211 lines
24 KiB
XML
211 lines
24 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745281" ID-ZooBank="CE0AC7CD-E55D-484C-9346-86F16C985DBE" approvalRequired="29" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="1" checkinTime="1679054310250" checkinUser="plazi" docAuthor="Serrano, Artur R. M." docDate="2023" docId="AE0487C8F532FFDA0986A4736388F811" docLanguage="en" docName="zootaxa.5256.1.1.pdf" docOrigin="Zootaxa 5256 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrocheilus assmanni Serrano 2023, n. sp." docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="523DFFB0F533FFD80911A31C612BFFA8" masterDocTitle="Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes" masterLastPageNumber="35" masterPageNumber="1" pageNumber="2" updateTime="1679055118150" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Serrano, Artur R. M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-4282-6073</mods:nameIdentifier>
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<mods:affiliation>cE 3 c-Center for Ecology, Evolution and Environmental Changes & CHANGE - Global Change and Sustainability Institute, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, R. Ernesto de Vasconcelos, Ed. C 2, Campo Grande, 1749 - 016 Lisboa, PORTUGAL. aserrano @ fc. ul. pt; http: // orcid. org / 0000 - 0003 - 4282 - 6073</mods:affiliation>
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<mods:nameIdentifier type="email">aserrano@fc.ul.pt</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5256</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>35</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7745281</mods:identifier>
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<mods:identifier type="ZooBank">CE0AC7CD-E55D-484C-9346-86F16C985DBE</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7745344" ID-Zenodo-Dep="7745344" LSID="urn:lsid:plazi:treatment:AE0487C8F532FFDA0986A4736388F811" httpUri="http://treatment.plazi.org/id/AE0487C8F532FFDA0986A4736388F811" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<subSubSection box="[151,514,1903,1930]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph blockId="1.[151,514,1903,1966]" box="[151,514,1903,1930]" pageId="1" pageNumber="2">
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<heading bold="true" box="[151,514,1903,1930]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis bold="true" box="[151,514,1903,1930]" pageId="1" pageNumber="2">
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<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[151,441,1903,1930]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="assmanni" status="n. sp.">
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<emphasis bold="true" box="[151,441,1903,1930]" italics="true" pageId="1" pageNumber="2">Macrocheilus assmanni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[448,514,1904,1930]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,320,1940,1966]" pageId="1" pageNumber="2" type="description">
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<paragraph blockId="1.[151,514,1903,1966]" box="[151,320,1940,1966]" pageId="1" pageNumber="2">
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<figureCitation box="[151,241,1940,1966]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="1" pageNumber="2">Figs. 1a</figureCitation>
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,
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<figureCitation box="[254,283,1940,1966]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="1" pageNumber="2">2a</figureCitation>
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,
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<figureCitation box="[294,320,1940,1966]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="1" pageNumber="2">4a</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph blockId="1.[151,1436,2012,2039]" lastBlockId="2.[151,1437,151,1977]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<emphasis bold="true" box="[151,324,2012,2038]" pageId="1" pageNumber="2">Type material.</emphasis>
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<materialsCitation collectingDate="1996-12-12" collectingDateMax="1996-12-17" collectingDateMin="1996-12-12" collectorName="Werner & Th. Assmann" country="Tanzania" lastPageId="2" lastPageNumber="3" location="Ruvuma" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Ruvuma" typeStatus="holotype">
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<typeStatus box="[330,434,2012,2038]" pageId="1" pageNumber="2">Holotype</typeStatus>
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female: “
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<collectingCountry box="[541,647,2012,2038]" name="Tanzania" pageId="1" pageNumber="2">Tanzania</collectingCountry>
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,
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<date box="[657,830,2012,2038]" pageId="1" pageNumber="2" value="1996-12-12" valueMax="1996-12-17" valueMin="1996-12-12">
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<collectingDate box="[657,830,2012,2038]" pageId="1" pageNumber="2" value="1996-12-12" valueMax="1996-12-17" valueMin="1996-12-12">12-17.XII.1996</collectingDate>
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</date>
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\
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<collectingRegion box="[845,943,2013,2039]" country="Tanzania" name="Ruvuma" pageId="1" pageNumber="2">Ruvuma</collectingRegion>
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, near Songea\ leg.
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<collectorName box="[1151,1233,2013,2039]" pageId="1" pageNumber="2">Werner</collectorName>
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& Lizler” \\ “Collection\
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<collectorName box="[241,387,151,177]" pageId="2" pageNumber="3">Th. Assmann</collectorName>
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\ Bleckede, CAB” (yellow label)\\
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</materialsCitation>
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<typeStatus box="[771,875,151,177]" pageId="2" pageNumber="3">Holotype</typeStatus>
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female\
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<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[971,1236,151,177]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="assmanni" status="n. sp.">
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<emphasis box="[971,1236,151,177]" italics="true" pageId="2" pageNumber="3">Macrocheilus assmanni</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1242,1303,151,177]" pageId="2" pageNumber="3">
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<taxonomicNameLabel box="[1242,1303,151,177]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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\ A. Serrano det. 2022 [h] [red card].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="diagnosis">
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[199,461,222,248]" pageId="2" pageNumber="3">Differential diagnosis.</emphasis>
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Head, pronotum, elytra and legs (except tarsi) black; antennae, distal half of mandibles, palps and tarsi reddish-brown coloured; anterior sub marginal and anterolateral surfaces of pronotum rugose-punctured; elytra without patches; shiny and pubescent.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="description">
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<paragraph blockId="2.[151,1437,151,1977]" box="[199,715,330,357]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[199,345,330,356]" pageId="2" pageNumber="3">Description.</emphasis>
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Length of the
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<typeStatus box="[509,611,331,357]" pageId="2" pageNumber="3">holotype</typeStatus>
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:
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<quantity box="[622,711,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="3" unit="mm" value="7.7">7.7 mm</quantity>
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.
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</paragraph>
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis box="[199,260,367,393]" italics="true" pageId="2" pageNumber="3">Head</emphasis>
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(
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<figureCitation box="[275,355,367,393]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="2" pageNumber="3">Fig. 1a</figureCitation>
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). Wider than long; length
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<quantity box="[650,753,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="2" pageNumber="3" unit="mm" value="1.44">1.44 mm</quantity>
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; width
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<quantity box="[836,939,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.98" pageId="2" pageNumber="3" unit="mm" value="1.98">1.98 mm</quantity>
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. Coarsely and deeply punctate, less dense in middle of the frons, each puncture with a long and fine brown-yellowish seta, without microsculpture; last maxillary palpomere slightly dilated, pubescent; eyes prominent, temples short with rows of large setae along hind margin of eyes. Clypeus less densely punctate, each puncture with a long and fine brown-yellowish seta, anterior half with a row of eight large setiferous punctures near the anterior margin. Labrum smooth, wider than long (length
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<quantity box="[1330,1432,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="3" unit="mm" value="0.77">0.77 mm</quantity>
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, width
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<quantity box="[224,329,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="3" unit="mm" value="0.91">0.91 mm</quantity>
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), anterior half rounded along margins, middle sides subparallel, posterior 1/3 sides converging to the base; at the widest point a large seta at each side near the rim, two major discal setae and two anterior marginal setae near the apex (
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<figureCitation box="[379,460,619,645]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="2" pageNumber="3">Fig. 2a</figureCitation>
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). A pair of supraorbital setae on each side. Mentum with a central tooth, acuminate and sharp. Antennae with the first and third antennomeres almost cylindrical, the second and fourth antennomeres subpyriforms, the remaining ones depressed, slightly longer than wide, with a thin median longitudinal smooth area; last antennomere more or less acuminate.
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</paragraph>
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis box="[199,280,763,789]" italics="true" pageId="2" pageNumber="3">Thorax</emphasis>
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. Pronotum cordiform (
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<figureCitation box="[536,615,763,789]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="2" pageNumber="3">Fig. 1a</figureCitation>
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), transverse, 1.6 times wider than long (length
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<quantity box="[1138,1240,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="3" unit="mm" value="1.22">1.22 mm</quantity>
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; width
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<quantity box="[1320,1424,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="2" pageNumber="3" unit="mm" value="1.95">1.95 mm</quantity>
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); surface covered with coarse punctures, each bearing one long and fine brown-yellowish seta, intervals smooth; anterior submarginal and anterolateral surfaces rugose-punctate, forming irregular meshes; anterior angles rounded, anterior margin arcuate; posterior margin arcuate in the middle, slightly oblique sinuate towards the posterior angles which are obtuse-rounded and bear a long seta; lateral margins sinuate just anterior to posterior angles, a lateral long seta near the widest part of pronotum, anterior angles with some large bristles; anterior transverse sulcus distinct, pronotal base with a pair of large foveae separated by a median faint sulcus reaching the posterior margin. Scutellum triangular, with scattered setiferous punctures. Prosternum coarsely punctured, without microsculpture.
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</paragraph>
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis box="[199,269,1051,1077]" italics="true" pageId="2" pageNumber="3">Elytra</emphasis>
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(
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<figureCitation box="[288,369,1051,1077]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="2" pageNumber="3">Fig. 1a</figureCitation>
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) subparallel-sided, 1.8 times longer than wide (length
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<quantity box="[1009,1114,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="3" unit="mm" value="5.08">5.08 mm</quantity>
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, width
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<quantity box="[1203,1310,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="2" pageNumber="3" unit="mm" value="2.85">2.85 mm</quantity>
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), widening slightly towards the posterior third, not covering the last tergite; only seven striae visible, well-marked and punctured, not reaching the base and the apex; intervals slightly convex, two rows of coarse setiferous punctures close to striae, pubescence sub-erected and directed posteriad; lateral margin rounded in the shoulders, ending in deep furrow densely punctuated towards the scutellum. Scutellar striae slightly punctured and visible between the two rows of first interval. Umbilicate series with about 20 pores, slightly separated in the middle (14+6), apex with 4 extra pores, each in the lengthening of 7
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<superScript attach="none" box="[598,611,1266,1280]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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, 6
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<superScript attach="none" box="[638,651,1266,1280]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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, 5
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<superScript attach="left" box="[678,691,1266,1280]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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and 4
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<superScript attach="left" box="[759,772,1266,1280]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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striae; apical transparent membrane extending from sutural angle to the lengthening of the imaginary 8
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<superScript attach="left" box="[634,647,1302,1316]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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interval.
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</paragraph>
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis box="[199,252,1339,1364]" italics="true" pageId="2" pageNumber="3">Legs</emphasis>
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blackish, knees and tarsomeres reddish-brown coloured; all tarsomeres pubescent, first three fore tarsomeres not dilated, fourth tarsomere triangulate; fore tibiae without blunt spines in the outer margin.
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</paragraph>
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis box="[199,370,1411,1437]" italics="true" pageId="2" pageNumber="3">Ventral surface</emphasis>
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. Almost all underside blackish, posterior margins of abdominal sternites red-brownish; whole underside, including abdominal sternites coarsely punctured and pubescent (yellowish color), except gula and ventral and lateral sides of neck. Pubescence of prosternum and proepisternum sub-erected and slightly directed anteriad; pubescence of the remaining ventral sclerites dressed and directed posteriad. Elytral epipleura moderately punctured, pubescence dressed and directed posteriad.
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</paragraph>
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis box="[199,392,1591,1617]" italics="true" pageId="2" pageNumber="3">Female genitalia</emphasis>
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(
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<figureCitation box="[408,492,1591,1617]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="2" pageNumber="3">Fig. 4a</figureCitation>
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). Gonocoxite 1 subtriangular; gonocoxite 2 short, largely sickle shaped, apex short and sharp; dorsal surface concave, strongly setose, inner margin with 1 seta; ventral surface convex, smooth.
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</paragraph>
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<paragraph blockId="2.[151,1437,151,1977]" box="[199,389,1663,1689]" pageId="2" pageNumber="3">
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<emphasis box="[199,257,1663,1689]" italics="true" pageId="2" pageNumber="3">Male</emphasis>
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. Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="etymology">
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[199,334,1699,1725]" pageId="2" pageNumber="3">Etymology.</emphasis>
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This species is named in honour of Professor Thorsten Assmann (Leuphana University, Lüneburg,
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<collectingCountry box="[151,260,1735,1761]" name="Germany" pageId="2" pageNumber="3">Germany</collectingCountry>
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), a well-known carabidologist.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="discussion">
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<paragraph blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[199,308,1771,1797]" pageId="2" pageNumber="3">Remarks</emphasis>
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. The new species is akin to
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<taxonomicName authority="Burgeon, 1937" authorityName="Burgeon" authorityYear="1937" box="[620,1045,1771,1797]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alluaudi">
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<emphasis box="[620,873,1771,1797]" italics="true" pageId="2" pageNumber="3">Macrocheilus alluaudi</emphasis>
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<bibRefCitation author="Burgeon, L." box="[880,1045,1771,1797]" pageId="2" pageNumber="3" pagination="319 - 405" refId="ref25317" refString="Burgeon, L. (1937) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres Carabides. Troisieme Partie. Annales du Musee du Congo Belge, Zoology, Serie 3 (2), 2 (5), 319 - 405." type="journal article" year="1937">Burgeon, 1937</bibRefCitation>
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</taxonomicName>
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by the black body colour (
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<figureCitation captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="2" pageNumber="3">
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Figs. 1f,
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<quantity box="[151,179,1807,1833]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="2" pageNumber="3" unit="g" value="1.0">1g</quantity>
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</figureCitation>
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), differing from this species by the smaller size and shape of last maxillary palpomere and labrum, among other external morphological characters. This slender and small new species differs from the remaining African species by the totally black body colour, with absence of yellow/orange elytral spots or bands, and by the distal surface of pronotum strongly rugose-punctuated, forming irregular meshes.
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,675,1950,1977]" pageId="2" pageNumber="3" type="distribution">
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<paragraph blockId="2.[151,1437,151,1977]" box="[199,675,1950,1977]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[199,345,1950,1976]" pageId="2" pageNumber="3">Distribution</emphasis>
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. Only known from
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<collectingCountry box="[567,671,1951,1977]" name="Tanzania" pageId="2" pageNumber="3">Tanzania</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |