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<document id="8BA4F012ABAA7949A818DAB143C9DB75" ID-CLB-Dataset="57399" ID-DOI="10.5852/ejt.2023.878.2153" ID-GBIF-Dataset="0dfb60c4-1de3-4062-9443-15b55e7e0c88" ID-ISSN="2118-9773" ID-Zenodo-Dep="8110550" ID-ZooBank="07EFE4B6-4184-41C7-82A5-747BF67BA093" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1688395572616" checkinUser="felipe" docAuthor="Grichanov, Igor Ya." docDate="2023" docId="282DC677E910FFB0C021CB8CFC647EAB" docLanguage="en" docName="ejt-878_1-52.pdf" docOrigin="European Journal of Taxonomy 878" docSource="http://zoobank.org/07efe4b6-4184-41c7-82a5-747bf67ba093" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Thinophilus medvedevi Grichanov 2023, sp. nov." docType="treatment" docUuid="EC22CD6D-73BB-4F90-86A9-D84A2BEB8E8A" docUuidSource="ZooBank" docVersion="12" lastPageNumber="43" masterDocId="D414BE0FE938FF9AC27ACA64FFC27B3F" masterDocTitle="A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="41" updateTime="1699533079317" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FF74E7B4A667A2DDCDF58B1F8AB2887A">A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species</mods:title>
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<mods:name id="58A90605BCF6C95726227282B2F6B537" type="personal">
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<mods:roleTerm id="433D0B95920E8AF6882329D2C5FAF3C7">Author</mods:roleTerm>
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<mods:namePart id="F585090642B635B465F04DF786DACDFE">Grichanov, Igor Ya.</mods:namePart>
<mods:affiliation id="04B5CA92507D820896B61CE143755727">All-Russian Institute of Plant Protection, Podbelskogo 3, St Petersburg, Pushkin, 196608, Russia.</mods:affiliation>
<mods:nameIdentifier id="52A52582764CC267F60910D51AFCC2A1" type="email">grichanov@mail.ru</mods:nameIdentifier>
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<mods:title id="F4F70274987AB3235566685B15868391">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7C65F3E3495EE967E3ADE25F5ACF45EF">
<mods:date id="2A2A46D38D1ABC7D385173D5AFBBC8E9">2023</mods:date>
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<mods:number id="BAF304EBD733E6B9D8A39A33FD9D4548">2023-07-03</mods:number>
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<mods:number id="DE35501A7E3A36AA33B0594982BF8E1C">878</mods:number>
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<mods:classification id="76383A3783714929097C28182D96F8B5">journal article</mods:classification>
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<mods:identifier id="554A8F94B8153DBA466B56279EBBB4B3" type="DOI">10.5852/ejt.2023.878.2153</mods:identifier>
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<treatment id="282DC677E910FFB0C021CB8CFC647EAB" ID-DOI="http://doi.org/10.5281/zenodo.8109797" ID-GBIF-Taxon="210883689" ID-Zenodo-Dep="8109797" LSID="urn:lsid:zoobank.org:act:EC22CD6D-73BB-4F90-86A9-D84A2BEB8E8A" httpUri="http://treatment.plazi.org/id/282DC677E910FFB0C021CB8CFC647EAB" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="E89E24EAE910FFB2C021CB8CFC1B793C" box="[603,985,488,515]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="A03B7761E910FFB2C021CB8CFC1B793C" blockId="40.[354,1235,488,585]" box="[603,985,488,515]" pageId="40" pageNumber="41">
<heading id="FB73C00DE910FFB2C021CB8CFC1B793C" box="[603,985,488,515]" centered="true" fontSize="11" level="2" pageId="40" pageNumber="41" reason="2">
<taxonomicName id="67840CE2E910FFB2C021CB8CFCB5793C" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[603,887,488,515]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="medvedevi" status="sp. nov.">
<emphasis id="92F0AB73E910FFB2C021CB8CFCB5793C" bold="true" box="[603,887,488,515]" italics="true" pageId="40" pageNumber="41">Thinophilus medvedevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E910FFB2C105CB8DFC1B793C" box="[895,985,489,515]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE910FFB2C318C868FB117919" box="[354,1235,524,550]" pageId="40" pageNumber="41" type="description">
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<uri id="D4157B63E910FFB2C318C868FB117919" box="[354,1235,524,550]" pageId="40" pageNumber="41">
urn:lsid:zoobank.org:act:
<uuid id="D4224DB4E910FFB2C0F8C868FB117919" box="[642,1235,524,550]" pageId="40" pageNumber="41">EC22CD6D-73BB-4F90-86A9-D84A2BEB8E8A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE910FFB2C08AC84BFC817976" box="[752,835,559,585]" pageId="40" pageNumber="41" type="description">
<paragraph id="A03B7761E910FFB2C08AC84BFC817976" blockId="40.[354,1235,488,585]" box="[752,835,559,585]" pageId="40" pageNumber="41">
<figureCitation id="38BF6BE4E910FFB2C08AC84BFC817976" box="[752,835,559,585]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="40" pageNumber="41">Fig. 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE910FFB2C2C7C814FE9178EC" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="A03B7761E910FFB2C2C7C814FEF579B5" blockId="40.[189,311,624,650]" box="[189,311,624,650]" pageId="40" pageNumber="41">
<heading id="FB73C00DE910FFB2C2C7C814FEF579B5" bold="true" box="[189,311,624,650]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="92F0AB73E910FFB2C2C7C814FEF579B5" bold="true" box="[189,311,624,650]" pageId="40" pageNumber="41">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E910FFB2C2C7C8FBFE9178EC" blockId="40.[189,1400,671,979]" pageId="40" pageNumber="41">
<taxonomicName id="67840CE2E910FFB2C2C7C8FBFE107986" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[189,466,671,697]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="medvedevi" status="sp. nov.">
<emphasis id="92F0AB73E910FFB2C2C7C8FBFE107986" box="[189,466,671,697]" italics="true" pageId="40" pageNumber="41">Thinophilus medvedevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E910FFB2C3A6C8C4FDFB7985" box="[476,569,672,698]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
is very close in size and habitus to
<taxonomicName id="67840CE2E910FFB2C18BC8C4FB597986" authorityName="Grichanov" authorityYear="2023" box="[1009,1179,671,697]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="quadrisetus">
<emphasis id="92F0AB73E910FFB2C18BC8C4FB597986" box="[1009,1179,671,697]" italics="true" pageId="40" pageNumber="41">T. quadrisetus</emphasis>
</taxonomicName>
known by female
<typeStatus id="7F3FC9C3E910FFB2C2C7C8A6FEE679E3" box="[189,292,706,732]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
from
<collectingCountry id="D89337F1E910FFB2C314C8A6FE1579E3" box="[366,471,706,732]" name="Tanzania" pageId="40" pageNumber="41">Tanzania</collectingCountry>
(
<collectingRegion id="6240B983E910FFB2C392C8A7FD5079E3" box="[488,658,706,733]" country="Tanzania" name="Dar es Salaam" pageId="40" pageNumber="41">Dar Es Salam</collectingRegion>
), differing from the latter in lateral bristles on scutellum, ⅓ to ½ as long as median bristles. The scutellum of
<taxonomicName id="67840CE2E910FFB2C081C882FC66783F" authorityName="Grichanov" authorityYear="2023" box="[763,932,742,768]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="quadrisetus">
<emphasis id="92F0AB73E910FFB2C081C882FC66783F" box="[763,932,742,768]" italics="true" pageId="40" pageNumber="41">T. quadrisetus</emphasis>
</taxonomicName>
bears two pairs of scutellars of almost equal length (
<bibRefCitation id="C4150A90E910FFB2C327C96DFE2D781C" author="Parent O." box="[349,495,777,803]" pageId="40" pageNumber="41" pagination="317 - 327" refId="ref25859" refString="Parent O. 1936. Dipteres dolichopodides. Nouvelle contribution a la faune ethiopienne. Revue de Zoologie et de Botanique africaines 28: 317 - 327." type="journal article" year="1936">Parent 1936</bibRefCitation>
). There are some fine differences between the species in colour characters and leg setation that may relate with individual variability.
<taxonomicName id="67840CE2E910FFB2C137C948FBA37879" authorityName="Grichanov" authorityYear="2023" box="[845,1121,812,838]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="medvedevi">
<emphasis id="92F0AB73E910FFB2C137C948FBA37879" box="[845,1121,812,838]" italics="true" pageId="40" pageNumber="41">Thinophilus medvedevi</emphasis>
</taxonomicName>
keys to
<taxonomicName id="67840CE2E910FFB2C6B0C949FABA7879" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[1226,1400,812,838]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="subpalpatus" status="sp. nov.">
<emphasis id="92F0AB73E910FFB2C6B0C949FABA7879" box="[1226,1400,812,838]" italics="true" pageId="40" pageNumber="41">T. subpalpatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E910FFB2C2C7C934FED87855" box="[189,282,848,874]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, differing in mid femur with rows of ventral setae, half as long as femur height; tarsomere 5 of all tarsi deep black; tarsomeres 14 of fore and mid tarsi yellow. The male of
<taxonomicName id="67840CE2E910FFB2C619C917FAD378B3" authorityName="Grichanov" authorityYear="2023" box="[1123,1297,882,908]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="subpalpatus">
<emphasis id="92F0AB73E910FFB2C619C917FAD378B3" box="[1123,1297,882,908]" italics="true" pageId="40" pageNumber="41">T. subpalpatus</emphasis>
</taxonomicName>
has mid femur with row of ventral setae on distal half, 2 × as long as femur height; tarsi gradually darkened towards tips.
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE910FFB2C2C7C99EFC307F7C" box="[189,1010,1018,1091]" pageId="40" pageNumber="41" type="etymology">
<paragraph id="A03B7761E910FFB2C2C7C99EFE877F2B" blockId="40.[189,325,1018,1044]" box="[189,325,1018,1044]" pageId="40" pageNumber="41">
<heading id="FB73C00DE910FFB2C2C7C99EFE877F2B" bold="true" box="[189,325,1018,1044]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="92F0AB73E910FFB2C2C7C99EFE877F2B" bold="true" box="[189,325,1018,1044]" pageId="40" pageNumber="41">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E910FFB2C2C7CE4DFC307F7C" blockId="40.[189,1010,1065,1091]" box="[189,1010,1065,1091]" pageId="40" pageNumber="41">The species is named after the collector, Andrey Medvedev (ZMUM).</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE910FFB2C2C7CE0EFBA97EBD" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="A03B7761E910FFB2C2C7CE0EFE6F7FBB" blockId="40.[189,429,1130,1156]" box="[189,429,1130,1156]" pageId="40" pageNumber="41">
<heading id="FB73C00DE910FFB2C2C7CE0EFE6F7FBB" bold="true" box="[189,429,1130,1156]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="92F0AB73E910FFB2C2C7CE0EFE6F7FBB" bold="true" box="[189,429,1130,1156]" pageId="40" pageNumber="41">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E910FFB2C290CEFDFEAA7FC5" blockId="40.[189,1399,1177,1274]" pageId="40" pageNumber="41">
<materialsCitation id="10EC7D3CE910FFB2C290CEFDFEAA7FC5" ID-GBIF-Occurrence="4145165318" collectingDate="2012-11-13" collectionCode="ZMUM" collectorName="A. Medvedev" country="Madagascar" latitude="-23.12" location="Ifaty" longLatPrecision="754" longitude="43.37" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="92F0AB73E910FFB2C290CEFDFE9F7F8C" bold="true" box="[234,349,1177,1203]" pageId="40" pageNumber="41">
<typeStatus id="7F3FC9C3E910FFB2C290CEFDFE9F7F8C" box="[234,349,1177,1203]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="D89337F1E910FFB2C2C7CED8FE4C7FE9" box="[189,398,1212,1238]" name="Madagascar" pageId="40" pageNumber="41">MADAGASCAR</collectingCountry>
<specimenCount id="B682BCE8E910FFB2C3D6CED9FE077FE8" box="[428,453,1213,1239]" pageId="40" pageNumber="41" type="male"></specimenCount>
; “
<collectingRegion id="6240B983E910FFB2C3A4CED8FD547FE9" box="[478,662,1212,1238]" country="Madagascar" name="Toliara" pageId="40" pageNumber="41">Toliara Region</collectingRegion>
, near
<location id="A55B21BAE910FFB2C09BCED8FCDE7FE9" LSID="urn:lsid:plazi:treatment:282DC677E910FFB0C021CB8CFC647EAB:A55B21BAE910FFB2C09BCED8FCDE7FE9" box="[737,796,1212,1238]" country="Madagascar" latitude="-23.12" longLatPrecision="754" longitude="43.37" name="Ifaty" pageId="40" pageNumber="41" stateProvince="Toliara">Ifaty</location>
;
<geoCoordinate id="C5B011A6E910FFB2C150CED8FC567FE9" box="[810,916,1212,1238]" degrees="23.12" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="555" value="-23.12">23.12° S</geoCoordinate>
,
<geoCoordinate id="C5B011A6E910FFB2C1DECED8FBD27FE8" box="[932,1040,1212,1239]" degrees="43.37" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="555" value="43.37">43.37° E</geoCoordinate>
;
<date id="D43A51A1E910FFB2C665CED8FB057FE9" box="[1055,1223,1212,1239]" pageId="40" pageNumber="41" value="2012-11-13">
<collectingDate id="C47EA849E910FFB2C665CED8FB057FE9" box="[1055,1223,1212,1239]" pageId="40" pageNumber="41" value="2012-11-13">13 Nov. 2012</collectingDate>
</date>
;
<collectorName id="0D7112B7E910FFB2C6AECED8FAB57FE9" box="[1236,1399,1212,1238]" pageId="40" pageNumber="41">A. Medvedev</collectorName>
leg.”;
<collectionCode id="C695EFA4E910FFB2C37FCE84FEA67FC5" box="[261,356,1248,1274]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="40" pageNumber="41">ZMUM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="A03B7761E910FFB2C290CF45FBA97EBD" blockId="40.[189,1398,1313,1410]" pageId="40" pageNumber="41">
<materialsCitation id="10EC7D3CE910FFB2C290CF45FB957E61" ID-GBIF-Occurrence="4145165346" collectingDate="2012-11-13" collectionCode="ZMUM" collectorName="A. Medvedev" country="Madagascar" latitude="-23.12" location="Ifaty" longLatPrecision="754" longitude="43.37" pageId="40" pageNumber="41" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="92F0AB73E910FFB2C290CF45FEAB7E04" bold="true" box="[234,361,1313,1339]" pageId="40" pageNumber="41">
<typeStatus id="7F3FC9C3E910FFB2C290CF45FEAB7E04" box="[234,361,1313,1339]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="D89337F1E910FFB2C2C7CF20FE4C7E61" box="[189,398,1348,1374]" name="Madagascar" pageId="40" pageNumber="41">MADAGASCAR</collectingCountry>
<specimenCount id="B682BCE8E910FFB2C3D0CF20FE197E61" box="[426,475,1348,1374]" pageId="40" pageNumber="41" type="male">1 ♂</specimenCount>
,
<specimenCount id="B682BCE8E910FFB2C393CF20FDEE7E61" box="[489,556,1348,1374]" pageId="40" pageNumber="41" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="C695EFA4E910FFB2C186CF20FB957E61" box="[1020,1111,1348,1374]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="40" pageNumber="41">ZMUM</collectionCode>
</materialsCitation>
<materialsCitation id="10EC7D3CE910FFB2C609CF20FBA57EBD" ID-GBIF-Occurrence="4145165305" collectingDate="2012-11-12" collectingDateMax="2012-11-19" collectingDateMin="2012-11-12" collectionCode="ZMUM" collectorName="A. Medvedev" country="Madagascar" latitude="-23.2" location="Toliara Region" longLatPrecision="754" longitude="43.62" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="paratype">
<specimenCount id="B682BCE8E910FFB2C609CF20FB667E61" box="[1139,1188,1348,1374]" pageId="40" pageNumber="41" type="male">1 ♂</specimenCount>
; “
<collectingRegion id="6240B983E910FFB2C6C6CF20FAB07E61" box="[1212,1394,1348,1374]" country="Madagascar" name="Toliara" pageId="40" pageNumber="41">Toliara Region</collectingRegion>
, Toliara env.;
<geoCoordinate id="C5B011A6E910FFB2C32CCF03FE7F7EBE" box="[342,445,1383,1409]" degrees="23.20" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="555" value="-23.2">23.20° S</geoCoordinate>
,
<geoCoordinate id="C5B011A6E910FFB2C3B0CF03FDF67EBD" box="[458,564,1383,1410]" degrees="43.62" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="555" value="43.62">43.62° E</geoCoordinate>
;
<date id="D43A51A1E910FFB2C03BCF03FCCD7EBE" box="[577,783,1383,1410]" pageId="40" pageNumber="41" value="2012-11-12" valueMax="2012-11-19" valueMin="2012-11-12">
<collectingDate id="C47EA849E910FFB2C03BCF03FCCD7EBE" box="[577,783,1383,1410]" pageId="40" pageNumber="41" value="2012-11-12" valueMax="2012-11-19" valueMin="2012-11-12">1219 Nov. 2012</collectingDate>
</date>
;
<collectorName id="0D7112B7E910FFB2C163CF03FC7B7EBE" box="[793,953,1383,1409]" pageId="40" pageNumber="41">A. Medvedev</collectorName>
leg.”;
<collectionCode id="C695EFA4E910FFB2C672CF0CFBA57EBD" box="[1032,1127,1384,1410]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="40" pageNumber="41">ZMUM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E89E24EAE910FFB0C2C7CFCCFC647EAB" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="description">
<paragraph id="A03B7761E910FFB2C2C7CFCCFE8D7EFD" blockId="40.[189,335,1448,1474]" box="[189,335,1448,1474]" pageId="40" pageNumber="41">
<heading id="FB73C00DE910FFB2C2C7CFCCFE8D7EFD" bold="true" box="[189,335,1448,1474]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="92F0AB73E910FFB2C2C7CFCCFE8D7EFD" bold="true" box="[189,335,1448,1474]" pageId="40" pageNumber="41">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A03B7761E910FFB2C290CFB3FE6C7ECE" blockId="40.[189,1399,1495,1557]" box="[234,430,1495,1521]" pageId="40" pageNumber="41">
<emphasis id="92F0AB73E910FFB2C290CFB3FEE87ECE" bold="true" box="[234,298,1495,1521]" pageId="40" pageNumber="41">Male</emphasis>
(
<figureCitation id="38BF6BE4E910FFB2C340CFB3FE677ECE" box="[314,421,1495,1521]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="40" pageNumber="41">Fig. 14A</figureCitation>
)
</paragraph>
<paragraph id="A03B7761E910FFB2C2C7CF9FFAB57D2A" blockId="40.[189,1399,1495,1557]" box="[189,1399,1530,1557]" pageId="40" pageNumber="41">
<smallCapsWord id="A6DDE1BDE910FFB2C2C7CF9FFEAC7D2C" baselines="1550,1550" box="[189,366,1531,1557]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="40" pageNumber="41">MEASUREMENTS</smallCapsWord>
. Body length
<quantity id="677CDA84E910FFB2C068CF9EFDAC7D2A" box="[530,622,1530,1557]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="40" pageNumber="41" unit="mm" value="5.8">5.8 mm</quantity>
; antenna length
<quantity id="677CDA84E910FFB2C151CF9EFC4A7D2A" box="[811,904,1530,1557]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="40" pageNumber="41" unit="mm" value="0.9">0.9 mm</quantity>
; wing length 5.0 mm; wing width
<quantity id="677CDA84E910FFB2C76CCF9EFAB07D2A" box="[1302,1394,1530,1557]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="40" pageNumber="41" unit="mm" value="1.6">1.6 mm</quantity>
.
</paragraph>
<paragraph id="A03B7761E910FFB2C2C7CC58FC687C4F" blockId="40.[187,1400,1596,1904]" pageId="40" pageNumber="41">
<smallCapsWord id="A6DDE1BDE910FFB2C2C7CC58FF3E7D6B" baselines="1615,1615" box="[189,252,1596,1622]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="40" pageNumber="41">HEAD</smallCapsWord>
(
<figureCitation id="38BF6BE4E910FFB2C374CC58FEBA7D69" box="[270,376,1596,1622]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="40" pageNumber="41">Fig. 14B</figureCitation>
). Postcranium blackish blue, white pollinose; frons, face and clypeus green-violet, grey pollinose; face under antennae 2 × as wide as height of postpedicel; clypeus half as long as epistoma, about 2 × as wide as long; palp yellow, bearing black bristly hairs; proboscis dark brown; 2 diverging ocellars; 1 strong vertical, 1 postvertical, stronger and longer than, and not in row with upper postoculars; upper postoculars iniseriate, black; middle and lower postoculars multiseriate, white, long; antenna almost entirely orange-yellow (
<figureCitation id="38BF6BE4E910FFB2C04FCC88FD667C39" box="[565,676,1772,1798]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="40" pageNumber="41">Fig. 14C</figureCitation>
); scape subtriangular; pedicel simple, convex on inner side; postpedicel browned at base of arista, rounded, with short pubescence, slightly higher than long (19/15); arista-like stylus dorsal, black and thick basally, white and thin distally, shortly pubescent; length ratio of scape to pedicel to postpedicel to stylus, 0.14/0.07/0.15/0.59.
</paragraph>
<paragraph id="A03B7761E911FFB3C2C7CB6DFB977A56" blockId="41.[189,1399,265,362]" pageId="41" pageNumber="42">
<smallCapsWord id="A6DDE1BDE911FFB3C2C7CB6DFEDB7A1E" baselines="284,284" box="[189,281,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="41" pageNumber="42">THORAX</smallCapsWord>
. Monochrome, black, whitish grey pollinose. No acrostichals; 7 dorsocentrals decreasing in length anteriorly; scutellum with 2 strong marginals and 2 laterals, ⅓ to ½ as long as median bristles; about 4 upper and about 11 lower, white propleural bristles of different length.
</paragraph>
<paragraph id="A03B7761E911FFB3C2C7CBF5FE637ACE" blockId="41.[189,1399,401,498]" pageId="41" pageNumber="42">
<smallCapsWord id="A6DDE1BDE911FFB3C2C7CBF5FF347A96" baselines="420,420" box="[189,246,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="41" pageNumber="42">LEGS</smallCapsWord>
. Fore coxa mostly yellow, black at base; mid and hind coxae black, yellow at apex, whitish grey pollinose; legs mostly yellow; segment 5 almost entirely black; hind tarsomeres 13 dark at apex; hind tarsomere 4 brown.
</paragraph>
<caption id="F4FB27E9E911FFB3C2C7CDECFBB27CD7" ID-DOI="http://doi.org/10.5281/zenodo.8110578" ID-Zenodo-Dep="8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="41" pageNumber="42" startId="41.[189,232,1928,1954]" targetBox="[242,1345,597,1888]" targetPageId="41">
<paragraph id="A03B7761E911FFB3C2C7CDECFBB27CD7" blockId="41.[188,1399,1928,2025]" pageId="41" pageNumber="42">
<emphasis id="92F0AB73E911FFB3C2C7CDECFEDC7C9D" bold="true" box="[189,286,1928,1954]" pageId="41" pageNumber="42">Fig. 14.</emphasis>
<taxonomicName id="67840CE2E911FFB3C353CDECFDFD7C9D" authority="Grichanov, 2023" authorityName="Grichanov" authorityYear="2023" box="[297,575,1928,1954]" class="Insecta" family="Dolichopodidae" genus="Thinophilus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="medvedevi" status="sp. nov.">
<emphasis id="92F0AB73E911FFB3C353CDECFDFD7C9D" box="[297,575,1928,1954]" italics="true" pageId="41" pageNumber="42">Thinophilus medvedevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89C31608E911FFB3C030CDECFD6A7C9D" box="[586,680,1928,1954]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="92F0AB73E911FFB3C0C8CDECFD2B7C9D" bold="true" box="[690,745,1928,1954]" pageId="41" pageNumber="42">AE</emphasis>
. Holotype, ♂ (ZMUM).
<emphasis id="92F0AB73E911FFB3C667CDECFB967C9D" bold="true" box="[1053,1108,1928,1954]" pageId="41" pageNumber="42">FG</emphasis>
. Paratype, ♂ (ZMUM).
<emphasis id="92F0AB73E911FFB3C2C7CDCFFF107CFA" bold="true" box="[189,210,1963,1989]" pageId="41" pageNumber="42">A</emphasis>
. Habitus.
<emphasis id="92F0AB73E911FFB3C33DCDCFFE997CFA" bold="true" box="[327,347,1963,1989]" pageId="41" pageNumber="42">B</emphasis>
. Head.
<emphasis id="92F0AB73E911FFB3C3C9CDCFFE0A7CFA" bold="true" box="[435,456,1963,1989]" pageId="41" pageNumber="42">C</emphasis>
. Antenna.
<emphasis id="92F0AB73E911FFB3C039CDCFFD9A7CFA" bold="true" box="[579,600,1963,1989]" pageId="41" pageNumber="42">D</emphasis>
. Mid tarsus, dorsal view.
<emphasis id="92F0AB73E911FFB3C1F8CDCFFC547CFA" bold="true" box="[898,918,1963,1989]" pageId="41" pageNumber="42">E</emphasis>
. Wing.
<emphasis id="92F0AB73E911FFB3C195CDCFFBC37CFA" bold="true" box="[1007,1025,1963,1989]" pageId="41" pageNumber="42">F</emphasis>
. Hypopygium after maceration, lateral view.
<emphasis id="92F0AB73E911FFB3C329CDAAFEA87CD7" bold="true" box="[339,362,1998,2024]" pageId="41" pageNumber="42">G</emphasis>
. Distal appendages of hypopygium after maceration, lateral view.
</paragraph>
</caption>
<paragraph id="A03B7761E912FFB0C2C7CB6DFF2B7AC9" blockId="42.[189,1399,265,502]" pageId="42" pageNumber="43">
<smallCapsWord id="A6DDE1BDE912FFB0C2C7CB6DFF347A1E" baselines="284,284" box="[189,246,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Fore" pageId="42" pageNumber="43">FORE</smallCapsWord>
<smallCapsWord id="A6DDE1BDE912FFB0C37ECB6AFEEE7A1E" baselines="284" box="[260,300,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="42" pageNumber="43">LEG</smallCapsWord>
. Coxa with white setae, with at most 1 dark apical seta; femur slightly thickened, with fine white ventral setae on basal half, with about 5 fine short black anteroventral setae at base, with about 5 posteroventral preapical black setae, as long as femur height; tibia bearing 2 anterodorsal, 2 posterodorsal and 34 apical short setae, devoid of black setulae anteriorly on distal ⅓; tarsomeres weakly thickened, with elongated setulae; tarsomeres 24 with white ventral pile; tarsomere 5 distinctly flattened; length of femur, tibia and tarsal segments (in mm): 1.8/1.57/0.63/0.33/0.2/0.18/ 0.2.
</paragraph>
<paragraph id="A03B7761E912FFB0C2C7C87AFEC279FA" blockId="42.[188,1401,542,709]" pageId="42" pageNumber="43">
<smallCapsWord id="A6DDE1BDE912FFB0C2C7C87AFF2F7909" baselines="561,561" box="[189,237,542,568]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mid" pageId="42" pageNumber="43">MID</smallCapsWord>
<smallCapsWord id="A6DDE1BDE912FFB0C28FC847FEDF7909" baselines="561" box="[245,285,547,566]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="42" pageNumber="43">LEG</smallCapsWord>
. Coxa with many black setae; femur with rather short black ventral setae; 1 anterior preapical bristle, posteroventral row of 5 black preapical setae; tibia bearing 2 anterodorsal; 2 posterodorsal, 1 anteroventral, 12 posteroventral, 4 apical bristles; tarsomere 4 slightly thickened; tarsomere 5 distinctly flattened (
<figureCitation id="38BF6BE4E912FFB0C34BC8E3FE63799E" box="[305,417,647,673]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="42" pageNumber="43">Fig. 14D</figureCitation>
); length of femur, tibia and tarsal segments (in mm): 1.82/1.74/1.05/0.39/0.26/0.19 /0.24.
</paragraph>
<paragraph id="A03B7761E912FFB0C2C7C888FBFB784F" blockId="42.[189,1399,748,880]" pageId="42" pageNumber="43">
<smallCapsWord id="A6DDE1BDE912FFB0C2C7C888FF35783A" baselines="767,768" box="[189,247,748,774]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hind" pageId="42" pageNumber="43">HIND</smallCapsWord>
<smallCapsWord id="A6DDE1BDE912FFB0C378C895FEE8783B" baselines="767" box="[258,298,753,772]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg" pageId="42" pageNumber="43">LEG</smallCapsWord>
. Coxa with 1 black exterior bristle; femur with 23 anterodorsal setae on distal half and 2 preapical posteroventral short setae, without ventrals; tibia bearing 3 anterodorsal, 3 posterodorsal, 2 ventral bristles, 3 apicals; tarsomere 4 slightly thickened; tarsomere 5 distinctly flattened; length of femur, tibia and tarsal segments (in mm): 2.1/2.26/0.66/0.61/0.37/0.25/0.27.
</paragraph>
<paragraph id="A03B7761E912FFB0C2C7C9F3FB457F25" blockId="42.[189,1399,919,1050]" pageId="42" pageNumber="43">
<smallCapsWord id="A6DDE1BDE912FFB0C2C7C9F3FF3C7890" baselines="938,938" box="[189,254,919,945]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Wing" pageId="42" pageNumber="43">WING</smallCapsWord>
(
<figureCitation id="38BF6BE4E912FFB0C36BC9F3FEBF788E" box="[273,381,919,945]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="42" pageNumber="43">Fig. 14E</figureCitation>
). Hyaline, without darker shades; veins yellow-brown, more yellowish at base; distal part of M
<subScript id="3C007524E912FFB0C355C9A9FE8878E4" attach="left" box="[303,330,973,987]" fontSize="6" pageId="42" pageNumber="43">1+2</subScript>
convex; tip of R
<subScript id="3C007524E912FFB0C06DC9A9FDF378E4" attach="left" box="[535,561,973,987]" fontSize="6" pageId="42" pageNumber="43">4+5</subScript>
parallel with M
<subScript id="3C007524E912FFB0C089C9A9FCCC78E4" attach="left" box="[755,782,973,987]" fontSize="6" pageId="42" pageNumber="43">1+2</subScript>
; ratio of part of costa between R
<subScript id="3C007524E912FFB0C6E6C9A9FB7578E4" attach="left" box="[1180,1207,973,987]" fontSize="6" pageId="42" pageNumber="43">2+3</subScript>
and R
<subScript id="3C007524E912FFB0C77DC9A9FAE378E4" attach="left" box="[1287,1313,973,987]" fontSize="6" pageId="42" pageNumber="43">4+5</subScript>
to that between R
<subScript id="3C007524E912FFB0C340C994FE9778C1" attach="left" box="[314,341,1008,1022]" fontSize="6" pageId="42" pageNumber="43">4+5</subScript>
and M
<subScript id="3C007524E912FFB0C3D3C994FE0678C1" attach="left" box="[425,452,1008,1022]" fontSize="6" pageId="42" pageNumber="43">1+2</subScript>
(in mm), 0.42/0.30; crossvein dm-m straight; ratio of dm-m to distal part of M
<subScript id="3C007524E912FFB0C71DC994FAB278C1" attach="right" box="[1383,1392,1008,1022]" fontSize="6" pageId="42" pageNumber="43">4</subScript>
, 0.43/0.61; anal vein distinct; halter yellow; lower calypter yellow, with white cilia.
</paragraph>
<paragraph id="A03B7761E912FFB0C2C7CE26FD277E33" blockId="42.[187,1399,1090,1292]" pageId="42" pageNumber="43">
<smallCapsWord id="A6DDE1BDE912FFB0C2C7CE26FEEE7F65" baselines="1109,1109" box="[189,300,1090,1116]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="42" pageNumber="43">ABDOMEN</smallCapsWord>
. With setae and hind-marginal bristles on tergites black, short; sternites with very short setulae. Hypopygium (
<figureCitation id="38BF6BE4E912FFB0C316CE01FE1A7F40" box="[364,472,1125,1151]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="42" pageNumber="43">Fig. 14F</figureCitation>
) black, cercus yellow; epandrial lobe fingerlike, with strong apical bristle; hypandrium short, apically concave; phallosoma broad, split at apex, not reaching apex of surstyli; phallus coiled, long and flat, band-like; surstylus slightly curved, widened distally (
<figureCitation id="38BF6BE4E912FFB0C6C5CECFFAEC7FF9" box="[1215,1326,1195,1222]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="42" pageNumber="43">Fig. 14G</figureCitation>
) with pointed apex, several short preapical bristles and setae; cerci dorsally fused at base, free and narrow distally, with long marginal bristles (
<figureCitation id="38BF6BE4E912FFB0C011CE96FD1A7E33" box="[619,728,1266,1292]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[242,1345,597,1888]" captionTargetId="figure-100@41.[242,1345,597,1889]" captionTargetPageId="41" captionText="Fig. 14. Thinophilus medvedevi sp. nov. AE. Holotype, ♂ (ZMUM). FG. Paratype, ♂ (ZMUM). A. Habitus. B. Head. C. Antenna.D. Mid tarsus, dorsal view. E. Wing.F. Hypopygium after maceration, lateral view. G. Distal appendages of hypopygium after maceration, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8110578" httpUri="https://zenodo.org/record/8110578/files/figure.png" pageId="42" pageNumber="43">Fig. 14G</figureCitation>
).
</paragraph>
<paragraph id="A03B7761E912FFB0C290CF57FE847E72" blockId="42.[189,1398,1331,1428]" box="[234,326,1331,1357]" pageId="42" pageNumber="43">
<emphasis id="92F0AB73E912FFB0C290CF57FE847E72" bold="true" box="[234,326,1331,1357]" pageId="42" pageNumber="43">Female</emphasis>
</paragraph>
<paragraph id="A03B7761E912FFB0C2C7CF32FC647EAB" blockId="42.[189,1398,1331,1428]" pageId="42" pageNumber="43">Similar to male except lacking male secondary sexual characters. Posteroventral preapical setae on fore femur weakly developed; fore tarsus without white ventral pile.</paragraph>
</subSubSection>
</treatment>
</document>