281 lines
34 KiB
XML
281 lines
34 KiB
XML
<document id="146379AFFE9366A216DD2122D6133724" ID-DOI="10.5281/zenodo.215031" ID-GBIF-Dataset="215f451e-15f2-440d-b4f1-a5ca804ec0cb" ID-ISSN="1175-5326" ID-Zenodo-Dep="215031" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460506771825" checkinUser="plazi" docAuthor="Spinelli, Gustavo R., Marino, Pablo I. & Borkent, Art" docDate="2012" docId="03A7878CE37CFFCA53D2BF84869BFCC6" docLanguage="en" docName="zt03419p052.pdf" docOrigin="Zootaxa 3419" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Forcipomyia (Metaforcipomyia) anniae Spinelli, Marino & Borkent, 2012, n. sp." docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="FF9EFFF4E376FFC65345BD358244FFF1" masterDocTitle="A revision of Biting Midges of the Subgenera Forcipomyia (Metaforcipomyia) and F. (Saliohelea) from Costa Rica (Diptera: Ceratopogonidae)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="11" updateTime="1698312853441" updateUser="plazi">
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<mods:title id="9123F5C9594683EA9C75DD63F3A4A45C">A revision of Biting Midges of the Subgenera Forcipomyia (Metaforcipomyia) and F. (Saliohelea) from Costa Rica (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="805D8605EA94D44BB64AE91D55232ACE">Spinelli, Gustavo R.</mods:namePart>
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<mods:namePart id="09B8194525C1E6176B6FCA1B6CEE6778">Marino, Pablo I.</mods:namePart>
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<mods:namePart id="565BAAE1B7E18FF01218C8D64279ECEF">Borkent, Art</mods:namePart>
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<mods:title id="BD6B41437E0090A8DA1AC5BAE87996D0">Zootaxa</mods:title>
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<mods:date id="B1303D5CC510FE309B5FE7CE37CFC3A4">2012</mods:date>
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<mods:number id="6D60E53E7F0B33E571E31396EB6A967C">3419</mods:number>
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<mods:identifier id="C5D0E4BAA58CD29B6F88024321719ABB" type="DOI">10.5281/zenodo.215031</mods:identifier>
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<treatment id="03A7878CE37CFFCA53D2BF84869BFCC6" ID-DOI="http://doi.org/10.5281/zenodo.6166110" ID-GBIF-Taxon="119522750" ID-Zenodo-Dep="6166110" LSID="urn:lsid:plazi:treatment:03A7878CE37CFFCA53D2BF84869BFCC6" httpUri="http://treatment.plazi.org/id/03A7878CE37CFFCA53D2BF84869BFCC6" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<subSubSection id="C3146511E37CFFCC53D2BF848322FD1D" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BB1369AE37CFFCC53D2BF8480FBFD3B" blockId="10.[151,703,688,748]" box="[151,703,688,715]" pageId="10" pageNumber="11">
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<heading id="D0F981F6E37CFFCC53D2BF8480FBFD3B" bold="true" box="[151,703,688,715]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B97AEA88E37CFFCC53D2BF8480FBFD3B" bold="true" box="[151,703,688,715]" pageId="10" pageNumber="11">
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<taxonomicName id="4C0E4D19E37CFFCC53D2BF848031FD3A" ID-CoL="8C6JB" box="[151,629,688,715]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="anniae" status="sp. nov." subGenus="Metaforcipomyia">
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<emphasis id="B97AEA88E37CFFCC53D2BF848376FD3A" bold="true" box="[151,306,689,715]" italics="true" pageId="10" pageNumber="11">Forcipomyia</emphasis>
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(
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<emphasis id="B97AEA88E37CFFCC5206BF848055FD3A" bold="true" box="[323,529,689,715]" italics="true" pageId="10" pageNumber="11">Metaforcipomyia</emphasis>
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)
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<emphasis id="B97AEA88E37CFFCC5167BF848031FD3A" bold="true" box="[546,629,689,715]" italics="true" pageId="10" pageNumber="11">anniae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957F3E37CFFCC5139BF8580FBFD3B" box="[636,703,688,714]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB1369AE37CFFCC53D2BFE18322FD1D" blockId="10.[151,703,688,748]" box="[151,358,724,748]" pageId="10" pageNumber="11">
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(
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<figureCitation id="13352A1FE37CFFCC53DABFE18363FD1D" box="[159,295,724,748]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Figs. 24–33</figureCitation>
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,
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<figureCitation id="13352A1FE37CFFCC5277BFE18319FD1D" box="[306,349,724,748]" captionStart="FIGURES 140 – 141" captionStartId="46.[151,264,2010,2033]" captionTargetBox="[284,1314,202,1982]" captionTargetId="figure@46.[276,1315,193,1983]" captionTargetPageId="46" captionText="FIGURES 140 – 141. Distribution map of species of Forcipomyia (Metaforcipomyia)." httpUri="https://zenodo.org/record/215052/files/figure.png" pageId="10" pageNumber="11">140</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146511E37CFFCC53D2BE2E8622FC8D" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="8BB1369AE37CFFCC53D2BE2E8622FC8D" blockId="10.[151,1436,795,2010]" pageId="10" pageNumber="11">
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<emphasis id="B97AEA88E37CFFCC53D2BE2E8355FCC5" bold="true" box="[151,273,795,820]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
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Male and female: only species of Neotropical
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<taxonomicName id="4C0E4D19E37CFFCC506DBE2886D6FCC5" authority="Metaforcipomyia" authorityName="Metaforcipomyia" box="[808,1170,796,820]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEA88E37CFFCC506DBE2881FCFCC5" box="[808,952,797,820]" italics="true" pageId="10" pageNumber="11">Forcipomyia</emphasis>
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(
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<emphasis id="B97AEA88E37CFFCC508CBE2986CDFCC5" box="[969,1161,796,820]" italics="true" pageId="10" pageNumber="11">Metaforcipomyia</emphasis>
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)
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</taxonomicName>
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with a hind tarsal ratio of 1.00–1.09, labrum tapering apically and legs with fore and midfemora with basal, narrow subapical dark bands, hind femur with broad basal dark band, and tibiae with subbasal and apical dark bands.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146511E37CFFCD5382BEB283EEFF06" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
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<paragraph id="8BB1369AE37CFFCC5382BEB28344FBA5" blockId="10.[151,1436,795,2010]" pageId="10" pageNumber="11">
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<emphasis id="B97AEA88E37CFFCC5382BEB2831DFC51" bold="true" box="[199,345,903,928]" pageId="10" pageNumber="11">Description.</emphasis>
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Male. Head. Brown. Eyes abutting medially for length of three ommatidia. Antenna (
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<figureCitation id="13352A1FE37CFFCC567ABEB287D7FC6E" box="[1343,1427,903,927]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 24</figureCitation>
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) pale brown with plume setae poorly developed, flagellomeres 2–4 spherical, flagellomere 10 1.42–1.59 (1.50, n = 4) X longer than flagellomere 11, flagellomeres 10–13 elongate; flagellomere 13 with apical nipple slightly constricted basally; AR 0.86–0.91 (0.88, n = 4). Palpus (
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<figureCitation id="13352A1FE37CFFCC51ADBEC1817DFBFD" box="[744,825,1012,1036]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 25</figureCitation>
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) pale brown; segment 3 slightly swollen at midlength, with shallow, small sensory pit opening on swollen portion; PR (3.00–3.14 (3.10, n = 4). Labrum tapering apically (
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<figureCitation id="13352A1FE37CFFCC53DAB90982B0FBA5" box="[159,244,1084,1108]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 26</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB1369AE37CFFCC538CB96A80BBFA6B" blockId="10.[151,1436,795,2010]" pageId="10" pageNumber="11">
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Thorax. Pale except scutellum, postscutellum brown; scutellum with 6 strong setae. Legs (
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<figureCitation id="13352A1FE37CFFCC5788B96A8764FB86" box="[1229,1312,1119,1143]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 27</figureCitation>
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) pale; fore and midfemora with basal, narrow subapical dark bands, hind femur with broad basal dark band, tibiae with subbasal, apical dark bands; apex of hind tibia with 6 spines, two farthest from spur longer; tarsomeres pale with pectinate scales; tarsomere 1 of foreleg with row of five spine-like bristles, tarsomere 1 of hind leg with patch of thick setae on basal portion; foreleg
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<collectingCountry id="F319760AE37CFFCC5143B9C4806EFAF9" box="[518,554,1265,1288]" name="Chile" pageId="10" pageNumber="11">TR</collectingCountry>
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1.56–1.63 (1.60, n = 4), midleg
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<collectingCountry id="F319760AE37CFFCC50F3B9C4819EFAF9" box="[950,986,1265,1288]" name="Chile" pageId="10" pageNumber="11">TR</collectingCountry>
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0.69–0.77 (0.72, n = 4), hind leg
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<collectingCountry id="F319760AE37CFFCC563DB9C487D8FAF9" box="[1400,1436,1265,1288]" name="Chile" pageId="10" pageNumber="11">TR</collectingCountry>
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1.00–1.09 (1.02, n = 4); claws curved, moderately stout. Wing (
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<figureCitation id="13352A1FE37CFFCC5021B82181F1FADD" box="[868,949,1300,1324]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 28</figureCitation>
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) plain, without pattern of pigmented membrane; M2 only visible at wing margin; first radial cell obliterated; second radial cell well developed; fork of cubitus situated slightly distad to level of apex of costa; wing length 0.62–0.70 (0.67, n = 4) mm; breadth 0.20–0.24 (0.22, n = 4) mm; CR 0.33–0.35 (0.34, n = 4). Halter whitish.
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</paragraph>
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<paragraph id="8BB1369AE37CFFCC538CB892868AF983" blockId="10.[151,1436,795,2010]" pageId="10" pageNumber="11">
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Abdomen. Tergites pale brown with brownish lateral spots. Genitalia (
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<figureCitation id="13352A1FE37CFFCC509DB8928668FA4E" box="[984,1068,1447,1471]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 29</figureCitation>
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): Brown. Tergite 9 short, extending to 1/3 length of gonocoxite; posterior margin truncate; cercus lobe-like, slightly produced beyond 1/2 length of gonocoxite; sternite 9 broad, posterior margin slightly concave. Gonocoxite slender, 2.2 X longer than greatest breadth; gonostylus 0.75 length of gonocoxite, nearly straight, tip pointed. Parameres absent, gonocoxal apodemes slightly curved. Aedeagus (
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<figureCitation id="13352A1FE37CFFCC528FBB028058F9BE" box="[458,540,1591,1615]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 30</figureCitation>
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) stout, lateral margins sclerotized, abruptly narrowed elongate, slender, posteriorly directed prong; basal arch extending to 1/10 of total length; lateral arms extending laterally.
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</paragraph>
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<paragraph id="8BB1369AE37CFFCC5382BB4A8288F846" blockId="10.[151,1436,795,2010]" pageId="10" pageNumber="11">
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Female. As for male, with following differences; antenna pale brown with most flagellomeres missing in available specimens. Palpus (
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<figureCitation id="13352A1FE37CFFCC52F7BB978042F94B" box="[434,518,1698,1722]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 31</figureCitation>
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) brown; segment 3 slightly swollen proximally with small, shallow sensory pit opening on swollen portion; segments 4, 5 completely fused, shorter than segment 3; PR 2.63–3.00 (2.81, n = 4). Thorax. Legs pale brown, fore and midfemora with broad basal, narrow subapical dark bands, hind femur with broad basal dark band, tibiae with subbasal, subapical dark bands; foreleg
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<collectingCountry id="F319760AE37CFFCC50A7BA258642F8D6" box="[994,1030,1808,1831]" name="Chile" pageId="10" pageNumber="11">TR</collectingCountry>
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1.71–1.86 (1.75, n = 4), midleg
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<collectingCountry id="F319760AE37CFFCC563DBA2587D8F8D6" box="[1400,1436,1808,1831]" name="Chile" pageId="10" pageNumber="11">TR</collectingCountry>
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0.75–0.83 (0.81, n = 4), hind leg
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<collectingCountry id="F319760AE37CFFCC514DBA068068F8BB" box="[520,556,1843,1866]" name="Chile" pageId="10" pageNumber="11">TR</collectingCountry>
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1.08–1.09 (1.09, n = 4), claws curved. Wing (
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<figureCitation id="13352A1FE37CFFCC576BBA07863BF8BB" box="[1070,1151,1842,1866]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="10" pageNumber="11">Fig. 32</figureCitation>
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) plain, without pattern of pigmented membrane; first radial cell reduced; second radial cell narrow; fork of cubitus situated distad to level of apex of costa; wing length 0.60–0.66 (0.63, n = 4) mm; breadth 0.22–0.28 (0.25, n = 4) mm; CR 0.33–0.36 (0.35, n = 4).
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</paragraph>
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<paragraph id="8BB1369AE37CFFCD5382BAF783EEFF06" blockId="10.[151,1436,795,2010]" lastBlockId="11.[151,1436,151,247]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
Abdomen. Tergites 1–7 yellowish brown with brownish lateral spots, 8–9 entirely brown. Genital sclerotization uncertain, difficult to see. Two spermathecae (
|
||
<figureCitation id="13352A1FE37DFFCD51ADBDA28179FF5E" box="[744,829,151,175]" captionStart="FIGURES 24 – 33" captionStartId="11.[151,264,1975,1998]" captionTargetBox="[202,1399,304,1940]" captionTargetId="figure@11.[170,1414,297,1957]" captionTargetPageId="11" captionText="FIGURES 24 – 33. Forcipomyia (Metaforcipomyia) anniae sp. n., 24 – 30, male, 31 – 33, female. 24 — flagellum; 25, 31 — palpus; 26 - labrum; 27 — legs (lateral view); 28, 32 — wing; 29 — genitalia (ventral view); 30 — aedeagus; 33 — spermathecae." httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="11" pageNumber="12">Fig. 33</figureCitation>
|
||
) subspherical, lightly sclerotized, with slender, long necks, slightly unequal in size, measuring 0.036 by
|
||
<quantity id="4CF69B7FE37DFFCD5190BD89810DFF25" box="[725,841,188,212]" metricMagnitude="-5" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="12" unit="mm" value="0.034">0.034 mm</quantity>
|
||
, neck
|
||
<quantity id="4CF69B7FE37DFFCD50CABD898646FF25" box="[911,1026,188,212]" metricMagnitude="-6" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="12" unit="mm" value="0.008">0.008 mm</quantity>
|
||
, and 0.032 by
|
||
<quantity id="4CF69B7FE37DFFCD57E1BD898753FF25" box="[1188,1303,188,212]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="mm" value="0.03">0.030 mm</quantity>
|
||
, neck
|
||
<quantity id="4CF69B7FE37DFFCD5618BD898282FF06" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="mm" value="0.006">0.006 mm</quantity>
|
||
. Cercus pale brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF716612E37DFFCD53D2BA82876EF81D" httpUri="https://zenodo.org/record/215035/files/figure.png" pageId="11" pageNumber="12" targetBox="[202,1399,304,1940]" targetPageId="11">
|
||
<paragraph id="8BB1369AE37DFFCD53D2BA82876EF81D" blockId="11.[151,1436,1975,2028]" pageId="11" pageNumber="12">
|
||
<emphasis id="B97AEA88E37DFFCD53D2BA828317F83C" bold="true" box="[151,339,1975,1998]" pageId="11" pageNumber="12">FIGURES 24–33.</emphasis>
|
||
<taxonomicName id="4C0E4D19E37DFFCD521CBA8C80ADF83F" box="[345,745,1976,1998]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="anniae" status="sp. nov." subGenus="Metaforcipomyia">
|
||
<emphasis id="B97AEA88E37DFFCD521CBA8C839FF83F" box="[345,475,1977,1998]" italics="true" pageId="11" pageNumber="12">Forcipomyia</emphasis>
|
||
(
|
||
<emphasis id="B97AEA88E37DFFCD52ACBA8D80D2F83F" box="[489,662,1976,1998]" italics="true" pageId="11" pageNumber="12">Metaforcipomyia</emphasis>
|
||
)
|
||
<emphasis id="B97AEA88E37DFFCD51E1BA8C80ADF83F" box="[676,745,1977,1998]" italics="true" pageId="11" pageNumber="12">anniae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97AEA88E37DFFCD51AABA8D816CF83C" bold="true" box="[751,808,1976,1997]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A24957F3E37DFFCD51AABA8D816CF83C" box="[751,808,1976,1997]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, 24–30, male, 31–33, female. 24—flagellum; 25, 31—palpus; 26-labrum; 27—legs (lateral view); 28, 32—wing; 29—genitalia (ventral view); 30—aedeagus; 33—spermathecae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3146511E37AFFCA5382BDA28281FDE6" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="8BB1369AE37AFFCA5382BDA286F2FF5E" blockId="12.[151,1436,151,824]" box="[199,1206,151,176]" pageId="12" pageNumber="13">
|
||
<emphasis id="B97AEA88E37AFFCA5382BDA28324FF41" bold="true" box="[199,352,151,176]" pageId="12" pageNumber="13">Distribution.</emphasis>
|
||
This species is known only from
|
||
<collectingCountry id="F319760AE37AFFCA519EBDA28111FF5E" box="[731,853,151,175]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
|
||
at 560–1150 meters (
|
||
<figureCitation id="13352A1FE37AFFCA5702BDA286EDFF5E" box="[1095,1193,151,175]" captionStart="FIGURES 140 – 141" captionStartId="46.[151,264,2010,2033]" captionTargetBox="[284,1314,202,1982]" captionTargetId="figure@46.[276,1315,193,1983]" captionTargetPageId="46" captionText="FIGURES 140 – 141. Distribution map of species of Forcipomyia (Metaforcipomyia)." httpUri="https://zenodo.org/record/215052/files/figure.png" pageId="12" pageNumber="13">Fig. 140</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB1369AE37AFFCA5382BD8E828EFE95" blockId="12.[151,1436,151,824]" pageId="12" pageNumber="13">
|
||
<emphasis id="B97AEA88E37AFFCA5382BD8E8396FF25" bold="true" box="[199,466,187,212]" pageId="12" pageNumber="13">Taxonomic discussion.</emphasis>
|
||
Males and females were collected at the same locality and date. This species is similar to
|
||
<taxonomicName id="4C0E4D19E37AFFCA53F1BDD4835CFF09" box="[180,280,224,248]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="rivalis">
|
||
<emphasis id="B97AEA88E37AFFCA53F1BDD48287FF09" box="[180,195,225,248]" italics="true" pageId="12" pageNumber="13">F</emphasis>
|
||
.
|
||
<emphasis id="B97AEA88E37AFFCA5394BDD5835CFF09" box="[209,280,224,248]" italics="true" pageId="12" pageNumber="13">rivalis</emphasis>
|
||
</taxonomicName>
|
||
by virtue of its thorax pigmentation, but
|
||
<taxonomicName id="4C0E4D19E37AFFCA51A2BDD48108FF09" box="[743,844,224,248]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="rivalis">
|
||
<emphasis id="B97AEA88E37AFFCA51A2BDD480B2FF09" box="[743,758,225,248]" italics="true" pageId="12" pageNumber="13">F</emphasis>
|
||
.
|
||
<emphasis id="B97AEA88E37AFFCA5041BDD58108FF09" box="[772,844,224,248]" italics="true" pageId="12" pageNumber="13">rivalis</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished by the labrum bearing a row of transverse, elongate spicules and a row of long setae on the posterior margin of male sternite 9. Characters to distinguish this species from
|
||
<taxonomicName id="4C0E4D19E37AFFCA5299BC1C803EFEB1" box="[476,634,296,320]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="atenasensis">
|
||
<emphasis id="B97AEA88E37AFFCA5299BC1C83AFFEB1" box="[476,491,297,320]" italics="true" pageId="12" pageNumber="13">F</emphasis>
|
||
.
|
||
<emphasis id="B97AEA88E37AFFCA52BCBC1C803EFEB1" box="[505,634,297,320]" italics="true" pageId="12" pageNumber="13">atenasensis</emphasis>
|
||
</taxonomicName>
|
||
are provided in the key and in the taxonomic discussion under that species.
|
||
</paragraph>
|
||
<paragraph id="8BB1369AE37AFFCA5382BC448281FDE6" blockId="12.[151,1436,151,824]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C0E4D19E37AFFCA5382BC4483EEFE79" box="[199,426,369,392]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anniae">
|
||
<emphasis id="B97AEA88E37AFFCA5382BC4483EEFE79" box="[199,426,369,392]" italics="true" pageId="12" pageNumber="13">Forcipomyia anniae</emphasis>
|
||
</taxonomicName>
|
||
is also very similar to
|
||
<taxonomicName id="4C0E4D19E37AFFCA51E8BC448168FE79" box="[685,812,368,392]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pluvialis">
|
||
<emphasis id="B97AEA88E37AFFCA51E8BC4480F8FE79" box="[685,700,369,392]" italics="true" pageId="12" pageNumber="13">F</emphasis>
|
||
.
|
||
<emphasis id="B97AEA88E37AFFCA518EBC458168FE79" box="[715,812,368,392]" italics="true" pageId="12" pageNumber="13">pluvialis</emphasis>
|
||
</taxonomicName>
|
||
. The male of the latter species differs by the pale basal third of the hind femur, by its variable thoracic pigmentation (pale, with patch of darker pigmentation anteriorly, more extensively partially pigmented or entirely darkly pigmented) contrasting with the unicolours pale scutum of
|
||
<taxonomicName id="4C0E4D19E37AFFCA53D2BCE88344FE05" box="[151,256,477,500]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anniae">
|
||
<emphasis id="B97AEA88E37AFFCA53D2BCE88344FE05" box="[151,256,477,500]" italics="true" pageId="12" pageNumber="13">F. anniae</emphasis>
|
||
</taxonomicName>
|
||
, and by the pale gonostylus. Characters to distinguish the females of both species may be found in the key.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146511E37AFFCA5382BF11869BFCC6" pageId="12" pageNumber="13" type="materials_examined">
|
||
<paragraph id="8BB1369AE37AFFCA5382BF118023FD1E" blockId="12.[151,1436,151,824]" pageId="12" pageNumber="13">
|
||
<emphasis id="B97AEA88E37AFFCA5382BF118331FDCD" bold="true" box="[199,373,547,572]" pageId="12" pageNumber="13">
|
||
<typeStatus id="54B58838E37AFFCA5382BF118344FDCD" box="[199,256,548,572]" pageId="12" pageNumber="13">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54B58838E37AFFCA52C4BF1183ADFDCD" box="[385,489,548,572]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
|
||
male on microscope slide, labeled “
|
||
<typeStatus id="54B58838E37AFFCA50CBBF118660FDCD" box="[910,1060,548,572]" pageId="12" pageNumber="13" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="4C0E4D19E37AFFCA576BBF118001FDAE" authority="Spinelli, Marino and Borkent" authorityName="Spinelli, Marino and Borkent" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anniae" subGenus="Metaforcipomyia">Forcipomyia (Metaforcipomyia) anniae Spinelli, Marino and Borkent</taxonomicName>
|
||
,
|
||
<collectingCountry id="F319760AE37AFFCA5110BF728092FDAE" box="[597,726,583,607]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
|
||
, Guanacaste prov.,
|
||
<collectingCountry id="F319760AE37AFFCA50FABF728657FDAE" box="[959,1043,583,607]" name="Liberia" pageId="12" pageNumber="13">Liberia</collectingCountry>
|
||
, Guanacaste NP, Estación Cacao,
|
||
<quantity id="4CF69B7FE37AFFCA53D2BF598376FD75" box="[151,306,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="12" pageNumber="13" unit="m" value="1075.0" valueMax="1150.0" valueMin="1000.0">1000–1150 m</quantity>
|
||
,
|
||
<date id="FFB0105AE37AFFCA5278BF5983ECFD75" box="[317,424,620,644]" pageId="12" pageNumber="13" value="1996-07">VII-1996</date>
|
||
, A. Masis, Malaise tp. LN 323150_375500 #47555, CD 5235” (
|
||
<collectionCode id="ED1FAE5FE37AFFCA5734BF5986FEFD75" box="[1137,1210,620,644]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="12" pageNumber="13">INBC</collectionCode>
|
||
).
|
||
<typeStatus id="54B58838E37AFFCA5789BF588778FD74" box="[1228,1340,621,645]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
|
||
,
|
||
<specimenCount id="9D08FD13E37AFFCA5603BF5987D8FD75" box="[1350,1436,620,644]" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D08FD13E37AFFCA538DBFBA8371FD56" box="[200,309,655,679]" pageId="12" pageNumber="13" type="female">4 females</specimenCount>
|
||
as follows: same data as
|
||
<typeStatus id="54B58838E37AFFCA511EBFBA8084FD56" box="[603,704,655,679]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
,
|
||
<specimenCount id="9D08FD13E37AFFCA5188BFBA8163FD56" box="[717,807,655,679]" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D08FD13E37AFFCA5071BFBA81E6FD56" box="[820,930,655,679]" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
|
||
(
|
||
<specimenCount id="9D08FD13E37AFFCA50F0BFBA8647FD56" box="[949,1027,655,679]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D08FD13E37AFFCA5755BFBA8632FD56" box="[1040,1142,655,679]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED1FAE5FE37AFFCA57C6BFBA868DFD56" box="[1155,1225,655,679]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="12" pageNumber="13">INBC</collectionCode>
|
||
;
|
||
<specimenCount id="9D08FD13E37AFFCA5793BFBA8761FD56" box="[1238,1317,655,679]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D08FD13E37AFFCA5676BFBA87DCFD56" box="[1331,1432,655,679]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED1FAE5FE37AFFCA53D2BF81829BFD3D" box="[151,223,692,716]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="12" pageNumber="13">CNCI</collectionCode>
|
||
).
|
||
<collectingCountry id="F319760AE37AFFCA53B4BF81832AFD3D" box="[241,366,692,716]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
|
||
, Limón prov., Est. Hitoy Cerere,
|
||
<quantity id="4CF69B7FE37AFFCA51A0BF818169FD3D" box="[741,813,692,716]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="13" unit="m" value="560.0">560 m</quantity>
|
||
,
|
||
<date id="FFB0105AE37AFFCA507FBF81818EFD3D" box="[826,970,692,716]" pageId="12" pageNumber="13" value="1998-07-12">12-VII-1998</date>
|
||
, E. Rojas, CD 5077,
|
||
<specimenCount id="9D08FD13E37AFFCA57F2BF818742FD3D" box="[1207,1286,692,716]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D08FD13E37AFFCA5654BF818739FD3D" box="[1297,1405,692,716]" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
|
||
(
|
||
<specimenCount id="9D08FD13E37AFFCA56D5BF818294FD1E" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D08FD13E37AFFCA539EBFE2837BFD1E" box="[219,319,727,751]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED1FAE5FE37AFFCA520FBFE283D5FD1E" box="[330,401,727,751]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="12" pageNumber="13">INBC</collectionCode>
|
||
;
|
||
<specimenCount id="9D08FD13E37AFFCA52D9BFE28044FD1E" box="[412,512,727,751]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED1FAE5FE37AFFCA514EBFE28018FD1E" box="[523,604,727,751]" pageId="12" pageNumber="13">MLPA</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB1369AE37AFFCA5382BFCE869BFCC6" blockId="12.[151,1436,151,824]" pageId="12" pageNumber="13">
|
||
<emphasis id="B97AEA88E37AFFCA5382BFCE8068FCE5" bold="true" box="[199,556,763,788]" pageId="12" pageNumber="13">Derivation of specific epithet.</emphasis>
|
||
This species is named to honour Annia Picado, in recognition of her superb work as a parataxonomist while at the Instituto Nacional de Biodiversidad de
|
||
<collectingCountry id="F319760AE37AFFCA50BBBE2A8633FCC6" box="[1022,1143,799,823]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
|
||
(
|
||
<collectionCode id="ED1FAE5FE37AFFCA57C2BE2A8697FCC6" box="[1159,1235,799,823]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="12" pageNumber="13">INBio</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |