700" pageId="4" pageNumber="698">
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<emphasis id="21F1BA2BD90EFF84D705B6F03FF0FA2B" bold="true" box="[813,947,1447,1471]" pageId="4" pageNumber="698">Description.</emphasis>
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Shell large, dextral, thick-walled, light chocolate brown in colour (
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<figureCitation id="8BBE7ABCD90EFF84D7DAB6903808FA4B" box="[1010,1099,1479,1503]" captionStart="Fig" captionStartId="1.[136,171,1950,1972]" captionTargetBox="[136,1476,218,1935]" captionTargetId="figure-7@1.[136,1476,218,1935]" captionTargetPageId="1" captionText="Fig. 1. Shells of Entadella Páll-Gergely & Hunyadi in Páll-Gergely et al., 2016, species. A, B, lectotype of Entadella athrix (Möllendorff, 1901) (SMF 8528); B, sculpture of the ventral side, not to scale; C–F, holotype of Entadella kilchomani Páll-Gergely, new species (NHMUK 20190591); D, protoconch; E, dorsal surface; F, ventral surface. Scale “a” represents 3 mm and refers to figs. D, E, and F; scale “b” represents 30 mm and refers to figs. A and C. Note that the close-up pictures were taken using different illumination; original colours are closer to the ones on the entire shell." figureDoi="http://doi.org/10.5281/zenodo.5343850" httpUri="https://zenodo.org/record/5343850/files/figure.png" pageId="4" pageNumber="698">Fig. 1C</figureCitation>
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). Shell rather flat with slightly elevated spire. Dorsal surface domed. Shell outline rounded from dorsal view, whorls grow regularly. Body whorl rounded, somewhat shouldered due to slight depression from ventrolateral direction. Whorls 6.75–7, convex, separated by moderately deep suture. Protoconch consisting of almost 2 whorls, smooth, with some slight indication of irregular growth lines (
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<figureCitation id="8BBE7ABCD90EFF84D7E9B5F03855F92B" box="[961,1046,1703,1727]" captionStart="Fig" captionStartId="1.[136,171,1950,1972]" captionTargetBox="[136,1476,218,1935]" captionTargetId="figure-7@1.[136,1476,218,1935]" captionTargetPageId="1" captionText="Fig. 1. Shells of Entadella Páll-Gergely & Hunyadi in Páll-Gergely et al., 2016, species. A, B, lectotype of Entadella athrix (Möllendorff, 1901) (SMF 8528); B, sculpture of the ventral side, not to scale; C–F, holotype of Entadella kilchomani Páll-Gergely, new species (NHMUK 20190591); D, protoconch; E, dorsal surface; F, ventral surface. Scale “a” represents 3 mm and refers to figs. D, E, and F; scale “b” represents 30 mm and refers to figs. A and C. Note that the close-up pictures were taken using different illumination; original colours are closer to the ones on the entire shell." figureDoi="http://doi.org/10.5281/zenodo.5343850" httpUri="https://zenodo.org/record/5343850/files/figure.png" pageId="4" pageNumber="698">Fig. 1D</figureCitation>
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). Teleoconch sculpture dominated by low, irregular growth wrinkles and slight spiral striation that appears on the dorsal side after 3.5 whorls (
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<figureCitation id="8BBE7ABCD90EFF84D129B5B03933F96B" box="[1281,1392,1767,1791]" captionStart="Fig" captionStartId="1.[136,171,1950,1972]" captionTargetBox="[136,1476,218,1935]" captionTargetId="figure-7@1.[136,1476,218,1935]" captionTargetPageId="1" captionText="Fig. 1. Shells of Entadella Páll-Gergely & Hunyadi in Páll-Gergely et al., 2016, species. A, B, lectotype of Entadella athrix (Möllendorff, 1901) (SMF 8528); B, sculpture of the ventral side, not to scale; C–F, holotype of Entadella kilchomani Páll-Gergely, new species (NHMUK 20190591); D, protoconch; E, dorsal surface; F, ventral surface. Scale “a” represents 3 mm and refers to figs. D, E, and F; scale “b” represents 30 mm and refers to figs. A and C. Note that the close-up pictures were taken using different illumination; original colours are closer to the ones on the entire shell." figureDoi="http://doi.org/10.5281/zenodo.5343850" httpUri="https://zenodo.org/record/5343850/files/figure.png" pageId="4" pageNumber="698">Fig. 1E, F</figureCitation>
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), and remains equally strong until peristome. Spiral striae much denser on ventral than on dorsal side, radial growth wrinkles lower on ventral than on dorsal side. Umbilicus open but very narrow, point-like, shows all whorls. Its edge covered by reflected peristome. Aperture semilunar, slightly elongated in direction of umbilicus, slightly oblique to the shell axis from lateral view. Peristome similar to shell in colour, strongly thickened and reflected, with a slight thickening on basal part of the
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<typeStatus id="CC3ED89BD90EFF84D7C9BB50387DF78B" box="[993,1086,2055,2079]" pageId="4" pageNumber="698" type="holotype">holotype</typeStatus>
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(not visible on
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<typeStatus id="CC3ED89BD90EFF84D0CFBB503905F78B" box="[1255,1350,2055,2079]" pageId="4" pageNumber="698" type="paratype">paratype</typeStatus>
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, which is seemingly a less matured shell). Parietal callus present only as transparent, fine calcium layer on the parietal shell wall, its surface is finely granulate, less glossy than rest of shell.
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</paragraph>
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<caption id="47FA36B1D90FFF85D4A0B4203980F853" ID-DOI="http://doi.org/10.5281/zenodo.5343862" ID-Zenodo-Dep="5343862" httpUri="https://zenodo.org/record/5343862/files/figure.png" pageId="5" pageNumber="699" startId="5.[136,171,1911,1933]" targetBox="[136,1474,311,1896]" targetPageId="5">
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||
<paragraph id="133A6639D90FFF85D4A0B4203980F853" blockId="5.[135,1475,1911,1991]" pageId="5" pageNumber="699">
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||
Fig. 5. Inner structure of genital organs of
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<taxonomicName id="D4851DBAD90FFF85D635B4203F2EF819" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[541,877,1911,1933]" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="699" phylum="Mollusca" rank="species" species="kilchomani" status="sp. nov.">
|
||
<emphasis id="21F1BA2BD90FFF85D635B4203EA8F818" box="[541,747,1911,1932]" italics="true" pageId="5" pageNumber="699">Entadella kilchomani</emphasis>
|
||
Páll-Gergely
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="3AC20750D90FFF85D75FB42F3FAFF819" box="[887,1004,1911,1933]" pageId="5" pageNumber="699" rank="species">new species</taxonomicNameLabel>
|
||
. A, penis, epiphallus and penial caecum (showed after opening the penial verge); B, intact penial verge; C, vagina. Abbreviations: bs, stalk of bursa copulatrix; e, epiphallus; f, flagellum; fp, flagellar papilla; f1, pointed part of flagellum; f2, blunt part of flagellum; p, penis; pv, penial verge; rm, retractor muscle; vd, vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="133A6639D90CFF86D459B24A3E9CFEA1" blockId="6.[113,735,285,309]" box="[113,735,285,309]" pageId="6" pageNumber="700">
|
||
<emphasis id="21F1BA2BD90CFF86D459B24A3D63FEA1" bold="true" box="[113,288,285,309]" pageId="6" pageNumber="700">Measurements.</emphasis>
|
||
D =
|
||
<quantity id="D47DCBDCD90CFF86D574B24A3DFCFEA1" box="[348,447,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" pageId="6" pageNumber="700" unit="mm" value="34.9">34.9 mm</quantity>
|
||
, H =
|
||
<quantity id="D47DCBDCD90CFF86D5D7B24A3E23FEA1" box="[511,608,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="6" pageNumber="700" unit="mm" value="19.3">19.3 mm</quantity>
|
||
(
|
||
<typeStatus id="CC3ED89BD90CFF86D658B24A3E97FEA1" box="[624,724,285,309]" pageId="6" pageNumber="700" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="133A6639D90CFF86D459B20A3DF6FB21" blockId="6.[112,754,349,1205]" pageId="6" pageNumber="700">
|
||
<emphasis id="21F1BA2BD90CFF86D459B20A3DE5FEE1" bold="true" box="[113,422,349,373]" pageId="6" pageNumber="700">Description of the anatomy.</emphasis>
|
||
<figureCitation id="8BBE7ABCD90CFF86D581B20A3DB1FEE1" box="[425,498,349,373]" captionStart="Fig" captionStartId="4.[114,149,992,1014]" captionTargetBox="[117,1450,190,978]" captionTargetId="figure-460@4.[101,1467,177,988]" captionTargetPageId="4" captionText="Fig. 4. Genitalia of Entadella kilchomani Páll-Gergely, new species. Abbreviations: bc, bursa copulatrix; e, epiphallus; f1, pointed part of penial caecum; f2, blunt part of penial caecum; go, genital opening; p, penis; rm, retractor muscle; so, spermoviduct; v, vagina; vd, vas deferens. Scale bar represents 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5343858" httpUri="https://zenodo.org/record/5343858/files/figure.png" pageId="6" pageNumber="700">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="8BBE7ABCD90CFF86D5D4B20A3E4FFEE1" box="[508,524,349,373]" captionStart="Fig" captionStartId="5.[136,171,1911,1933]" captionTargetBox="[136,1474,311,1896]" captionTargetId="figure-73@5.[136,1474,311,1896]" captionTargetPageId="5" captionText="Fig. 5. Inner structure of genital organs of Entadella kilchomani Páll-Gergely, new species. A, penis, epiphallus and penial caecum (showed after opening the penial verge); B, intact penial verge; C, vagina. Abbreviations: bs, stalk of bursa copulatrix; e, epiphallus; f, flagellum; fp, flagellar papilla; f1, pointed part of flagellum; f2, blunt part of flagellum; p, penis; pv, penial verge; rm, retractor muscle; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5343862" httpUri="https://zenodo.org/record/5343862/files/figure.png" pageId="6" pageNumber="700">5</figureCitation>
|
||
. Right ommatophoral retractor crosses penis and vagina. Atrium very short. Penis with a short, slender distal, and a slightly longer, much thicker proximal part. Inner penial wall with irregular, low, longitudinal folds having perpendicular projections resulting in a reticulated surface. Penial verge large, thickened, clubshaped, with a slit-like opening on the lateral side. Epiphallus thick, but thinner than proximal part of penis, slightly shorter than penis, internally with 3–4 elevated, longitudinal folds that are discernible inside penial verge as well. Retractor muscle inserts on distal part of epiphallus.
|
||
<collectingRegion id="D141A8DBD90CFF86D5D1B1C93E61FD21" box="[505,546,670,693]" country="Hungary" name="Vas" pageId="6" pageNumber="700">Vas</collectingRegion>
|
||
deferens normally developed. A thin membrane was found between vas deferens and penis-epiphallus, and vas deferens and distal end of flagellum. Flagellum strongly developed, approximately as long as penis and epiphallus combined, its end divided into a pointed and a blunter tip. Longitudinal folds of epiphallus continue into pointed end of flagellum, whereas the blunt flagellum tip seemingly forms an independent cavity with thick, transversely sculptured wall, and two small flagellar papillae. Vagina short, shorter than penis, internally with ca. 12 longitudinal, wavy folds, which continue inside stalk of bursa copulatrix for some distance (possibly the vagina and the distal end of bursa’s stalk form a single functional unit). The wavy folds abruptly transform into straight folds inside lumen of bursa’s stalk. Stalk of bursa very long, bursa oval, large. Spermoviduct normally developed. Albumen gland and talon were not examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5B9F35B2D90CFF86D459B78A3E02F9A1" pageId="6" pageNumber="700" type="diagnosis">
|
||
<paragraph id="133A6639D90CFF86D459B78A3E02F9A1" blockId="6.[112,754,1245,1589]" pageId="6" pageNumber="700">
|
||
<emphasis id="21F1BA2BD90CFF86D459B78A3D2EFB61" bold="true" box="[113,365,1245,1269]" pageId="6" pageNumber="700">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="D4851DBAD90CFF86D55CB78A3EAEFB61" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[372,749,1245,1269]" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="700" phylum="Mollusca" rank="species" species="kilchomani" status="sp. nov.">
|
||
<emphasis id="21F1BA2BD90CFF86D55CB78A3E19FB61" box="[372,602,1245,1269]" italics="true" pageId="6" pageNumber="700">Entadella kilchomani</emphasis>
|
||
Páll-Gergely
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="3AC20750D90CFF86D459B7A93CB8FA81" box="[113,251,1277,1301]" pageId="6" pageNumber="700" rank="species">new species</taxonomicNameLabel>
|
||
, differs from
|
||
<taxonomicName id="D4851DBAD90CFF86D5B5B7AA3EDDFA81" authorityName="Pall-Gergely & Hunyadi" authorityYear="2016" box="[413,670,1277,1301]" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="700" phylum="Mollusca" rank="species" species="entadiformis">
|
||
<emphasis id="21F1BA2BD90CFF86D5B5B7AA3EDDFA81" box="[413,670,1277,1301]" italics="true" pageId="6" pageNumber="700">Entadella entadiformis</emphasis>
|
||
</taxonomicName>
|
||
by the more strongly built, less depressed shell having regularly shaped whorls, the lack of tubercles on the protoconch, the less elongated aperture, and the much stronger peristome.
|
||
<taxonomicName id="D4851DBAD90CFF86D459B62A3D5CFA01" baseAuthorityName="Mollendorff" baseAuthorityYear="1901" box="[113,287,1405,1429]" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="700" phylum="Mollusca" rank="species" species="athrix">
|
||
<emphasis id="21F1BA2BD90CFF86D459B62A3D5CFA01" box="[113,287,1405,1429]" italics="true" pageId="6" pageNumber="700">Entadella athrix</emphasis>
|
||
</taxonomicName>
|
||
is smaller (D:
|
||
<quantity id="D47DCBDCD90CFF86D595B62A3E19FA01" box="[445,602,1405,1429]" metricMagnitude="-2" metricUnit="m" metricValue="2.565" metricValueMax="2.67" metricValueMin="2.46" pageId="6" pageNumber="700" unit="mm" value="25.65" valueMax="26.7" valueMin="24.6">24.6–26.7 mm</quantity>
|
||
, H:
|
||
<quantity id="D47DCBDCD90CFF86D6ADB62A3CDEFA21" metricMagnitude="-2" metricUnit="m" metricValue="1.4449999999999998" metricValueMax="1.51" metricValueMin="1.38" pageId="6" pageNumber="700" unit="mm" value="14.45" valueMax="15.1" valueMin="13.8">13.8–15.1 mm</quantity>
|
||
, n = 4 [MNHN 2012-27112, MNHN 2012-27125]), has a wider umbilicus, and a much thinner peristome.
|
||
<taxonomicName id="D4851DBAD90CFF86D6A0B6EA3C89FA61" authority="Páll-Gergely & Hunyadi, 2019" authorityName="Páll-Gergely & Hunyadi" authorityYear="2019" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="700" phylum="Mollusca" rank="species" species="concava" status="sp. nov.">
|
||
<emphasis id="21F1BA2BD90CFF86D6A0B6EA3C89FA61" italics="true" pageId="6" pageNumber="700">Entadella concava</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="3AC20750D90CFF86D4F0B6893D1FFA61" box="[216,348,1501,1525]" pageId="6" pageNumber="700" rank="species">new species</taxonomicNameLabel>
|
||
, is smaller than
|
||
<taxonomicName id="D4851DBAD90CFF86D623B68A3CBFF981" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="700" phylum="Mollusca" rank="species" species="kilchomani" status="sp. nov.">
|
||
<emphasis id="21F1BA2BD90CFF86D623B68A3EB2FA61" box="[523,753,1501,1525]" italics="true" pageId="6" pageNumber="700">Entadella kilchomani</emphasis>
|
||
Páll-Gergely
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="3AC20750D90CFF86D52CB6A93DC6F981" box="[260,389,1533,1557]" pageId="6" pageNumber="700" rank="species">new species</taxonomicNameLabel>
|
||
, more depressed, and has a slightly wider umbilicus and sunken initial whorls.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5B9F35B2D90CFF86D705B3EA3819FE81" pageId="6" pageNumber="700" type="etymology">
|
||
<paragraph id="133A6639D90CFF86D705B3EA3819FE81" blockId="6.[811,1450,189,277]" pageId="6" pageNumber="700">
|
||
<emphasis id="21F1BA2BD90CFF86D705B3EA3FEEFF41" bold="true" box="[813,941,189,213]" pageId="6" pageNumber="700">Etymology.</emphasis>
|
||
<taxonomicName id="D4851DBAD90CFF86D799B3EA395DFF41" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2019" box="[945,1310,189,213]" class="Gastropoda" family="Camaenidae" genus="Entadella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="700" phylum="Mollusca" rank="species" species="kilchomani" status="sp. nov.">
|
||
<emphasis id="21F1BA2BD90CFF86D799B3EA38D1FF41" box="[945,1170,189,213]" italics="true" pageId="6" pageNumber="700">Entadella kilchomani</emphasis>
|
||
Páll-Gergely
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="3AC20750D90CFF86D10EB3E939E4FF41" box="[1318,1447,189,213]" pageId="6" pageNumber="700" rank="species">new species</taxonomicNameLabel>
|
||
, is named after Kilchoman Distillery, the favourite whisky distillery of the first author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |