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<document id="178F3F7E64ABBEAB7C042BFF28CBA3AE" ID-CLB-Dataset="57399" ID-DOI="10.5852/ejt.2023.878.2153" ID-GBIF-Dataset="0dfb60c4-1de3-4062-9443-15b55e7e0c88" ID-ISSN="2118-9773" ID-Zenodo-Dep="8110550" ID-ZooBank="07EFE4B6-4184-41C7-82A5-747BF67BA093" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1688395572616" checkinUser="felipe" docAuthor="Grichanov, Igor Ya." docDate="2023" docId="282DC677E924FF84C029C8D3FC2A7FEE" docLanguage="en" docName="ejt-878_1-52.pdf" docOrigin="European Journal of Taxonomy 878" docSource="http://zoobank.org/07efe4b6-4184-41c7-82a5-747bf67ba093" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Thinophilus longicercus Grichanov 2023, sp. nov." docType="treatment" docUuid="812BFAC4-FB3B-4F00-BD46-86BC4CDCFD4C" docUuidSource="ZooBank" docVersion="11" lastPageNumber="31" masterDocId="D414BE0FE938FF9AC27ACA64FFC27B3F" masterDocTitle="A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="29" updateTime="1699533079317" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D3EA3F3EBE2F489E21907F0408AAE738">A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species</mods:title>
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<mods:namePart id="406C009BD9E26F3AA20B10479A2BC3DD">Grichanov, Igor Ya.</mods:namePart>
<mods:affiliation id="B7848DEB5FBF4379285CD4E5F85AC24B">All-Russian Institute of Plant Protection, Podbelskogo 3, St Petersburg, Pushkin, 196608, Russia.</mods:affiliation>
<mods:nameIdentifier id="9808B7CC4B623FF5BD3D28D526341B61" type="email">grichanov@mail.ru</mods:nameIdentifier>
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<mods:title id="45D094F1CC0A9E69914F7B4B8A157A32">European Journal of Taxonomy</mods:title>
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<mods:part id="AA8095806E36B9E8DFE5391BCA1A8A62">
<mods:date id="387277AC1114C4D0F9915A0F19B9C02B">2023</mods:date>
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<mods:number id="D4EA23CBFD2AB8B96D351ED669FCA5FF">2023-07-03</mods:number>
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<mods:number id="B5764E915A6CAB29B544D24EE3ABDE6C">878</mods:number>
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<treatment id="282DC677E924FF84C029C8D3FC2A7FEE" ID-DOI="http://doi.org/10.5281/zenodo.8119226" ID-GBIF-Taxon="210883713" ID-Zenodo-Dep="8119226" LSID="urn:lsid:zoobank.org:act:812BFAC4-FB3B-4F00-BD46-86BC4CDCFD4C" httpUri="http://treatment.plazi.org/id/282DC677E924FF84C029C8D3FC2A7FEE" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="E89E24EAE924FF86C029C8D3FC2379ED" box="[595,993,695,722]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="A03B7761E924FF86C029C8D3FC2379ED" blockId="28.[353,1234,695,792]" box="[595,993,695,722]" pageId="28" pageNumber="29">
<heading id="FB73C00DE924FF86C029C8D3FC2379ED" box="[595,993,695,722]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="67840CE2E924FF86C029C8D3FCBD79ED" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[595,895,695,722]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="longicercus" status="sp. nov.">
<emphasis id="92F0AB73E924FF86C029C8D3FCBD79ED" bold="true" box="[595,895,695,722]" italics="true" pageId="28" pageNumber="29">Thinophilus longicercus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E924FF86C1FCC8DCFC2379ED" box="[902,993,696,722]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="E89E24EAE924FF86C31BC8BEFCFE7827" pageId="28" pageNumber="29" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="A03B7761E924FF86C08DC899FCFE7827" blockId="28.[353,1234,695,792]" box="[759,828,765,792]" pageId="28" pageNumber="29">
<figureCitation id="38BF6BE4E924FF86C08DC899FCFE7827" box="[759,828,765,792]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,845,1887]" captionTargetId="figure-189@29.[189,1398,845,1889]" captionTargetPageId="29" captionText="Fig. 9. Thinophilus longicercus sp. nov., holotype, ♂ (RBINS). A. Habitus. B. Fore leg, posterior view. C. Hind femur. D. Wing. E. Hypopygium after maceration with basal half of cercus, lateral view. F. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="28" pageNumber="29">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE924FF86C2C7C95BFE577F2B" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="A03B7761E924FF86C2C7C95BFEF57866" blockId="28.[189,311,831,857]" box="[189,311,831,857]" pageId="28" pageNumber="29">
<heading id="FB73C00DE924FF86C2C7C95BFEF57866" bold="true" box="[189,311,831,857]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="92F0AB73E924FF86C2C7C95BFEF57866" bold="true" box="[189,311,831,857]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E924FF86C2C7C909FE577F2B" blockId="28.[188,1398,877,1045]" pageId="28" pageNumber="29">
<taxonomicName id="67840CE2E924FF86C2C7C909FE1978B8" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[189,475,877,903]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="longicercus" status="sp. nov.">
<emphasis id="92F0AB73E924FF86C2C7C909FE1978B8" box="[189,475,877,903]" italics="true" pageId="28" pageNumber="29">Thinophilus longicercus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E924FF86C398C90AFDFE78B7" box="[482,572,878,904]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
keys to
<taxonomicName id="67840CE2E924FF86C0E6C90AFCF078B8" authorityName="ZIN" baseAuthorityName="Curran" baseAuthorityYear="1924" box="[668,818,877,903]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="92F0AB73E924FF86C0E6C90AFCF078B8" box="[668,818,877,903]" italics="true" pageId="28" pageNumber="29">T. imperialis</emphasis>
</taxonomicName>
(see key above), both species having male cercus very long, thin distally, extending to the base of abdomen, and sternites 3 and 4 of abdomen with tuft of mainly black hairs. Fore tibia bears 34 dorsal bristles, not longer than tibia width in
<taxonomicName id="67840CE2E924FF86C6D6C9D1FA8078F1" authorityName="ZIN" baseAuthorityName="Curran" baseAuthorityYear="1924" box="[1196,1346,948,974]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="92F0AB73E924FF86C6D6C9D1FA8078F1" box="[1196,1346,948,974]" italics="true" pageId="28" pageNumber="29">T. imperialis</emphasis>
</taxonomicName>
, but fore tibia with 34 dorsal bristles, 2 × as long as tibia width in
<taxonomicName id="67840CE2E924FF86C1F6C9BCFBF078CE" authorityName="Grichanov" authorityYear="2023" box="[908,1074,983,1009]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="longicercus">
<emphasis id="92F0AB73E924FF86C1F6C9BCFBF078CE" box="[908,1074,983,1009]" italics="true" pageId="28" pageNumber="29">T. longicercus</emphasis>
</taxonomicName>
; surstylus shape is different in the two species.
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE924FF86C2C7CE58FAC27FBB" box="[189,1280,1084,1157]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="A03B7761E924FF86C2C7CE58FE877F69" blockId="28.[189,325,1084,1110]" box="[189,325,1084,1110]" pageId="28" pageNumber="29">
<heading id="FB73C00DE924FF86C2C7CE58FE877F69" bold="true" box="[189,325,1084,1110]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="92F0AB73E924FF86C2C7CE58FE877F69" bold="true" box="[189,325,1084,1110]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E924FF86C2C7CE0EFAC27FBB" blockId="28.[189,1280,1130,1157]" box="[189,1280,1130,1157]" pageId="28" pageNumber="29">
The specific epithet
<taxonomicName id="67840CE2E924FF86C3CECE0EFDFF7FBB" authority="(Latin)" authorityName="Grichanov" authorityYear="2023" baseAuthorityName="Latin" box="[436,573,1130,1156]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="longicercus">
<emphasis id="92F0AB73E924FF86C3CECE0EFDFF7FBB" box="[436,573,1130,1156]" italics="true" pageId="28" pageNumber="29">longicercus</emphasis>
</taxonomicName>
(Latin) refers to the long
<emphasis id="92F0AB73E924FF86C104CE0FFC087FBB" box="[894,970,1131,1156]" italics="true" pageId="28" pageNumber="29">cercus</emphasis>
of the male hypopygium.
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE924FF86C2C7CECFFD007E61" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="A03B7761E924FF86C2C7CECFFE6F7FFA" blockId="28.[189,429,1195,1221]" box="[189,429,1195,1221]" pageId="28" pageNumber="29">
<heading id="FB73C00DE924FF86C2C7CECFFE6F7FFA" bold="true" box="[189,429,1195,1221]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="92F0AB73E924FF86C2C7CECFFE6F7FFA" bold="true" box="[189,429,1195,1221]" pageId="28" pageNumber="29">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E924FF86C290CEBEFD007E61" blockId="28.[189,1400,1242,1375]" pageId="28" pageNumber="29">
<materialsCitation id="10EC7D3CE924FF86C290CEBEFD007E61" ID-GBIF-Occurrence="4145165366" collectingDate="1955-12" collectionCode="RBINS" collectorName="B. Stuckenberg" country="Madagascar" county="Fenoarivo Atsinanana" determinerName="P. Vanschuytbroeck" latitude="-17.366667" location="Lagoon" longLatPrecision="1279" longitude="49.416668" municipality="Fenerive" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="92F0AB73E924FF86C290CEBEFE9F7FCB" bold="true" box="[234,349,1242,1268]" pageId="28" pageNumber="29">
<typeStatus id="7F3FC9C3E924FF86C290CEBEFE9F7FCB" box="[234,349,1242,1268]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="D89337F1E924FF86C2C7CE9AFE4C7E27" box="[189,398,1278,1304]" name="Madagascar" pageId="28" pageNumber="29">MADAGASCAR</collectingCountry>
<specimenCount id="B682BCE8E924FF86C3D2CE9AFE7C7E27" box="[424,446,1278,1304]" pageId="28" pageNumber="29" type="male"></specimenCount>
(partly covered with mould); “
<location id="A55B21BAE924FF86C150CE9AFC447E27" LSID="urn:lsid:plazi:treatment:282DC677E924FF84C029C8D3FC2A7FEE:A55B21BAE924FF86C150CE9AFC447E27" box="[810,902,1278,1304]" country="Madagascar" county="Fenoarivo Atsinanana" latitude="-17.366667" longLatPrecision="1279" longitude="49.416668" municipality="Fenerive" name="Lagoon" pageId="28" pageNumber="29">Lagoon</location>
shore;
<collectingMunicipality id="405FED1BE924FF86C1A7CE99FB867E28" box="[989,1092,1277,1303]" pageId="28" pageNumber="29">Fenerive</collectingMunicipality>
[=
<collectingCounty id="495A0FEDE924FF86C617CE99FAB07E28" box="[1133,1394,1277,1303]" pageId="28" pageNumber="29">Fenoarivo Atsinanana</collectingCounty>
;
<geoCoordinate id="C5B011A6E924FF86C2C7CF7BFEE77E04" box="[189,293,1311,1339]" degrees="17" direction="south" minutes="22" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="-17.366667">17°22 S</geoCoordinate>
,
<geoCoordinate id="C5B011A6E924FF86C349CF7BFE5D7E04" box="[307,415,1311,1339]" degrees="49" direction="east" minutes="25" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="49.416668">49°25 E</geoCoordinate>
];
<date id="D43A51A1E924FF86C3CFCF45FDF17E04" box="[437,563,1312,1339]" pageId="28" pageNumber="29" value="1955-12">
<collectingDate id="C47EA849E924FF86C3CFCF45FDF17E04" box="[437,563,1312,1339]" pageId="28" pageNumber="29" value="1955-12">Dec. 1955</collectingDate>
</date>
;
<collectorName id="0D7112B7E924FF86C03ACF45FD347E04" box="[576,758,1313,1339]" pageId="28" pageNumber="29">B. Stuckenberg</collectorName>
leg.;
<taxonomicName id="67840CE2E924FF86C141CF45FB477E04" authority="Loew" authorityName="Loew" authorityYear="1857" box="[827,1157,1313,1339]" class="Insecta" family="Dolichopodidae" genus="Hydrophorus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bisetus">
<emphasis id="92F0AB73E924FF86C141CF45FBF17E04" box="[827,1075,1313,1339]" italics="true" pageId="28" pageNumber="29">Hydrophorus bisetus</emphasis>
Loew
</taxonomicName>
;
<determinerName id="55E79BA5E924FF86C6EBCF45FABA7E04" box="[1169,1400,1313,1339]" pageId="28" pageNumber="29">P. Vanschuytbroeck</determinerName>
det. 1957; I.R.Sc.N.B. I.G. 20938”;
<collectionCode id="C695EFA4E924FF86C01ECF20FD7C7E61" box="[612,702,1348,1374]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE924FF84C2C7CFE1FEFB78C8" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="A03B7761E924FF86C2C7CFE1FE8D7EA0" blockId="28.[189,335,1413,1439]" box="[189,335,1413,1439]" pageId="28" pageNumber="29">
<heading id="FB73C00DE924FF86C2C7CFE1FE8D7EA0" bold="true" box="[189,335,1413,1439]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="92F0AB73E924FF86C2C7CFE1FE8D7EA0" bold="true" box="[189,335,1413,1439]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E924FF86C290CFD0FE5D7EF1" blockId="28.[189,1399,1460,1522]" box="[234,415,1460,1486]" pageId="28" pageNumber="29">
<emphasis id="92F0AB73E924FF86C290CFD0FEE87EF1" bold="true" box="[234,298,1460,1486]" pageId="28" pageNumber="29">Male</emphasis>
(
<figureCitation id="38BF6BE4E924FF86C340CFD0FE557EF1" box="[314,407,1460,1486]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,845,1887]" captionTargetId="figure-189@29.[189,1398,845,1889]" captionTargetPageId="29" captionText="Fig. 9. Thinophilus longicercus sp. nov., holotype, ♂ (RBINS). A. Habitus. B. Fore leg, posterior view. C. Hind femur. D. Wing. E. Hypopygium after maceration with basal half of cercus, lateral view. F. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="28" pageNumber="29">Fig. 9A</figureCitation>
)
</paragraph>
<paragraph id="A03B7761E924FF86C2C7CFBCFAB57ECD" blockId="28.[189,1399,1460,1522]" box="[189,1399,1495,1522]" pageId="28" pageNumber="29">
<smallCapsWord id="A6DDE1BDE924FF86C2C7CFBCFEAC7ECF" baselines="1515,1515" box="[189,366,1496,1522]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="28" pageNumber="29">MEASUREMENTS</smallCapsWord>
. Body length
<quantity id="677CDA84E924FF86C068CFB3FDAC7ECD" box="[530,622,1495,1522]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="28" pageNumber="29" unit="mm" value="6.3">6.3 mm</quantity>
; antenna length
<quantity id="677CDA84E924FF86C151CFB3FC4A7ECD" box="[811,904,1495,1522]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="28" pageNumber="29" unit="mm" value="1.1">1.1 mm</quantity>
; wing length
<quantity id="677CDA84E924FF86C65ECFB3FB427ECD" box="[1060,1152,1495,1522]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="28" pageNumber="29" unit="mm" value="5.2">5.2 mm</quantity>
; wing width
<quantity id="677CDA84E924FF86C76CCFB3FAB07ECD" box="[1302,1394,1495,1522]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="28" pageNumber="29" unit="mm" value="1.6">1.6 mm</quantity>
.
</paragraph>
<paragraph id="A03B7761E924FF86C2C7CC7DFBF87C16" blockId="28.[187,1400,1560,1833]" pageId="28" pageNumber="29">
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. Postcranium bluish black, grey pollinose; frons bluish black, weakly pollinose; face and clypeus greenish black, brownish grey pollinose; face under antennae 1.8 × as wide as height of postpedicel; clypeus half as long as epistoma, 1.6 × as wide as long; palp yellow, bearing black bristly hairs; proboscis black; 2 diverging ocellars; 1 vertical, 1 postvertical, much stronger and longer than, and not in row with upper postoculars; upper postoculars uniseriate, black; lower postoculars multiseriate, white, long; antennal scape, pedicel and postpedicel black dorsally, orange ventrally; scape invisible; pedicel simple, convex on inner side; postpedicel rounded, with short pubescence, slightly higher than long (18/14); arista-like stylus dorsal, black, thick basally, thin distally, shortly pubescent.
</paragraph>
<paragraph id="A03B7761E924FF86C2C7CD35FB507C8E" blockId="28.[189,1398,1872,1969]" pageId="28" pageNumber="29">
<smallCapsWord id="A6DDE1BDE924FF86C2C7CD35FEDB7C56" baselines="1892,1892" box="[189,281,1873,1899]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="28" pageNumber="29">THORAX</smallCapsWord>
. Metallic bluish black, grey dusted; mesonotum darker, with matt black postalar spot; no acrostichals; 67 dorsocentrals decreasing in length anteriorly; scutellum with 2 strong marginals and 2 minute laterals; few upper and lower, white propleural bristles of different length.
</paragraph>
<paragraph id="A03B7761E925FF87C2C7CB6DFB8A7926" blockId="29.[189,1400,265,538]" pageId="29" pageNumber="30">
<smallCapsWord id="A6DDE1BDE925FF87C2C7CB6DFF347A1E" baselines="284,284" box="[189,246,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="29" pageNumber="30">LEGS</smallCapsWord>
. Fore coxa brown-black; mid and hind coxae black, orange-yellow at apex, grey pollinose; legs mostly dirty yellow, darker at knees; tibiae and tarsomeres 13 brown or black at tips; tarsomeres 45 brown-black. Fore leg (
<figureCitation id="38BF6BE4E925FF87C3AECB2BFDF37A56" box="[468,561,335,361]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,845,1887]" captionTargetId="figure-189@29.[189,1398,845,1889]" captionTargetPageId="29" captionText="Fig. 9. Thinophilus longicercus sp. nov., holotype, ♂ (RBINS). A. Habitus. B. Fore leg, posterior view. C. Hind femur. D. Wing. E. Hypopygium after maceration with basal half of cercus, lateral view. F. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="29" pageNumber="30">Fig. 9B</figureCitation>
). Coxa with long black setae and apical bristles; femur thickened, with ventral and posteroventral rows of strong bristles, about half as long as femur height; tibia bearing 34 long erect dorsal bristles, ventral row of short thick spinules, posteroventral row of elongate setulae, 2 long apicoventral bristles, devoid of black setulae anteriorly; basitarsus with ventral row of short thick spinules and posteroventral row of elongate setulae; segment 5 weakly widened and flattened; length of femur, tibia and tarsal segments (in mm): 1.63/1.42/0.64/0.29/0.19/0.16/0.17.
</paragraph>
<paragraph id="A03B7761E925FF87C2C7C825FDE079FB" blockId="29.[188,1398,575,708]" pageId="29" pageNumber="30">
<smallCapsWord id="A6DDE1BDE925FF87C2C7C825FF2F7965" baselines="596,597" box="[189,237,577,603]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mid" pageId="29" pageNumber="30">MID</smallCapsWord>
<smallCapsWord id="A6DDE1BDE925FF87C28EC822FEDE7966" baselines="596" box="[244,284,582,601]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="29" pageNumber="30">LEG</smallCapsWord>
. Coxa with black setae and bristles; femur with 2 ventral rows of short setae, at most ⅓ as long as femur height; tibia bearing 3 anterodorsal; 2 posterodorsal, 3 anteroventral, 2 posteroventral, 4 apical bristles; segment 5 inconspicuously widened; length of femur, tibia and tarsal segments (in mm): 1.89/ 1.97/1.02/0.34/0.26/0.22/0.22.
</paragraph>
<caption id="F4FB27E9E925FF87C2C7CDECFC1E7CD7" ID-DOI="http://doi.org/10.5281/zenodo.8110568" ID-Zenodo-Dep="8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1928,1954]" targetBox="[189,1398,845,1887]" targetPageId="29">
<paragraph id="A03B7761E925FF87C2C7CDECFC1E7CD7" blockId="29.[189,1399,1928,2025]" pageId="29" pageNumber="30">
<emphasis id="92F0AB73E925FF87C2C7CDECFED27C9D" bold="true" box="[189,272,1928,1954]" pageId="29" pageNumber="30">Fig. 9.</emphasis>
<taxonomicName id="67840CE2E925FF87C366CDECFDFD7C9D" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[284,575,1928,1954]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="longicercus" status="sp. nov.">
<emphasis id="92F0AB73E925FF87C366CDECFDFD7C9D" box="[284,575,1928,1954]" italics="true" pageId="29" pageNumber="30">Thinophilus longicercus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E925FF87C036CDECFD6F7C9D" box="[588,685,1928,1954]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS).
<emphasis id="92F0AB73E925FF87C1ADCDECFC2E7C9D" bold="true" box="[983,1004,1928,1954]" pageId="29" pageNumber="30">A</emphasis>
. Habitus.
<emphasis id="92F0AB73E925FF87C614CDECFB407C9D" bold="true" box="[1134,1154,1928,1954]" pageId="29" pageNumber="30">B</emphasis>
. Fore leg, posterior view.
<emphasis id="92F0AB73E925FF87C379CDCFFEDA7CFA" bold="true" box="[259,280,1963,1989]" pageId="29" pageNumber="30">C</emphasis>
. Hind femur.
<emphasis id="92F0AB73E925FF87C3C6CDCFFE137CFA" bold="true" box="[444,465,1963,1989]" pageId="29" pageNumber="30">D</emphasis>
. Wing.
<emphasis id="92F0AB73E925FF87C054CDCFFD807CFA" bold="true" box="[558,578,1963,1989]" pageId="29" pageNumber="30">E</emphasis>
. Hypopygium after maceration with basal half of cercus, lateral view.
<emphasis id="92F0AB73E925FF87C2C7CDAAFF0D7CD7" bold="true" box="[189,207,1998,2024]" pageId="29" pageNumber="30">F</emphasis>
. Distal appendages of hypopygium after maceration, ventral view.
</paragraph>
</caption>
<paragraph id="A03B7761E926FF84C2C7CB6DFC6F7AB2" blockId="30.[188,1398,265,397]" pageId="30" pageNumber="31">
<smallCapsWord id="A6DDE1BDE926FF84C2C7CB6DFF357A1D" baselines="284,285" box="[189,247,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hind" pageId="30" pageNumber="31">HIND</smallCapsWord>
<smallCapsWord id="A6DDE1BDE926FF84C287CB6AFEE77A1E" baselines="284" box="[253,293,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="30" pageNumber="31">LEG</smallCapsWord>
. Coxa with 1 black exterior bristle; femur (
<figureCitation id="38BF6BE4E926FF84C162CB6DFCB37A1C" box="[792,881,265,291]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,845,1887]" captionTargetId="figure-189@29.[189,1398,845,1889]" captionTargetPageId="29" captionText="Fig. 9. Thinophilus longicercus sp. nov., holotype, ♂ (RBINS). A. Habitus. B. Fore leg, posterior view. C. Hind femur. D. Wing. E. Hypopygium after maceration with basal half of cercus, lateral view. F. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="30" pageNumber="31">Fig. 9C</figureCitation>
) ventrally with anteroventral row of bristles, about femur height; 3 anterodorsal bristles; tibia bearing 4 anterodorsal, 4 posterodorsal bristles, 2 anteroventral, 3 posteroventral, 4 apicals; segment 5 inconspicuously widened; length of femur, tibia and tarsal segments (in mm): 2.73/2.8/0.67/0.54/0.37/0.24/0.26.
</paragraph>
<paragraph id="A03B7761E926FF84C2C7CBD1FC2A7907" blockId="30.[189,1399,436,568]" pageId="30" pageNumber="31">
<smallCapsWord id="A6DDE1BDE926FF84C2C7CBD1FF3C7AF2" baselines="456,456" box="[189,254,437,463]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Wing" pageId="30" pageNumber="31">WING</smallCapsWord>
(
<figureCitation id="38BF6BE4E926FF84C374CBD0FEAE7AF1" box="[270,364,436,463]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,845,1887]" captionTargetId="figure-189@29.[189,1398,845,1889]" captionTargetPageId="29" captionText="Fig. 9. Thinophilus longicercus sp. nov., holotype, ♂ (RBINS). A. Habitus. B. Fore leg, posterior view. C. Hind femur. D. Wing. E. Hypopygium after maceration with basal half of cercus, lateral view. F. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="30" pageNumber="31">Fig. 9D</figureCitation>
). Fumose, without distinct dark spots; veins brown; distal part of M
<subScript id="3C007524E926FF84C605CBA3FB587AEA" attach="left" box="[1151,1178,455,469]" fontSize="6" pageId="30" pageNumber="31">1+2</subScript>
convex; tip of R
<subScript id="3C007524E926FF84C726CBA3FAB47AEA" attach="left" box="[1372,1398,455,469]" fontSize="6" pageId="30" pageNumber="31">4+5</subScript>
parallel with M
<subScript id="3C007524E926FF84C30FCB8EFE527AC7" attach="left" box="[373,400,490,504]" fontSize="6" pageId="30" pageNumber="31">1+2</subScript>
; ratio of part of costa between R
<subScript id="3C007524E926FF84C160CB8EFCF77AC7" attach="left" box="[794,821,490,504]" fontSize="6" pageId="30" pageNumber="31">2+3</subScript>
and R
<subScript id="3C007524E926FF84C1FFCB8EFC5D7AC7" attach="left" box="[901,927,490,504]" fontSize="6" pageId="30" pageNumber="31">4+5</subScript>
to that between R
<subScript id="3C007524E926FF84C600CB8EFB577AC7" attach="left" box="[1146,1173,490,504]" fontSize="6" pageId="30" pageNumber="31">4+5</subScript>
and M
<subScript id="3C007524E926FF84C691CB8EFAC47AC7" attach="left" box="[1259,1286,490,504]" fontSize="6" pageId="30" pageNumber="31">1+2</subScript>
(in mm), 0.5/0.28; crossvein dm-m straight; ratio of dm-m to distal part of M
<subScript id="3C007524E926FF84C1AFC869FC1C7924" attach="left" box="[981,990,525,539]" fontSize="6" pageId="30" pageNumber="31">4</subScript>
, 0.4/0.58; anal vein distinct; halter dirty yellow; lower calypter dirty yellow, with white and brown cilia.
</paragraph>
<paragraph id="A03B7761E926FF84C2C7C83BFEE278AC" blockId="30.[185,1399,607,915]" pageId="30" pageNumber="31">
<smallCapsWord id="A6DDE1BDE926FF84C2C7C83BFEEE7947" baselines="626,627" box="[189,300,607,633]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="30" pageNumber="31">ABDOMEN</smallCapsWord>
. Shining green-black dorsally, weakly dusted; shining blue-violet laterally; tergites 24 ventrally whitish pollinose; setae and hind-marginal bristles black, short; sternites 3 and 4 of male abdomen with tuft of mainly black long hairs. Hypopygium (
<figureCitation id="38BF6BE4E926FF84C09AC8C1FC9B79FF" box="[736,857,677,704]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,845,1887]" captionTargetId="figure-189@29.[189,1398,845,1889]" captionTargetPageId="29" captionText="Fig. 9. Thinophilus longicercus sp. nov., holotype, ♂ (RBINS). A. Habitus. B. Fore leg, posterior view. C. Hind femur. D. Wing. E. Hypopygium after maceration with basal half of cercus, lateral view. F. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="30" pageNumber="31">Fig. 9EF</figureCitation>
) black with black appendages; epandrial lobe reduced to small subtriangular projection, glabrous; hypandrium short, fused with epandrium, apically concave; phallosoma narrow, weakly projected; phallus long and simple, concealed; surstylus flat and broad, at middle slightly wider than at base (lateral view), with several short inner and ventral bristles at base, with few short outer bristles at apex; cerci dorsally separated, very long, extending to base of abdomen, broad on basal third, filiform distally, with marginal bristles (
<figureCitation id="38BF6BE4E926FF84C664C931FBB8784F" box="[1054,1146,853,880]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,845,1887]" captionTargetId="figure-189@29.[189,1398,845,1889]" captionTargetPageId="29" captionText="Fig. 9. Thinophilus longicercus sp. nov., holotype, ♂ (RBINS). A. Habitus. B. Fore leg, posterior view. C. Hind femur. D. Wing. E. Hypopygium after maceration with basal half of cercus, lateral view. F. Distal appendages of hypopygium after maceration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8110568" httpUri="https://zenodo.org/record/8110568/files/figure.png" pageId="30" pageNumber="31">Fig. 9E</figureCitation>
; distal half of cercus broken).
</paragraph>
<paragraph id="A03B7761E926FF84C290C9DEFE8478EB" blockId="30.[189,326,954,1015]" box="[234,326,954,980]" pageId="30" pageNumber="31">
<emphasis id="92F0AB73E926FF84C290C9DEFE8478EB" bold="true" box="[234,326,954,980]" pageId="30" pageNumber="31">Female</emphasis>
</paragraph>
<paragraph id="A03B7761E926FF84C2C7C9B9FEFB78C8" blockId="30.[189,326,954,1015]" box="[189,313,989,1015]" pageId="30" pageNumber="31">Unknown.</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE926FF84C2C7CE7BFC2A7FEE" pageId="30" pageNumber="31" type="discussion">
<paragraph id="A03B7761E926FF84C2C7CE7BFEC67F06" blockId="30.[189,260,1055,1081]" box="[189,260,1055,1081]" pageId="30" pageNumber="31">
<heading id="FB73C00DE926FF84C2C7CE7BFEC67F06" bold="true" box="[189,260,1055,1081]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="92F0AB73E926FF84C2C7CE7BFEC67F06" bold="true" box="[189,260,1055,1081]" pageId="30" pageNumber="31">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E926FF84C2C7CE29FC2A7FEE" blockId="30.[189,1399,1101,1233]" pageId="30" pageNumber="31">
The
<typeStatus id="7F3FC9C3E926FF84C28ECE29FE997F58" box="[244,347,1101,1127]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
bears an identification label by P. Vanschuytbroeck, “
<taxonomicName id="67840CE2E926FF84C19CCE29FAF37F58" authority="Loew" authorityName="Loew" authorityYear="1857" box="[998,1329,1101,1127]" class="Insecta" family="Dolichopodidae" genus="Hydrophorus" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="bisetus">
<emphasis id="92F0AB73E926FF84C19CCE29FB1C7F58" box="[998,1246,1101,1127]" italics="true" pageId="30" pageNumber="31">Hydrophorus bisetus</emphasis>
Loew
</taxonomicName>
”. The latter species was reported from
<collectingCountry id="D89337F1E926FF84C03BCE14FD0D7FB5" box="[577,719,1136,1162]" name="Madagascar" pageId="30" pageNumber="31">Madagascar</collectingCountry>
from
<specimenCount id="B682BCE8E926FF84C162CE15FC6B7FB5" box="[792,937,1136,1163]" pageId="30" pageNumber="31" type="generic">4 specimens</specimenCount>
(
<bibRefCitation id="C4150A90E926FF84C1C1CE14FB0B7FB4" author="Vanschuytbroeck P." box="[955,1225,1136,1163]" pageId="30" pageNumber="31" pagination="1 - 7" refId="ref26297" refString="Vanschuytbroeck P. 1957. Dolichopodidae de Madagascar (Diptera, Brachycera, Orthorrhapha). Bulletin de l'Institut royal des Sciences naturelles de Belgique 33 (25): 1 - 7." type="journal article" year="1957">Vanschuytbroeck 1957</bibRefCitation>
), but only
<specimenCount id="B682BCE8E926FF84C736CE15FF357F91" pageId="30" pageNumber="31" type="male">one male</specimenCount>
was found in RBINS collection, belonging to the new species described here.
<taxonomicName id="67840CE2E926FF84C6FACEF0FAB47F91" authorityName="Loew" authorityYear="1857" box="[1152,1398,1172,1198]" class="Insecta" family="Dolichopodidae" genus="Hydrophorus" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="bisetus">
<emphasis id="92F0AB73E926FF84C6FACEF0FAB47F91" box="[1152,1398,1172,1198]" italics="true" pageId="30" pageNumber="31">Hydrophorus bisetus</emphasis>
</taxonomicName>
was excluded from the Afrotropical Region by
<bibRefCitation id="C4150A90E926FF84C09CCED3FC267FEE" author="Dyte C. E. &amp; Smith K. G. V." box="[742,996,1207,1233]" pageId="30" pageNumber="31" pagination="443 - 463" refId="ref24826" refString="Dyte C. E. &amp; Smith K. G. V. 1980. 33. Family Dolichopodidae: 443 - 463. In: Crosskey R. W. (ed.) Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London." type="book chapter" year="1980">Dyte &amp; Smith (1980)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>