308 lines
46 KiB
XML
308 lines
46 KiB
XML
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<mods:title id="60494A837BAFEF367A40077190C424FD">The new Southeast Asian goblin spider genus Aposphragisma (Araneae, Oonopidae): diversity and phylogeny</mods:title>
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<mods:namePart id="788A534308588C866D64EF11A64F895F">THOMA, MARCO</mods:namePart>
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<mods:namePart id="A0ACB1E8F0286EE9F0F5DD0070F73AE2">KRANZ-BALTENSPERGER, YVONNE</mods:namePart>
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<mods:namePart id="D8B17F57186A1D11E71EB1AC43F77A4B">FRICK, HOLGER</mods:namePart>
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<mods:title id="7FE199761BF19D4AECD1EE058C612BA5">Zootaxa</mods:title>
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<mods:date id="E47734FB5EE1C3E0BA5C240D612C9392">2014</mods:date>
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<treatment id="094F426DFFD2FF9A6FC9F770871CFD29" ID-DOI="http://doi.org/10.5281/zenodo.4431947" ID-GBIF-Taxon="176566783" ID-Zenodo-Dep="4431947" LSID="urn:lsid:plazi:treatment:094F426DFFD2FF9A6FC9F770871CFD29" httpUri="http://treatment.plazi.org/id/094F426DFFD2FF9A6FC9F770871CFD29" lastPageId="71" lastPageNumber="72" pageId="68" pageNumber="69">
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<subSubSection id="C9FCA0F0FFD2FF996FC9F770802AFA87" box="[151,637,1296,1323]" pageId="68" pageNumber="69" type="nomenclature">
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<heading id="DA114417FFD2FF996FC9F770802AFA87" bold="true" box="[151,637,1296,1323]" fontSize="11" level="2" pageId="68" pageNumber="69" reason="4">
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<taxonomicName id="46E688F8FFD2FF996FC9F7708045FA87" ID-CoL="FVF3" authority="Thoma" authorityName="Thoma" authorityYear="2014" box="[151,530,1296,1323]" class="Arachnida" family="Oonopidae" genus="Aposphragisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="sepilok" status="sp. nov.">
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<emphasis id="B3922F69FFD2FF996FC9F77083E7FA86" bold="true" box="[151,432,1296,1322]" italics="true" pageId="68" pageNumber="69">Aposphragisma sepilok</emphasis>
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Thoma
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</taxonomicName>
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,
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<taxonomicNameLabel id="A8A19212FFD2FF996D41F771802AFA87" box="[543,637,1297,1323]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C9FCA0F0FFD2FF996FC9F7538379FAE0" box="[151,302,1331,1356]" pageId="68" pageNumber="69" type="description">
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<paragraph id="8159F37BFFD2FF996FC9F7538379FAE0" blockId="68.[151,637,1296,1356]" box="[151,302,1331,1356]" pageId="68" pageNumber="69">
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(
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<figureCitation id="19DDEFFEFFD2FF996FC1F7538371FAE0" box="[159,294,1331,1356]" captionStart-0="FIGURE 46" captionStart-1="FIGURE 47" captionStartId-0="69.[151,250,1871,1893]" captionStartId-1="70.[151,250,1472,1494]" captionTargetBox-0="[178,1416,231,1822]" captionTargetBox-1="[173,1423,218,1409]" captionTargetId-0="figure-87@69.[151,1436,193,1850]" captionTargetId-1="figure-265@70.[151,1436,193,1451]" captionTargetPageId-0="69" captionTargetPageId-1="70" captionText-0="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." captionText-1="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4431999" figureDoi-1="http://doi.org/10.5281/zenodo.4432001" httpUri-0="https://zenodo.org/record/4431999/files/figure.png" httpUri-1="https://zenodo.org/record/4432001/files/figure.png" pageId="68" pageNumber="69">Figs. 46–47</figureCitation>
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)
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</paragraph>
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<subSubSection id="C9FCA0F0FFD2FF996FC9F71B83B9F9C0" pageId="68" pageNumber="69" type="materials_examined">
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<emphasis id="B3922F69FFD2FF996FC9F71B831FFA38" bold="true" box="[151,328,1403,1428]" pageId="68" pageNumber="69">Type material.</emphasis>
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:
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</emphasis>
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<specimenCount id="97E038F2FFD2FF996EB5F71B8076FA38" box="[491,545,1403,1428]" pageId="68" pageNumber="69" type="male">male</specimenCount>
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(
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),
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(
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),
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W of Sandakan
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</locationDeviation>
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,
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<location id="8439A5A0FFD2FF996A13F71B83DCFA1B" LSID="urn:lsid:plazi:treatment:094F426DFFD2FF9A6FC9F770871CFD29:8439A5A0FFD2FF996A13F71B83DCFA1B" country="Malaysia" latitude="5.8333335" longLatPrecision="1305" longitude="117.933334" name="KabiliSepilok forest reserve" pageId="68" pageNumber="69" stateProvince="Sabah">KabiliSepilok forest reserve</location>
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[
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; inf.],
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<collectingDate id="E51C2C53FFD2FF996DEDF7FF8166FA14" box="[691,817,1439,1464]" pageId="68" pageNumber="69" value="1982-04-23">23.IV.1982</collectingDate>
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,
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<collectedFrom id="32FAB086FFD2FF996C62F7FF860CFA70" pageId="68" pageNumber="69">secondary lowland forest, “forêt près du ‘pond’ (étang formant la reserve d’eau pour Sepilok), tamisage de feuilles mortes et de bois pourri”</collectedFrom>
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, leg.
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<collectorName id="2C1396ADFFD2FF996BC9F7A4875DFA77" box="[1175,1290,1476,1499]" pageId="68" pageNumber="69">B. Hauser</collectorName>
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, deposited at
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<collectionCode id="E7F76BBEFFD2FF996FC9F78782A7F9AC" box="[151,240,1511,1536]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="68" pageNumber="69" type="Museum">MHNG</collectionCode>
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;
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</materialsCitation>
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<materialsCitation id="318EF926FFD2FF996E5BF787815CF9E4" ID-GBIF-Occurrence="3014709325" collectedFrom="foret pres de l'Orang-Utan Rehabilitation Station, Eusideroxylon zwangeri et Pometia pinnata" collectingDate="1983-03-12" collectionCode="MHNG" collectorName="B. Hauser" country="Malaysia" latitude="5.8333335" location="Kabili-Sepilok-Forest reserve" longLatPrecision="1305" longitude="117.933334" municipality="Kabili-Sepilok-Forest reserve" pageId="68" pageNumber="69" specimenCode="PBI_OON 00015274" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
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:
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</emphasis>
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(
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),
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<collectingRegion id="43223D99FFD2FF996C14F78781D8F9AC" box="[842,911,1511,1536]" country="Malaysia" name="Sabah" pageId="68" pageNumber="69">Sabah</collectingRegion>
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(
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<collectingCountry id="F9F1B3EBFFD2FF996CF9F7878673FA53" box="[935,1060,1511,1536]" name="Malaysia" pageId="68" pageNumber="69">MALAYSIA</collectingCountry>
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),
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<collectingMunicipality id="613D6901FFD2FF996B1BF78787CBFA53" box="[1093,1436,1511,1536]" country="Malaysia" latitude="5.8333335" longLatPrecision="1305" longitude="117.933334" name="Kabili-Sepilok-Forest reserve" pageId="68" pageNumber="69" stateProvince="Sabah">Kabili-Sepilok-Forest reserve</collectingMunicipality>
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[
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<geoCoordinate id="E4D295BCFFD2FF996FC3F46B82A9F988" box="[157,254,1547,1572]" degrees="05" direction="north" minutes="50" orientation="latitude" pageId="68" pageNumber="69" precision="925" value="5.8333335">05°50′N</geoCoordinate>
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<geoCoordinate id="E4D295BCFFD2FF996E5AF46B8338F988" box="[260,367,1547,1572]" degrees="117" direction="east" minutes="56" orientation="longitude" pageId="68" pageNumber="69" precision="925" value="117.933334">117°56′E</geoCoordinate>
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; inf.],
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<date id="F558D5BBFFD2FF996EE9F46B8060F988" box="[439,567,1547,1572]" pageId="68" pageNumber="69" value="1983-03-12">
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<collectingDate id="E51C2C53FFD2FF996EE9F46B8060F988" box="[439,567,1547,1572]" pageId="68" pageNumber="69" value="1983-03-12">12.III.1983</collectingDate>
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||
</date>
|
||
,
|
||
<collectedFrom id="32FAB086FFD2FF996D1DF46B8304F9EB" pageId="68" pageNumber="69">
|
||
forêt près de l’Orang-Utan Rehabilitation Station,
|
||
<taxonomicName id="46E688F8FFD2FF996B2DF46B87D6F98F" box="[1139,1409,1547,1572]" class="Magnoliopsida" family="Lauraceae" genus="Eusideroxylon" kingdom="Plantae" order="Laurales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="zwangeri">
|
||
<emphasis id="B3922F69FFD2FF996B2DF46B87D6F98F" box="[1139,1409,1547,1572]" italics="true" pageId="68" pageNumber="69">Eusideroxylon zwangeri</emphasis>
|
||
</taxonomicName>
|
||
et
|
||
<taxonomicName id="46E688F8FFD2FF996FC9F4508304F9EB" box="[151,339,1584,1607]" class="Magnoliopsida" family="Sapindaceae" genus="Pometia" kingdom="Plantae" order="Sapindales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="pinnata">
|
||
<emphasis id="B3922F69FFD2FF996FC9F4508304F9EB" box="[151,339,1584,1607]" italics="true" pageId="68" pageNumber="69">Pometia pinnata</emphasis>
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
, leg.
|
||
<collectorName id="2C1396ADFFD2FF996EC9F450805BF9EB" box="[407,524,1584,1607]" pageId="68" pageNumber="69">B. Hauser</collectorName>
|
||
, deposited at
|
||
<collectionCode id="E7F76BBEFFD2FF996DF3F44F8151F9E4" box="[685,774,1583,1608]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="68" pageNumber="69" type="Museum">MHNG</collectionCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="318EF926FFD2FF996C4AF44F83B9F9C0" ID-GBIF-Occurrence="3014709367" collectionCode="MHNG" pageId="68" pageNumber="69" specimenCode="PBI_OON 00031409" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="97E038F2FFD2FF996C4AF44F812FF9E4" box="[788,888,1583,1608]" pageId="68" pageNumber="69" type="female">1 female</specimenCount>
|
||
(
|
||
<specimenCode id="D1405B00FFD2FF996CD4F44F86D6F9E4" box="[906,1153,1583,1608]" pageId="68" pageNumber="69">PBI_OON 00031409</specimenCode>
|
||
), collected together with holotype, deposited at
|
||
<collectionCode id="E7F76BBEFFD2FF996ECAF43383BDF9C0" box="[404,490,1619,1644]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="68" pageNumber="69" type="Museum">MHNG</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9FCA0F0FFD2FF996F99F4178714F923" box="[199,1347,1655,1680]" pageId="68" pageNumber="69" type="etymology">
|
||
<paragraph id="8159F37BFFD2FF996F99F4178714F923" blockId="68.[151,1436,1403,2004]" box="[199,1347,1655,1680]" pageId="68" pageNumber="69">
|
||
<emphasis id="B3922F69FFD2FF996F99F4178319F93C" bold="true" box="[199,334,1655,1680]" pageId="68" pageNumber="69">Etymology.</emphasis>
|
||
The species epithet refers to the
|
||
<typeStatus id="5E5D4DD9FFD2FF996DE0F41880B9F93C" box="[702,750,1656,1680]" pageId="68" pageNumber="69">type</typeStatus>
|
||
locality at Kabili-Sepilok-Forest reserve, on Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9FCA0F0FFD2FF996F99F4FB8327F8E8" pageId="68" pageNumber="69" type="diagnosis">
|
||
<paragraph id="8159F37BFFD2FF996F99F4FB8327F8E8" blockId="68.[151,1436,1403,2004]" pageId="68" pageNumber="69">
|
||
<emphasis id="B3922F69FFD2FF996F99F4FB8316F918" bold="true" box="[199,321,1691,1716]" pageId="68" pageNumber="69">Diagnosis.</emphasis>
|
||
<taxonomicName id="46E688F8FFD2FF996E14F4FC83EEF918" authorityName="Thoma" authorityYear="2014" box="[330,441,1691,1716]" class="Arachnida" family="Oonopidae" genus="Aposphragisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="sepilok" status="sp. nov.">
|
||
<emphasis id="B3922F69FFD2FF996E14F4FC83EEF918" box="[330,441,1691,1716]" italics="true" pageId="68" pageNumber="69">A. sepilok</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A8A19212FFD2FF996E9CF4FC804CF91F" box="[450,539,1692,1715]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the stripe-clade and can be distinguished from other species of this clade by the combination of the following traits: carapace without spikes; carapace margin with blunt denticles, sluice restricted to posterior margin (
|
||
<figureCitation id="19DDEFFEFFD2FF996D67F48380F6F950" box="[569,673,1763,1788]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="68" pageNumber="69">Fig. 46B</figureCitation>
|
||
); epigastric scutum in females posterolaterally with a pair of single curved ridges (cr) (
|
||
<figureCitation id="19DDEFFEFFD2FF996E32F5678386F88C" box="[364,465,1799,1824]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="68" pageNumber="69">Fig. 47E</figureCitation>
|
||
). The granulate surface of the carapace sides and the tiny eyes seperate the new species from all congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9FCA0F0FFD2FF9A6F99F52F83E0FDCC" lastPageId="71" lastPageNumber="72" pageId="68" pageNumber="69" type="description">
|
||
<paragraph id="8159F37BFFD2FF996F99F52F811FF8C4" blockId="68.[151,1436,1403,2004]" box="[199,840,1871,1896]" pageId="68" pageNumber="69">
|
||
<emphasis id="B3922F69FFD2FF996F99F52F830EF8C4" bold="true" box="[199,345,1871,1896]" pageId="68" pageNumber="69">Description.</emphasis>
|
||
Description based on
|
||
<specimenCount id="97E038F2FFD2FF996D0CF52F80CAF8C4" box="[594,669,1871,1896]" pageId="68" pageNumber="69" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="97E038F2FFD2FF996D8AF52F8113F8C4" box="[724,836,1871,1896]" pageId="68" pageNumber="69" type="female">2 females</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8159F37BFFD2FF996F99F51383C0F878" blockId="68.[151,1436,1403,2004]" pageId="68" pageNumber="69">
|
||
<emphasis id="B3922F69FFD2FF996F99F5138341F820" bold="true" box="[199,278,1907,1932]" pageId="68" pageNumber="69">MALE.</emphasis>
|
||
With the characters of the genus except as noted. Body length
|
||
<quantity id="461E5E9EFFD2FF996CA5F5138633F827" box="[1019,1124,1907,1932]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="68" pageNumber="69" unit="mm" value="1.39">1.39 mm</quantity>
|
||
, carapace length
|
||
<quantity id="461E5E9EFFD2FF996A70F51387CFF827" box="[1326,1432,1907,1932]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="68" pageNumber="69" unit="mm" value="0.61">0.61 mm</quantity>
|
||
. Sclerotized parts uniformly coloured light orange-brown, close to colour 38 ‘tawny’; legs slightly paler than body, pale orange-yellowish.
|
||
</paragraph>
|
||
<caption id="D599A3F3FFD3FF986FC9F52F809CF86D" ID-DOI="http://doi.org/10.5281/zenodo.4431999" ID-Zenodo-Dep="4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="69" pageNumber="70" startId="69.[151,250,1871,1893]" targetBox="[178,1416,231,1822]" targetPageId="69">
|
||
<paragraph id="8159F37BFFD3FF986FC9F52F809CF86D" blockId="69.[151,1435,1871,1985]" pageId="69" pageNumber="70">
|
||
<emphasis id="B3922F69FFD3FF986FC9F52F8375F8C8" bold="true" box="[151,290,1871,1893]" pageId="69" pageNumber="70">FIGURE 46.</emphasis>
|
||
<taxonomicName id="46E688F8FFD3FF986E75F52F804DF8C9" authorityName="Thoma" authorityYear="2014" box="[299,538,1871,1893]" class="Arachnida" family="Oonopidae" genus="Aposphragisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="sepilok" status="sp. nov.">
|
||
<emphasis id="B3922F69FFD3FF986E75F52F804DF8C9" box="[299,538,1871,1893]" italics="true" pageId="69" pageNumber="70">Aposphragisma sepilok</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A8A19212FFD3FF986D7DF5308022F8C9" box="[547,629,1872,1893]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D599A3F3FFD0FF9B6FC9F7A0813FF99E" ID-DOI="http://doi.org/10.5281/zenodo.4432001" ID-Zenodo-Dep="4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="70" pageNumber="71" startId="70.[151,250,1472,1494]" targetBox="[173,1423,218,1409]" targetPageId="70">
|
||
<paragraph id="8159F37BFFD0FF9B6FC9F7A0813FF99E" blockId="70.[151,1436,1472,1586]" pageId="70" pageNumber="71">
|
||
<emphasis id="B3922F69FFD0FF9B6FC9F7A08374FA79" bold="true" box="[151,291,1472,1494]" pageId="70" pageNumber="71">FIGURE 47.</emphasis>
|
||
<taxonomicName id="46E688F8FFD0FF9B6E72F7A0804AFA7A" authorityName="Thoma" authorityYear="2014" box="[300,541,1472,1494]" class="Arachnida" family="Oonopidae" genus="Aposphragisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="sepilok" status="sp. nov.">
|
||
<emphasis id="B3922F69FFD0FF9B6E72F7A0804AFA7A" box="[300,541,1472,1494]" italics="true" pageId="70" pageNumber="71">Aposphragisma sepilok</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A8A19212FFD0FF9B6D78F7A1802EFA7A" box="[550,633,1473,1494]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8159F37BFFD0FF9B6F99F43981D7F876" blockId="70.[151,1437,1625,2010]" pageId="70" pageNumber="71">
|
||
CEPHALOTHORAX. Carapace: surface of elevated portion of pars cephalica smooth, sides granulate (
|
||
<figureCitation id="19DDEFFEFFD0FF9B6A78F43982FCF93A" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="70" pageNumber="71">Figs. 46A, E</figureCitation>
|
||
); posterolateral surface without spikes; posterior edge of pars cephalica with slightly stronger modified hair bases, appearing as small denticles; carapace margin with sluice restricted to posterior margin, with blunt denticles (
|
||
<figureCitation id="19DDEFFEFFD0FF9B6FC1F4A58350F972" box="[159,263,1733,1758]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="70" pageNumber="71">Fig. 46A</figureCitation>
|
||
); few non-marginal pars thoracica setae on carapace sides, very scattered. Eyes: tiny, especially PME and PLE (
|
||
<figureCitation id="19DDEFFEFFD0FF9B6F80F489831AF8AE" box="[222,333,1769,1794]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="70" pageNumber="71">Fig. 46A</figureCitation>
|
||
); posterior eye-row straight from above, procurved from front (
|
||
<figureCitation id="19DDEFFEFFD0FF9B6B15F48986BBF8AE" box="[1099,1260,1769,1794]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="70" pageNumber="71">Figs. 46A, G</figureCitation>
|
||
); ALE largest, separated by their radius to diameter; ALE-PLE separated by less than ALE radius; PME separated by their radius to diameter; PLE-PME separated by less than PME radius. Sternum finely ornamented except smooth median stripe which is about half as long as sternum (
|
||
<figureCitation id="19DDEFFEFFD0FF9B6DFBF5358159F8C2" box="[677,782,1877,1902]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="70" pageNumber="71">Fig. 46C</figureCitation>
|
||
); infracoxal grooves present; posterior margin with broad single extension, covered with few blunt denticles (
|
||
<figureCitation id="19DDEFFEFFD0FF9B6DA8F5198135F83E" box="[758,866,1913,1938]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="70" pageNumber="71">Fig. 46C</figureCitation>
|
||
). Pleura: surface smooth with pairs of large pits dorsally of each coxa I, II and III. Mouthparts: chelicerae promargin with row of flattened setae, distally extending into a short inwards pointing tooth-like projection (tlp) (
|
||
<figureCitation id="19DDEFFEFFD0FF9B6C52F5A18124F876" box="[780,883,1985,2010]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="70" pageNumber="71">Fig. 46C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8159F37BFFD1FF9A6F99F2F78021FEB1" blockId="71.[151,1437,151,645]" pageId="71" pageNumber="72">
|
||
ABDOMEN. Book lung covers (bc) large, elliptical to pear-shaped (sides slightly concave), about 3 times longer than wide (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6E42F2DC83DEFF79" box="[284,393,188,213]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="71" pageNumber="72">Fig. 46H</figureCitation>
|
||
). Epigastric scutum dorsally with a widely oval subterminal ridge (sr) (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6B8DF2DC8717FF79" box="[1235,1344,188,213]" captionStart="FIGURE 46" captionStartId="69.[151,250,1871,1893]" captionTargetBox="[178,1416,231,1822]" captionTargetId="figure-87@69.[151,1436,193,1850]" captionTargetPageId="69" captionText="FIGURE 46. Aposphragisma sepilok sp. nov. Male: A. carapace dorsal view; C. prosoma ventral view; E. carapace lateral view; G. carapace front view; H. carapace lateral view. Female: B. carapace dorsal view; D. prosoma ventral view; F. carapace lateral view. Bc, book lung cover; ctg, continuous transverse groove; sr, subterminal widely oval scutal ridge; tlp, tooth-like projection of male chelicerae. Scale bars: A–H 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4431999" httpUri="https://zenodo.org/record/4431999/files/figure.png" pageId="71" pageNumber="72">Fig. 46H</figureCitation>
|
||
); scutopedicel region with slightly swollen hair bases. Postepigastric scutum long, semicircular, with long posteriorly directed lateral apodemes (lap) (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6D5EF364803EFEB1" box="[512,617,260,285]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Fig. 47D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8159F37BFFD1FF9A6F99F347877AFE01" blockId="71.[151,1437,151,645]" pageId="71" pageNumber="72">
|
||
GENITALIA. Epigastric region: sperm pore situated between anterior and posterior spiracles (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6BBDF3478719FEEC" box="[1251,1358,295,320]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Fig. 47D</figureCitation>
|
||
). Palp: basal segments as well as bulb and cymbium lighter in colour than rest of body; bulb rather stout (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6BA4F32C870FFEC9" box="[1274,1368,332,357]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Figs. 47</figureCitation>
|
||
A–B). Embolus (em) tip spatulate (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6E8DF30F806EFE24" box="[467,569,367,392]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Fig. 47C</figureCitation>
|
||
); embolic spine (esp) with pointed tip (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6CB2F30F8607FE24" box="[1004,1104,367,392]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Fig. 47C</figureCitation>
|
||
). Conductor (con) with broad tip (appearing ‘snout’-like in lateral view;
|
||
<figureCitation id="19DDEFFEFFD1FF9A6D2DF3F4808EFE01" box="[627,729,404,429]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Fig. 47C</figureCitation>
|
||
). Embolus-conductor-complex about as long as bulb.
|
||
</paragraph>
|
||
<paragraph id="8159F37BFFD1FF9A6F99F3D78631FDB4" blockId="71.[151,1437,151,645]" pageId="71" pageNumber="72">
|
||
<emphasis id="B3922F69FFD1FF9A6F99F3D78366FE7C" bold="true" box="[199,305,439,464]" pageId="71" pageNumber="72">FEMALE.</emphasis>
|
||
As in male except as noted. Body length
|
||
<quantity id="461E5E9EFFD1FF9A6C71F3D7818EFE63" box="[815,985,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" metricValueMax="1.54" metricValueMin="1.52" pageId="71" pageNumber="72" unit="mm" value="1.53" valueMax="1.54" valueMin="1.52">1.52–1.54 mm</quantity>
|
||
, carapace length
|
||
<quantity id="461E5E9EFFD1FF9A6BF7F3D78704FE63" box="[1193,1363,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.1" metricValueMin="7.0" pageId="71" pageNumber="72" unit="mm" value="0.705" valueMax="0.71" valueMin="0.7">0.70–0.71 mm</quantity>
|
||
. PME separated by less than their radius. Epigastric scutum posterolaterally with a single curved ridge (cr) (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6A76F3BC87C7FE59" box="[1320,1424,476,501]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Fig. 47E</figureCitation>
|
||
). Postepigastric scutum with short posteriorly directed lateral apodemes (lap) (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6CAAF39F860DFDB4" box="[1012,1114,511,536]" captionStart="FIGURE 47" captionStartId="70.[151,250,1472,1494]" captionTargetBox="[173,1423,218,1409]" captionTargetId="figure-265@70.[151,1436,193,1451]" captionTargetPageId="70" captionText="FIGURE 47. Aposphragisma sepilok sp. nov. Male: A–B. palp prolateral view (B: hairs omitted except trichobothria); C. embolus-conductor-complex prolateral view; D. abdomen ventral view. Female: E. abdomen ventral view. Cb, conical bulge; con, conductor; cr, curved ridge; em, embolus; esp, embolic spine; lap, postepigastric lateral apodemes; spp, spermophor; spo, spermophor opening. Scale bars: A–B 100 µm; C 50 µm; D–E 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4432001" httpUri="https://zenodo.org/record/4432001/files/figure.png" pageId="71" pageNumber="72">Fig. 47E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8159F37BFFD1FF9A6F99F04483E0FDCC" blockId="71.[151,1437,151,645]" pageId="71" pageNumber="72">
|
||
<emphasis id="B3922F69FFD1FF9A6F99F044838CFD91" bold="true" box="[199,475,548,573]" pageId="71" pageNumber="72">Intraspecific variation.</emphasis>
|
||
Body size variable (see above); one female slightly darker; number of denticles on carapace margin variable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9FCA0F0FFD1FF9A6F99F00C871CFD29" box="[199,1355,620,645]" pageId="71" pageNumber="72" type="distribution">
|
||
<paragraph id="8159F37BFFD1FF9A6F99F00C871CFD29" blockId="71.[151,1437,151,645]" box="[199,1355,620,645]" pageId="71" pageNumber="72">
|
||
<emphasis id="B3922F69FFD1FF9A6F99F00C8337FD29" bold="true" box="[199,352,620,645]" pageId="71" pageNumber="72">Distribution.</emphasis>
|
||
NE-Sabah,
|
||
<collectingCountry id="F9F1B3EBFFD1FF9A6EB6F00C8034FD2F" box="[488,611,620,643]" name="Malaysia" pageId="71" pageNumber="72">MALAYSIA</collectingCountry>
|
||
; all specimens collected at Kabili-Sepilok forest reserve (
|
||
<figureCitation id="19DDEFFEFFD1FF9A6BB2F00C876CFD29" box="[1260,1339,620,645]" captionStart="FIGURE 49" captionStartId="73.[151,250,1482,1504]" captionTargetBox="[173,1409,203,1453]" captionTargetId="figure-249@73.[155,1427,185,1464]" captionTargetPageId="73" captionText="FIGURE 49. Distribution of Aposphragisma gen. nov. in Southeast Asia. Stripe clade: 1) A. baltenspergerae sp. nov., 2) A. borgulai sp. nov., 3) A. brunomanseri sp. nov., 4) A. dayak sp. nov., 5) A. dentatum sp. nov., 6) A. hausammannae sp. nov., 7) A. helvetiorum sp. nov., 8) A. kolleri sp. nov., 9) A. menzi sp. nov., 10) A. monoceros sp. nov., 11) A. retifer sp. nov., 12) A. salewskii sp. nov., 13) A. sepilok sp. nov. Non-stripe clade species: 14) A. confluens sp. nov., 15) A. draconigenum sp. nov., 16) A. nocturnum sp. nov., 17) A. rimba sp. nov., 18) A. scimitar sp. nov., 19) A. stannum sp. nov. Scale bars: 500 km." figureDoi="http://doi.org/10.5281/zenodo.4432005" httpUri="https://zenodo.org/record/4432005/files/figure.png" pageId="71" pageNumber="72">Fig. 49</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |