309 lines
28 KiB
XML
309 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.45.95845" ID-Pensoft-Pub="1314-2615-45-17" ID-Pensoft-UUID="22E19A47A21D5DA492B3F6947BA63B3A" ID-ZooBank="2914532A9A22475698F37D885262AA44" ModsDocID="1314-2615-45-17" checkinTime="1675174029439" checkinUser="pensoft" docAuthor="Moutaouakil, Soumia, Boulanouar, Mohamed, Ghamizi, Mohamed, Lips, Josiane & Ferreira, Rodrigo Lopes" docDate="2023" docId="122FB1D9B75B51A9A043BAF8EF9C9668" docLanguage="en" docName="SubterBiol 45: 17-37" docOrigin="Subterranean Biology 45" docPubDate="2023-01-30" docSource="http://dx.doi.org/10.3897/subtbiol.45.95845" docTitle="Castellanethes ougougensis Moutaouakil & Boulanouar 2023, sp. nov." docType="treatment" docUuid="B2EE7107-4D7B-46C2-99C5-1668C3688183" docUuidSource="ZooBank" docVersion="1" id="22E19A47A21D5DA492B3F6947BA63B3A" lastPageNumber="17" masterDocId="22E19A47A21D5DA492B3F6947BA63B3A" masterDocTitle="Two new sympatric cave species of Castellanethes (Isopoda, Oniscidea, Olibrinidae) from Western High Atlas of Morocco" masterLastPageNumber="37" masterPageNumber="17" pageNumber="17" updateTime="1675174029439" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new sympatric cave species of Castellanethes (Isopoda, Oniscidea, Olibrinidae) from Western High Atlas of Morocco</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moutaouakil, Soumia</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5146-3804</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Marrakech - Research center, Marrakech, Morocco & Water, Biodiversity and Climate Change Laboratory, Faculty of Sciences Semlalia, Cadi Ayyad University, Marrakech, Morocco</mods:affiliation>
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<mods:nameIdentifier type="email">moutaouakil.soumia@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Boulanouar, Mohamed</mods:namePart>
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<mods:affiliation>Natural History Museum of Marrakech - Research center, Marrakech, Morocco & Water, Biodiversity and Climate Change Laboratory, Faculty of Sciences Semlalia, Cadi Ayyad University, Marrakech, Morocco</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ghamizi, Mohamed</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9396-4260</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Marrakech - Research center, Marrakech, Morocco & Water, Biodiversity and Climate Change Laboratory, Faculty of Sciences Semlalia, Cadi Ayyad University, Marrakech, Morocco</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lips, Josiane</mods:namePart>
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<mods:affiliation>Ecole Normale Superieure of Marrakech, Cadi Ayyad University, Marrakech, Morocco</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ferreira, Rodrigo Lopes</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3288-4405</mods:nameIdentifier>
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<mods:affiliation>Groupe d'Etudes de Biospeologie, Federation Francaise de Speleologie, Lyon, France</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Subterranean Biology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>45</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>17</mods:start>
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<mods:end>37</mods:end>
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<mods:url>http://dx.doi.org/10.3897/subtbiol.45.95845</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.45.95845</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2615-45-17</mods:identifier>
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<mods:identifier type="Pensoft-UUID">22E19A47A21D5DA492B3F6947BA63B3A</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:B2EE7107-4D7B-46C2-99C5-1668C3688183" httpUri="http://treatment.plazi.org/id/122FB1D9B75B51A9A043BAF8EF9C9668" lastPageNumber="17" pageId="0" pageNumber="17">
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<subSubSection pageId="0" pageNumber="17" type="nomenclature">
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<paragraph pageId="0" pageNumber="17">
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<taxonomicName LSID="https://zoobank.org/B2EE7107-4D7B-46C2-99C5-1668C3688183" authority="Moutaouakil & Boulanouar" authorityName="Moutaouakil & Boulanouar" authorityYear="2023" class="Malacostraca" family="Olibrinidae" genus="Castellanethes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castellanethes ougougensis" order="Isopoda" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="ougougensis" status="sp. nov.">Castellanethes ougougensis Moutaouakil & Boulanouar</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="17">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="17" type="description">
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<paragraph pageId="0" pageNumber="17">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave A adult specimen, dorsal view B, C maxillula D maxilla E maxilliped. Scale bars: 1 mm (A); 0.1 mm (B-E)." figureDoi="10.3897/subtbiol.45.95845.figure2" httpUri="https://binary.pensoft.net/fig/802831" pageId="0" pageNumber="17">Figs 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.45.95845.figure3" httpUri="https://binary.pensoft.net/fig/802832" pageId="0" pageNumber="17">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave A male pleopod 1 B female pleopod 1 exopod C male pleopod 2 D female pleopod 2 exopod E male pleopod 3 F male pleopod 4 G male pleopod 5. Scale bar: 0.1 mm." figureDoi="10.3897/subtbiol.45.95845.figure4" httpUri="https://binary.pensoft.net/fig/802833" pageId="0" pageNumber="17">, 4</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Castellanethes soloisensis from Goran Cave A antenna D antennula. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B antenna E antennula. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C antenna F antennula. Scale bars: 1 mm (A); 500 µm (B, C); 100 µm (D-F)." figureDoi="10.3897/subtbiol.45.95845.figure8" httpUri="https://binary.pensoft.net/fig/802837" pageId="0" pageNumber="17">, 8B, E</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Castellanethes soloisensis from Goran Cave A left mandible B right mandible. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave C left mandible D right mandible. Castellanethes ighousi sp. nov. from Imi Ougoug Cave E left mandible F right mandible. Scale bar: 200 µm (A-F)." figureDoi="10.3897/subtbiol.45.95845.figure9" httpUri="https://binary.pensoft.net/fig/802838" pageId="0" pageNumber="17">, 9C, D</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Castellanethes soloisensis from Goran Cave A telson + uropods D male pleopod 1 G male pleopod 2. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B telson + uropods E male pleopod 1 H male pleopod 2. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C telson + uropods F male pleopod 1 I male pleopod 2. Scale bar: 500 µm (A, C, G); 400 µm (B); 300 µm (D, F, I); 200 µm (E, H)." figureDoi="10.3897/subtbiol.45.95845.figure10" httpUri="https://binary.pensoft.net/fig/802839" pageId="0" pageNumber="17">, 10B, E, H</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Castellanethes soloisensis ♂ specimen from Goran Cave A pereopod 1 B pereopod 6 C pereopod 7. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave D pereopod 1 E pereopod 6 F pereopod 7. Castellanethes ighousi sp. nov. ♂ specimen from Imi Ougoug Cave G pereopod 1 H pereopod 6 I pereopod 7. Scale bar: 1 mm (H, I); 500 µm (A-G)." figureDoi="10.3897/subtbiol.45.95845.figure11" httpUri="https://binary.pensoft.net/fig/802840" pageId="0" pageNumber="17">, 11D-F</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Presentation of the type locality (Imi Ougoug Cave) A entrance of the Imi Ougoug Cave B Imi Ougoug Cave topography C external area at Talmat River D aquatic habitat inside Imi Ougoug Cave E terrestrial habitat F Castellanethes ighousi in a pond G Scolopendromorpha / isopod's predator H Castellanethes ougougensis." figureDoi="10.3897/subtbiol.45.95845.figure12" httpUri="https://binary.pensoft.net/fig/802841" pageId="0" pageNumber="17">, 12H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="17" type="materials_examined">
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<paragraph pageId="0" pageNumber="17">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="17">
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<materialsCitation collectingDate="2019-12-29" collectorName="Moutaouakil S." country="Morocco" elevation="773" location="Agadir-Imi Ougoug Cave" specimenCount="1" specimenCount-male="1" typeStatus="Holotype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="17">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Morocco">Morocco</collectingCountry>
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•
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<specimenCount type="male">1 ♂</specimenCount>
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(mounted on a slide),
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<location LSID="urn:lsid:plazi:treatment:122FB1D9B75B51A9A043BAF8EF9C9668:0D708A3D81960DF1BB1AEF45C1092A62" country="Morocco" name="Agadir-Imi Ougoug Cave">Agadir-Imi Ougoug Cave</location>
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, alt.
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<quantity metricMagnitude="2" metricUnit="m" metricValue="7.7299999999999995" unit="m" value="773.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="7.7299999999999995" unit="m" value="773.0">773 m</elevation>
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</quantity>
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,
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<collectingDate value="2019-12-29">29 Dec 2019</collectingDate>
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,
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<collectorName>Moutaouakil S.</collectorName>
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leg. MHNM ZAI01
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</materialsCitation>
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.
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<materialsCitation collectingDate="2020-02-26" collectorName="Moutaouakil S." country="Morocco" location="Imi Ougoug Cave" specimenCount="2" specimenCount-female="2" typeStatus="Paratypes">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="17">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Morocco">Morocco</collectingCountry>
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•
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<specimenCount type="female">2 ♀</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:122FB1D9B75B51A9A043BAF8EF9C9668:31987C0C1AD7E7CDB93EE00A4935542B" country="Morocco" name="Imi Ougoug Cave">Imi Ougoug Cave</location>
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,
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<collectingDate value="2020-02-26">26 Feb 2020</collectingDate>
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,
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<collectorName>Moutaouakil S.</collectorName>
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leg., MHNM ZAI03 and MHNM ZAI04
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</materialsCitation>
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•
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<materialsCitation collectingDate="2019-10-27" collectorName="Lips J." country="Morocco" location="Ifri N'Tyaline Cave" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:122FB1D9B75B51A9A043BAF8EF9C9668:C02DB6888B956071A80A9A7A96B22019" country="Morocco" name="Ifri N'Tyaline Cave">
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Ifri
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<normalizedToken originalValue="N’Tyaline">N'Tyaline</normalizedToken>
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Cave
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</location>
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,
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<collectingDate value="2019-10-27">27 Oct 2019</collectingDate>
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,
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<collectorName>Lips J.</collectorName>
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leg., MHNM ZAI05
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</materialsCitation>
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•
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<materialsCitation collectingDate="2019-10-31" collectorName="Moutaouakil S." country="Morocco" location="Win Timdouine Cave" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:122FB1D9B75B51A9A043BAF8EF9C9668:5433F0E3170E0498112ABDB70D5229E9" country="Morocco" name="Win Timdouine Cave">Win Timdouine Cave</location>
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,
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<collectingDate value="2019-10-31">31 Oct 2019</collectingDate>
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,
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<collectorName>Moutaouakil S.</collectorName>
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leg., ZAI06
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</materialsCitation>
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•
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<materialsCitation collectingDate="2019-11-04" collectorName="Moutaouakil S." country="Morocco" location="Taghrart Ounakrim Cave" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:122FB1D9B75B51A9A043BAF8EF9C9668:5CC119A35630BE99FD146F5465378017" country="Morocco" name="Taghrart Ounakrim Cave">Taghrart Ounakrim Cave</location>
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,
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<collectingDate value="2019-11-04">04 Nov 2019</collectingDate>
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,
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<collectorName>Moutaouakil S.</collectorName>
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leg., MHNM ZAI07
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</materialsCitation>
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•
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<materialsCitation collectingDate="2020-02-25" collectorName="Moutaouakil S." country="Morocco" location="Tigmi N'Dou Akkal Cave" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:122FB1D9B75B51A9A043BAF8EF9C9668:A86057B51C0B4FC6ED4D601DF55A2205" country="Morocco" name="Tigmi N'Dou Akkal Cave">
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Tigmi
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<normalizedToken originalValue="N’Dou">N'Dou</normalizedToken>
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Akkal Cave
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</location>
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,
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<collectingDate value="2020-02-25">25 Feb 2020</collectingDate>
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,
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<collectorName>Moutaouakil S.</collectorName>
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leg., MHNM ZAI08
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="17" type="diagnosis">
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<paragraph pageId="0" pageNumber="17">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="17">
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Telson with double-rounded tips. Antennula with eight petaliform aesthetascs. Antennal flagellum with six articles. Molar penicil of mandible with two plumose setae. Male pleopod 1 endopod with four wide scales at apex inner margin. Male pleopod 2 endopod thick and about 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as exopod.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="17" type="description">
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<paragraph pageId="0" pageNumber="17">Description.</paragraph>
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<paragraph pageId="0" pageNumber="17">
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Maximum body length: ♂ 5 mm, ♀ 6 mm. Body outline as in Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave A adult specimen, dorsal view B, C maxillula D maxilla E maxilliped. Scale bars: 1 mm (A); 0.1 mm (B-E)." figureDoi="10.3897/subtbiol.45.95845.figure2" httpUri="https://binary.pensoft.net/fig/802831" pageId="0" pageNumber="17">2A</figureCitation>
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. Colourless, dorsal surface tuberculate with tubercles arranged as follows: cephalon with four rows of tubercles; pereonite 1 with three rows of tubercles; pereonites 2-7 with two rows; pleonites 1 and 2 with one row of tubercles each, pleonites 3-5 and telson smooth, the latter about four times wider than long (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Castellanethes soloisensis from Goran Cave A telson + uropods D male pleopod 1 G male pleopod 2. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B telson + uropods E male pleopod 1 H male pleopod 2. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C telson + uropods F male pleopod 1 I male pleopod 2. Scale bar: 500 µm (A, C, G); 400 µm (B); 300 µm (D, F, I); 200 µm (E, H)." figureDoi="10.3897/subtbiol.45.95845.figure10" httpUri="https://binary.pensoft.net/fig/802839" pageId="0" pageNumber="17">10B</figureCitation>
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). Pleon narrower than pereon. Antennula with three articles (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Castellanethes soloisensis from Goran Cave A antenna D antennula. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B antenna E antennula. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C antenna F antennula. Scale bars: 1 mm (A); 500 µm (B, C); 100 µm (D-F)." figureDoi="10.3897/subtbiol.45.95845.figure8" httpUri="https://binary.pensoft.net/fig/802837" pageId="0" pageNumber="17">8E</figureCitation>
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), second article with one row of short setae on outer margin, third article with one row of eight petaliform aesthetascs along outer margin. Antenna long and slender (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Castellanethes soloisensis from Goran Cave A antenna D antennula. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B antenna E antennula. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C antenna F antennula. Scale bars: 1 mm (A); 500 µm (B, C); 100 µm (D-F)." figureDoi="10.3897/subtbiol.45.95845.figure8" httpUri="https://binary.pensoft.net/fig/802837" pageId="0" pageNumber="17">8B</figureCitation>
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), flagellum of six articles. Mandible with molar penicil consisting of two branches (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Castellanethes soloisensis from Goran Cave A left mandible B right mandible. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave C left mandible D right mandible. Castellanethes ighousi sp. nov. from Imi Ougoug Cave E left mandible F right mandible. Scale bar: 200 µm (A-F)." figureDoi="10.3897/subtbiol.45.95845.figure9" httpUri="https://binary.pensoft.net/fig/802838" pageId="0" pageNumber="17">9C, D</figureCitation>
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), basal one more than twice as long as the other; left mandible with 2+1 penicils, right with 1+1 penicils. Maxillula (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave A adult specimen, dorsal view B, C maxillula D maxilla E maxilliped. Scale bars: 1 mm (A); 0.1 mm (B-E)." figureDoi="10.3897/subtbiol.45.95845.figure2" httpUri="https://binary.pensoft.net/fig/802831" pageId="0" pageNumber="17">2B, C</figureCitation>
|
||
) outer endite with 12 simple teeth and one long seta, inner branch without setae, penicils or spines. Maxilla (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave A adult specimen, dorsal view B, C maxillula D maxilla E maxilliped. Scale bars: 1 mm (A); 0.1 mm (B-E)." figureDoi="10.3897/subtbiol.45.95845.figure2" httpUri="https://binary.pensoft.net/fig/802831" pageId="0" pageNumber="17">2D</figureCitation>
|
||
) rounded and covered with thin setae and one row of thick setae. Maxilliped (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave A adult specimen, dorsal view B, C maxillula D maxilla E maxilliped. Scale bars: 1 mm (A); 0.1 mm (B-E)." figureDoi="10.3897/subtbiol.45.95845.figure2" httpUri="https://binary.pensoft.net/fig/802831" pageId="0" pageNumber="17">2E</figureCitation>
|
||
) endite with long apical penicil; epipodite bearing fringe of thin setae on outer margin. Pereopods with long plumose dactylar seta (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Castellanethes ougougensis sp. nov. ♂ specimen from Imi Ougoug Cave A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.45.95845.figure3" httpUri="https://binary.pensoft.net/fig/802832" pageId="0" pageNumber="17">3A-G</figureCitation>
|
||
). Uropods (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Castellanethes soloisensis from Goran Cave A telson + uropods D male pleopod 1 G male pleopod 2. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B telson + uropods E male pleopod 1 H male pleopod 2. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C telson + uropods F male pleopod 1 I male pleopod 2. Scale bar: 500 µm (A, C, G); 400 µm (B); 300 µm (D, F, I); 200 µm (E, H)." figureDoi="10.3897/subtbiol.45.95845.figure10" httpUri="https://binary.pensoft.net/fig/802839" pageId="0" pageNumber="17">10B</figureCitation>
|
||
) with protopod slightly longer than wide; exopod bearing one row of scales on inner margin; endopod wider and longer than exopod.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="17">
|
||
<emphasis bold="true" pageId="0" pageNumber="17">Male.</emphasis>
|
||
Pleopod 1 (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave A male pleopod 1 B female pleopod 1 exopod C male pleopod 2 D female pleopod 2 exopod E male pleopod 3 F male pleopod 4 G male pleopod 5. Scale bar: 0.1 mm." figureDoi="10.3897/subtbiol.45.95845.figure4" httpUri="https://binary.pensoft.net/fig/802833" pageId="0" pageNumber="17">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Castellanethes soloisensis from Goran Cave A telson + uropods D male pleopod 1 G male pleopod 2. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B telson + uropods E male pleopod 1 H male pleopod 2. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C telson + uropods F male pleopod 1 I male pleopod 2. Scale bar: 500 µm (A, C, G); 400 µm (B); 300 µm (D, F, I); 200 µm (E, H)." figureDoi="10.3897/subtbiol.45.95845.figure10" httpUri="https://binary.pensoft.net/fig/802839" pageId="0" pageNumber="17">10E</figureCitation>
|
||
) endopod with distal part pointed bent inwards, bearing four large transversal scales on the inner side; exopod triangular with no scale-setae (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave A male pleopod 1 B female pleopod 1 exopod C male pleopod 2 D female pleopod 2 exopod E male pleopod 3 F male pleopod 4 G male pleopod 5. Scale bar: 0.1 mm." figureDoi="10.3897/subtbiol.45.95845.figure4" httpUri="https://binary.pensoft.net/fig/802833" pageId="0" pageNumber="17">4B</figureCitation>
|
||
). Pleopod 2 (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave A male pleopod 1 B female pleopod 1 exopod C male pleopod 2 D female pleopod 2 exopod E male pleopod 3 F male pleopod 4 G male pleopod 5. Scale bar: 0.1 mm." figureDoi="10.3897/subtbiol.45.95845.figure4" httpUri="https://binary.pensoft.net/fig/802833" pageId="0" pageNumber="17">4C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Castellanethes soloisensis from Goran Cave A telson + uropods D male pleopod 1 G male pleopod 2. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave B telson + uropods E male pleopod 1 H male pleopod 2. Castellanethes ighousi sp. nov. from Imi Ougoug Cave C telson + uropods F male pleopod 1 I male pleopod 2. Scale bar: 500 µm (A, C, G); 400 µm (B); 300 µm (D, F, I); 200 µm (E, H)." figureDoi="10.3897/subtbiol.45.95845.figure10" httpUri="https://binary.pensoft.net/fig/802839" pageId="0" pageNumber="17">10H</figureCitation>
|
||
) endopod thick and about 1/5 as long as exopod; exopod with one to two setae on apex (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Castellanethes ougougensis sp. nov. from Imi Ougoug Cave A male pleopod 1 B female pleopod 1 exopod C male pleopod 2 D female pleopod 2 exopod E male pleopod 3 F male pleopod 4 G male pleopod 5. Scale bar: 0.1 mm." figureDoi="10.3897/subtbiol.45.95845.figure4" httpUri="https://binary.pensoft.net/fig/802833" pageId="0" pageNumber="17">4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/subtbiol.45.95845.figure2" httpUri="https://binary.pensoft.net/fig/802831" pageId="0" pageNumber="17" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="17">
|
||
<emphasis bold="true" pageId="0" pageNumber="17">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Moutaouakil & Boulanouar" authorityYear="2023" class="Malacostraca" family="Olibrinidae" genus="Castellanethes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castellanethes ougougensis" order="Isopoda" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="ougougensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="17">Castellanethes ougougensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. ♂ specimen from Imi Ougoug Cave
|
||
<emphasis bold="true" pageId="0" pageNumber="17">A</emphasis>
|
||
adult specimen, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="17">B, C</emphasis>
|
||
maxillula
|
||
<emphasis bold="true" pageId="0" pageNumber="17">D</emphasis>
|
||
maxilla
|
||
<emphasis bold="true" pageId="0" pageNumber="17">E</emphasis>
|
||
maxilliped. Scale bars: 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="17">A</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="17">B-E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/subtbiol.45.95845.figure3" httpUri="https://binary.pensoft.net/fig/802832" pageId="0" pageNumber="17" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="17">
|
||
<emphasis bold="true" pageId="0" pageNumber="17">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Moutaouakil & Boulanouar" authorityYear="2023" class="Malacostraca" family="Olibrinidae" genus="Castellanethes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castellanethes ougougensis" order="Isopoda" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="ougougensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="17">Castellanethes ougougensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. ♂ specimen from Imi Ougoug Cave
|
||
<emphasis bold="true" pageId="0" pageNumber="17">A</emphasis>
|
||
pereopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="17">B</emphasis>
|
||
pereopod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="17">C</emphasis>
|
||
pereopod 3
|
||
<emphasis bold="true" pageId="0" pageNumber="17">D</emphasis>
|
||
pereopod 4
|
||
<emphasis bold="true" pageId="0" pageNumber="17">E</emphasis>
|
||
pereopod 5
|
||
<emphasis bold="true" pageId="0" pageNumber="17">F</emphasis>
|
||
pereopod 6
|
||
<emphasis bold="true" pageId="0" pageNumber="17">G</emphasis>
|
||
pereopod 7. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/subtbiol.45.95845.figure4" httpUri="https://binary.pensoft.net/fig/802833" pageId="0" pageNumber="17" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="17">
|
||
<emphasis bold="true" pageId="0" pageNumber="17">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Moutaouakil & Boulanouar" authorityYear="2023" class="Malacostraca" family="Olibrinidae" genus="Castellanethes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castellanethes ougougensis" order="Isopoda" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="ougougensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="17">Castellanethes ougougensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Imi Ougoug Cave
|
||
<emphasis bold="true" pageId="0" pageNumber="17">A</emphasis>
|
||
male pleopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="17">B</emphasis>
|
||
female pleopod 1 exopod
|
||
<emphasis bold="true" pageId="0" pageNumber="17">C</emphasis>
|
||
male pleopod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="17">D</emphasis>
|
||
female pleopod 2 exopod
|
||
<emphasis bold="true" pageId="0" pageNumber="17">E</emphasis>
|
||
male pleopod 3
|
||
<emphasis bold="true" pageId="0" pageNumber="17">F</emphasis>
|
||
male pleopod 4
|
||
<emphasis bold="true" pageId="0" pageNumber="17">G</emphasis>
|
||
male pleopod 5. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="17" type="etymology">
|
||
<paragraph pageId="0" pageNumber="17">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="17">The name refers to the cave where the specimens were found first: Imi Ougoug Cave.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |