180 lines
18 KiB
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180 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.929.49987" ID-GBIF-Dataset="97c79ed9-33fd-49b8-90c5-95a159839db2" ID-PMC="PMC7192954" ID-Pensoft-Pub="1313-2970-929-1" ID-Pensoft-UUID="EFB92B457B1D56A1996483C5EECFAAF0" ID-PubMed="32377146" ID-ZooBank="4FCA3A27–973C49F7–8264–6A34D279EC5C" ModsDocID="1313-2970-929-1" checkinTime="1587627147117" checkinUser="pensoft" docAuthor="Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki & Bjoern Stelbrink," docDate="2020" docId="E2D86FBA658857FC93211203D47458A8" docLanguage="en" docName="ZooKeys 929: 1-17" docOrigin="ZooKeys 929" docSource="http://dx.doi.org/10.3897/zookeys.929.49987" docTitle="Erhaia nainitalensis Davis & Rao 1997" docType="treatment" docVersion="4" id="EFB92B457B1D56A1996483C5EECFAAF0" lastPageNumber="11" masterDocId="EFB92B457B1D56A1996483C5EECFAAF0" masterDocTitle="The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya" masterLastPageNumber="17" masterPageNumber="1" pageNumber="8" updateTime="1668168753491" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gittenberger, Edmund</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier type="email">egittenberger@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leda, Pema</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wangchuk, Jigme</mods:namePart>
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<mods:affiliation>Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3503-4369</mods:nameIdentifier>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gyeltshen, Choki</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6790-6278</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bjoern Stelbrink,</mods:namePart>
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<mods:affiliation>Department of Animal Ecology & Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany & Zoological Institute, Department of Environmental Sciences, University of Basel, Vesalgasse 1, 4051 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7471-4992</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>929</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>17</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.929.49987</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.929.49987</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-929-1</mods:identifier>
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<treatment ID-GBIF-Taxon="163735475" LSID="urn:lsid:plazi:treatment:E2D86FBA658857FC93211203D47458A8" httpUri="http://treatment.plazi.org/id/E2D86FBA658857FC93211203D47458A8" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
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<subSubSection pageId="7" pageNumber="8" type="nomenclature">
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<paragraph pageId="7" pageNumber="8">
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<taxonomicName LSID="E2D86FBA-6588-57FC-9321-1203D47458A8" authority="Davis & Rao, 1997" authorityName="Davis & Rao" authorityYear="1997" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia nainitalensis" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="nainitalensis">Erhaia nainitalensis Davis & Rao, 1997</taxonomicName>
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="7" pageNumber="8">Fig. 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="reference_group">
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName authorityName="Davis & Rao" authorityYear="1997" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia nainitalensis" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="nainitalensis">
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<pageBreakToken pageId="8" pageNumber="9" start="start">Erhaia</pageBreakToken>
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nainitalensis
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</taxonomicName>
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Davis & Rao, 1997: 276 ("India, Uttar Pradesh, Nainital District, Padampuiri"; "
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<geoCoordinate degrees="29" direction="north" minutes="23" orientation="latitude" precision="925" value="29.383333">29°23'N</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="east" minutes="30" orientation="longitude" precision="925" value="79.5">79°30'E</geoCoordinate>
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")
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName authorityName="Nesemann & S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia chandeshwariensis" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="chandeshwariensis">Erhaia chandeshwariensis</taxonomicName>
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Nesemann and S. Sharma, in
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<bibRefCitation author="Nesemann, H" journalOrPublisher="Molecular Biology and Evolution" pageId="14" pageNumber="15" refId="B13" refString="Nesemann, H, Sharma, S, Sharma, G, Khanal, SN, Pradhan, B, Shah, DN, Tachamo, RD, 2007. Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57-102." title="Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57 - 102." year="2007">Nesemann et al. 2007</bibRefCitation>
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: 64, 78 fig. 4 ("Nepal, Central Zone, Kavre District, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa"; the altitude is not indicated).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="materials_examined">
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<paragraph pageId="8" pageNumber="9">Material examined</paragraph>
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<paragraph pageId="8" pageNumber="9">
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(photos only). Holotype and 4 paratypes of
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<taxonomicName authorityName="Nesemann & S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia chandeshwariensis" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="chandeshwariensis">
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<emphasis italics="true" pageId="8" pageNumber="9">Erhaia chandeshwariensis</emphasis>
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</taxonomicName>
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(NHMW 103315 and 103316).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="10" type="discussion">
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<paragraph pageId="9" pageNumber="10">
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<pageBreakToken pageId="9" pageNumber="10" start="start">Discussion</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="9" pageNumber="10">
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According to
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<bibRefCitation author="Davis, GM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="14" pageNumber="15" pagination="273 - 299" publicationUrl="http://www.jstor.org/stable/4065055" refId="B5" refString="Davis, GM, Rao, S, 1997. Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China. Proceedings of the Academy of Natural Sciences of Philadelphia 148: 273 - 299, http://www.jstor.org/stable/4065055" title="Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China." url="http://www.jstor.org/stable/4065055" volume="148" year="1997">Davis and Rao (1997</bibRefCitation>
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: 277, legends to figure 2) the holotype of
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<taxonomicName lsidName="E. nainitalensis" pageId="9" pageNumber="10" rank="species" species="nainitalensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. nainitalensis</emphasis>
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</taxonomicName>
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is 2.28 mm high. However, elsewhere in the same article (
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<bibRefCitation author="Davis, GM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="14" pageNumber="15" pagination="273 - 299" publicationUrl="http://www.jstor.org/stable/4065055" refId="B5" refString="Davis, GM, Rao, S, 1997. Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China. Proceedings of the Academy of Natural Sciences of Philadelphia 148: 273 - 299, http://www.jstor.org/stable/4065055" title="Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China." url="http://www.jstor.org/stable/4065055" volume="148" year="1997">Davis and Rao 1997</bibRefCitation>
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: 279, table 1) the shell height of a single male animal is reported as 1.62 mm whereas 1.88-1.95-2.14 mm (minimum-mean-maximum) is indicated for five female snails. The difference in measurements between the single male and the five females might be indicative of sexual dimorphism. Shells of the species are also supposed to be
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<normalizedToken originalValue="“minute”">"minute"</normalizedToken>
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, what is defined as 2.0 mm high or smaller (
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<bibRefCitation author="Davis, GM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="14" pageNumber="15" pagination="273 - 299" publicationUrl="http://www.jstor.org/stable/4065055" refId="B5" refString="Davis, GM, Rao, S, 1997. Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China. Proceedings of the Academy of Natural Sciences of Philadelphia 148: 273 - 299, http://www.jstor.org/stable/4065055" title="Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China." url="http://www.jstor.org/stable/4065055" volume="148" year="1997">Davis and Rao 1997</bibRefCitation>
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: 275, 289, table 5). We suppose that the dimensions of the holotype of
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<taxonomicName lsidName="E. chandeshwariensis" pageId="9" pageNumber="10" rank="species" species="chandeshwariensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. chandeshwariensis</emphasis>
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</taxonomicName>
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, i.e. H 1.94 mm, fall within the range of the measurements of
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<taxonomicName lsidName="E. nainitalensis" pageId="9" pageNumber="10" rank="species" species="nainitalensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. nainitalensis</emphasis>
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</taxonomicName>
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. Three of the four paratypes of
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<taxonomicName lsidName="E. chandeshwariensis" pageId="9" pageNumber="10" rank="species" species="chandeshwariensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. chandeshwariensis</emphasis>
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</taxonomicName>
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(NHMW 103316/4) are c. 1.9 mm high, whereas the fourth shell is damaged, missing the apical whorls.
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<bibRefCitation author="Davis, GM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="14" pageNumber="15" pagination="273 - 299" publicationUrl="http://www.jstor.org/stable/4065055" refId="B5" refString="Davis, GM, Rao, S, 1997. Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China. Proceedings of the Academy of Natural Sciences of Philadelphia 148: 273 - 299, http://www.jstor.org/stable/4065055" title="Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China." url="http://www.jstor.org/stable/4065055" volume="148" year="1997">Davis and Rao (1997</bibRefCitation>
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: 278, fig. 3) figure a smooth columella for
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<taxonomicName lsidName="E. nainitalensis" pageId="9" pageNumber="10" rank="species" species="nainitalensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. nainitalensis</emphasis>
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</taxonomicName>
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, but mention (p. 289) a "discernable glassy thickening of the columella".
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</paragraph>
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<paragraph pageId="9" pageNumber="10">
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Judging the nominal taxa on the basis of photographs of shells and additional data in the literature, we conclude that in general shape and apertural characters, i.e. a narrow umbilical chink, a virtually smooth columella, and a thickened outer and inner lip, the holotype of
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<taxonomicName lsidName="E. chandeshwariensis" pageId="9" pageNumber="10" rank="species" species="chandeshwariensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. chandeshwariensis</emphasis>
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</taxonomicName>
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cannot be distinguished from the shells of
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<taxonomicName lsidName="E. nainitalensis" pageId="9" pageNumber="10" rank="species" species="nainitalensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. nainitalensis</emphasis>
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</taxonomicName>
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that are figured by
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<bibRefCitation author="Davis, GM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="14" pageNumber="15" pagination="273 - 299" publicationUrl="http://www.jstor.org/stable/4065055" refId="B5" refString="Davis, GM, Rao, S, 1997. Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China. Proceedings of the Academy of Natural Sciences of Philadelphia 148: 273 - 299, http://www.jstor.org/stable/4065055" title="Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China." url="http://www.jstor.org/stable/4065055" volume="148" year="1997">Davis and Rao (1997</bibRefCitation>
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: 277, figs 2A-F, 280, figs 4A-D). The fact that
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<bibRefCitation author="Davis, GM" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="14" pageNumber="15" pagination="273 - 299" publicationUrl="http://www.jstor.org/stable/4065055" refId="B5" refString="Davis, GM, Rao, S, 1997. Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China. Proceedings of the Academy of Natural Sciences of Philadelphia 148: 273 - 299, http://www.jstor.org/stable/4065055" title="Discovery of Erhaia (Gastropoda: Pomatiopsidae) in northern India with description of a new genus of Erhaiini from China." url="http://www.jstor.org/stable/4065055" volume="148" year="1997">Davis and Rao (1997</bibRefCitation>
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: 276) indicate
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<normalizedToken originalValue="3¾">33/4</normalizedToken>
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-4 whorls for
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<taxonomicName lsidName="E. nainitalensis" pageId="9" pageNumber="10" rank="species" species="nainitalensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. nainitalensis</emphasis>
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</taxonomicName>
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, whereas Nesemann and Sharma (2007: 64) mention
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<normalizedToken originalValue="3½">31/2</normalizedToken>
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whorls for
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<taxonomicName lsidName="E. chandeshwariensis" pageId="9" pageNumber="10" rank="species" species="chandeshwariensis">
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<emphasis italics="true" pageId="9" pageNumber="10">E. chandeshwariensis</emphasis>
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</taxonomicName>
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, might be due to different counting methods.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="family pomatiopsidae stimpson, 1865">
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<paragraph pageId="10" pageNumber="11">
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<pageBreakToken pageId="10" pageNumber="11" start="start">Family</pageBreakToken>
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<taxonomicName family="Pomatiopsidae" lsidName="" pageId="10" pageNumber="11" rank="family">Pomatiopsidae</taxonomicName>
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Stimpson, 1865
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |