217 lines
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217 lines
22 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.969.56214" ID-GBIF-Dataset="8a3ab2f8-451e-4089-a523-7fe9afbdd897" ID-PMC="PMC7515932" ID-Pensoft-Pub="1313-2970-969-43" ID-Pensoft-UUID="555EE844482D585496554D2AD30BB501" ID-PubMed="33013167" ID-ZooBank="C26EB0BB52214953A0AD0822A9050369" ModsDocID="1313-2970-969-43" checkinTime="1600380584787" checkinUser="pensoft" docAuthor="Cumming, Royce T., Tirant, Stephane Le, Teemsma, Sierra N., Hennemann, Frank H., Willemse, Luc & Buescher, Thies H." docDate="2020" docId="D779887611475C8F98AE1232313315AD" docLanguage="en" docName="ZooKeys 969: 43-84" docOrigin="ZooKeys 969" docSource="http://dx.doi.org/10.3897/zookeys.969.56214" docTitle="Nanophyllium miyashitai Cumming & Tirant & Teemsma & Hennemann & Willemse & Büscher 2020, sp. nov." docType="treatment" docUuid="2C3C5029-9E78-4687-A680-BDC9645DD3CB" docUuidSource="ZooBank" docVersion="4" id="555EE844482D585496554D2AD30BB501" lastPageNumber="43" masterDocId="555EE844482D585496554D2AD30BB501" masterDocTitle="Lost lovers linked at long last: elusive female Nanophyllium mystery solved after a century of being placed in a different genus (Phasmatodea, Phylliidae)" masterLastPageNumber="84" masterPageNumber="43" pageNumber="43" updateTime="1668169498602" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Lost lovers linked at long last: elusive female Nanophyllium mystery solved after a century of being placed in a different genus (Phasmatodea, Phylliidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cumming, Royce T.</mods:namePart>
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<mods:affiliation>Associate Researcher, Montreal Insectarium, 4581 rue Sherbrooke est, Montre ́ al, Que ́ bec, H 1 X 2 B 2, Canada & Ph. D. Student, Richard Gilder Graduate School, American Museum of Natural History, New York, NY 10024, USA & Ph. D. program in Biology, Graduate Center, City University of New York, NY, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-7930-1292</mods:nameIdentifier>
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<mods:nameIdentifier type="email">roycecumming@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tirant, Stephane Le</mods:namePart>
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<mods:affiliation>Collection manager, Montreal Insectarium, 4581 rue Sherbrooke, Montre ́ al, Que ́ bec, H 1 X 2 B 2, Canada</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Teemsma, Sierra N.</mods:namePart>
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<mods:affiliation>Associate Researcher, Montreal Insectarium, 4581 rue Sherbrooke est, Montre ́ al, Que ́ bec, H 1 X 2 B 2, Canada</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hennemann, Frank H.</mods:namePart>
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<mods:affiliation>Tannenwaldallee 53, 61348 Bad Homburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Willemse, Luc</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, PO Box 9517, NL- 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0517-9778</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Buescher, Thies H.</mods:namePart>
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<mods:affiliation>Department of Functional Morphology and Biomechanics, Zoological Institute, Kiel University, Am Botanischen Garten 9, 24118 Kiel, Germany</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0639-4699</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>969</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>43</mods:start>
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<mods:end>84</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.969.56214</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.969.56214</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-969-43</mods:identifier>
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<mods:identifier type="ZooBank">C26EB0BB52214953A0AD0822A9050369</mods:identifier>
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<mods:identifier type="Pensoft-UUID">555EE844482D585496554D2AD30BB501</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="167677368" LSID="urn:lsid:zoobank.org:act:2C3C5029-9E78-4687-A680-BDC9645DD3CB" httpUri="http://treatment.plazi.org/id/D779887611475C8F98AE1232313315AD" lastPageNumber="43" pageId="0" pageNumber="43">
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<subSubSection pageId="0" pageNumber="43" type="nomenclature">
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<paragraph pageId="0" pageNumber="43">
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<taxonomicName LSID="D7798876-1147-5C8F-98AE-1232313315AD" authority="Cumming & Tirant & Teemsma & Hennemann & Willemse & Büscher, 2020" authorityName="Cumming & Tirant & Teemsma & Hennemann & Willemse & Büscher" authorityYear="2020" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanophyllium miyashitai" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="miyashitai" status="sp. nov.">Nanophyllium miyashitai</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">Figure 20</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="type material">
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<paragraph pageId="0" pageNumber="43">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="43">Holotype</emphasis>
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: ♂, Papua New Guinea: Morobe Province, Wau: IX, 2000. From the collection of Tetsuo Miyashita, Japan. Deposited in the Montreal Insectarium (Quebec, Canada) type collection.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="differentiation">
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<paragraph pageId="0" pageNumber="43">Differentiation.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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With the interior lobe of the profemora rounded, not angular, and the mesofemoral interior lobe broad and reaching fully end to end in a rounded triangle, this new species falls within the
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<taxonomicName authorityName="Cumming" authorityYear="2016" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="stellae" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="stellae">
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<emphasis italics="true" pageId="0" pageNumber="43">stellae</emphasis>
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</taxonomicName>
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species group. This is the first species from the
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<taxonomicName authorityName="Cumming" authorityYear="2016" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="stellae" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="stellae">
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<emphasis italics="true" pageId="0" pageNumber="43">stellae</emphasis>
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</taxonomicName>
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species group recorded from Papua New Guinea. The other two species are known from Jayapura, Irian Jaya, Indonesia (very near the border with Papua New Guinea so it is likely they also occur there but to date, we have not confirmed any specimens). This new species can be differentiated from the other two species in the
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<taxonomicName authorityName="Cumming" authorityYear="2016" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="stellae" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="stellae">
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<emphasis italics="true" pageId="0" pageNumber="43">stellae</emphasis>
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</taxonomicName>
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species group by the mesopleurae which have a prominent anterior tubercle followed by four additional small tubercles (only a single anterior tubercle in the other two species) and tegmina that are shorter, only about half the length of the metathorax (almost the length of the metathorax in the other two species).
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</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Like the other members of the
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<taxonomicName authorityName="Cumming" authorityYear="2016" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanophyllium stellae" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="stellae">
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<emphasis italics="true" pageId="0" pageNumber="43">Nanophyllium stellae</emphasis>
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</taxonomicName>
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species group, the holotype is a male specimen and the female is unknown. It is expected that the female is larger than other known female
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<taxonomicName class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanophyllium" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="43">Nanophyllium</emphasis>
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</taxonomicName>
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as the
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<taxonomicName authorityName="Cumming" authorityYear="2016" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="stellae" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="stellae">
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<emphasis italics="true" pageId="0" pageNumber="43">stellae</emphasis>
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</taxonomicName>
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species group members are larger than the
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<taxonomicName authorityName="Redtenbacher" authorityYear="1906" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="pygmaeum" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="pygmaeum">
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<emphasis italics="true" pageId="0" pageNumber="43">pygmaeum</emphasis>
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</taxonomicName>
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species group members.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="description">
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<paragraph pageId="0" pageNumber="43">Description.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">
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Male.
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<emphasis italics="true" pageId="0" pageNumber="43">Coloration</emphasis>
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.
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</emphasis>
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Antennae dark brown, a similar brown to that found throughout the head and thorax. The majority of the dorsal aspect throughout the remainder of the body and legs is of a similar lighter brown, but not a light as the stripe of light brown running along the sagittal plane along the head and thorax. Alae and tegmina have a similar dark brown to that found on the antennae. Throughout the ventral surface the coloration is the same as that found on the legs. Granulation on the body is mostly of a lighter brown than the surface it is found on.
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</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">
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<emphasis italics="true" pageId="0" pageNumber="43">Morphology</emphasis>
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.
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</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="43">Head</emphasis>
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. Head capsule slightly longer than wide, with a vertex that is heavily granulose, which includes the two posteromedian tubercles which are no larger than the surrounding granulation around them (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20F</figureCitation>
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). Three well-developed ocelli are slightly posterior to the compound eyes which are ovular and slightly protrude from the head capsule (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20F</figureCitation>
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).
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<emphasis italics="true" pageId="0" pageNumber="43">Antennae</emphasis>
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. Antennae in the holotype are both damaged and repaired so the original number of antennomeres is unknown. The antennae are longer than the outstretched forelegs and the left antennae consist of 21 antennomeres and the right of 19 (including the scapus and pedicellus). Scapus and pedicellus with short clear setae and the scapus has a notable spur on the anterior rim lateral side. All segments beyond the scapus and pedicellus covered in stiff dark setae each longer than the segment is wide until the terminal four segments where the setae begin to steadily decrease in size until the terminal segment which has dense short setae.
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<emphasis italics="true" pageId="0" pageNumber="43">Thorax</emphasis>
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. Pronotum wider than long (width to length, 1: 0.75) with parallel lateral margins, and all margins slightly granulose. Surface of the pronotum heavily granulose like the vertex of the head capsule. Prescutum significantly wider than long (width to length, 3.3: 1), with converging lateral margins with a granular surface of at least five nodes (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20F</figureCitation>
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). Surface of prescutum slightly granular but lacking significant features. Mesopleurae gently diverging, anterior edge armed with a single tubercle, remainder of the rim with four small tubercles with a single seta protruding from the tip of each. Mesopleurae surface irregularly granular with a single distinct pit in the center. Pro-, meso-, and metasternum covered in irregularly spaced granules.
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<emphasis italics="true" pageId="0" pageNumber="43">Wings</emphasis>
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. Tegmina short, only reaching about halfway through the metathorax. Alae developed; exposed section of folded alae moderately sclerotized.
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<emphasis italics="true" pageId="0" pageNumber="43">Abdomen</emphasis>
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. Abdominal segments with folding in the holotype so shape description is only approximate. Abdominal segment II slightly tapering, III gradually widening, IV widening for the first quarter, then parallel, V through the first half of VI parallel, VII converging, VIII-IX parallel to subparallel. Anal abdominal segment X longer than wide with a broad rounded apex (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20D</figureCitation>
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). Poculum broad, about as broad as segment IX, ending in a broad rounded apex that reaches the anterior margin of segment X (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20E</figureCitation>
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). Cercus about as wide as the vomer but slightly shorter, margins marked with a row of thin tan setae and a dorsal surface that is heavily granular. Vomer long, reaching the majority of the length to the apex, with sides gradually converging to the hooked apex (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20E</figureCitation>
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).
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<emphasis italics="true" pageId="0" pageNumber="43">Legs</emphasis>
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. Profemora, interior lobe rounded with three small nubby, evenly spaced teeth (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20C</figureCitation>
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). Exterior lobe wider than the interior lobe and with a slight recurve and an edge that is smooth with a row of single stout setae along the entire length. Protibiae lacking an exterior lobe, interior lobe a rounded scalene triangle spanning the entire length of the protibia (Fig.
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Holotype male Nanophyllium miyashitai sp. nov. A full body dorsal B full body ventral C right front leg D abdominal segment X and dorsal view of cerci E genitalia, ventral F head through thorax, dorsal." figureDoi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43">20C</figureCitation>
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). Mesofemora, exterior lobe smoothly arcing the length of the mesofemora, interior lobe smoothly triangular with five to six small nubby teeth on the distal half and about one and a half times as wide as exterior lobe. Interior and exterior lobe of metafemora smoothly arcing with interior lobe about twice as wide as the exterior lobe and the interior with a few small nubby teeth near the distal end. Meso- and metatibiae lacking exterior and interior lobes.
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</paragraph>
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<caption doi="10.3897/zookeys.969.56214.figure20" httpUri="https://binary.pensoft.net/fig/453795" pageId="0" pageNumber="43" start="Figure 20" startId="F20">
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Figure 20.</emphasis>
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Holotype male
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<taxonomicName authorityName="Cumming & Tirant & Teemsma & Hennemann & Willemse & Büscher" authorityYear="2020" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanophyllium miyashitai" order="Phasmida" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="miyashitai">
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<emphasis italics="true" pageId="0" pageNumber="43">Nanophyllium miyashitai</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
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full body dorsal
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<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
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full body ventral
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<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
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right front leg
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<emphasis bold="true" pageId="0" pageNumber="43">D</emphasis>
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abdominal segment X and dorsal view of cerci
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<emphasis bold="true" pageId="0" pageNumber="43">E</emphasis>
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genitalia, ventral
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<emphasis bold="true" pageId="0" pageNumber="43">F</emphasis>
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head through thorax, dorsal.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="description">
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<paragraph pageId="0" pageNumber="43">Measurements of holotype</paragraph>
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<paragraph pageId="0" pageNumber="43">
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[
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<emphasis bold="true" pageId="0" pageNumber="43">mm].</emphasis>
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Length of body (including cerci and head, excluding antennae) 40.0, length/width of head 2.6/2.5, antennae (repaired) 16.4, pronotum 1.8, mesonotum 1.8, length of tegmina 5.9, length of alae 30.5, greatest width of abdomen 8.0, profemora 7.4, mesofemora 6.8, metafemora 7.3, protibiae 3.9, mesotibiae 5.0, metatibiae 6.7.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="distribution">
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<paragraph pageId="0" pageNumber="43">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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Currently only known from the type locality of Wau, Morobe Province, Papua New Guinea (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map showing localities for the known and herein described species." figureDoi="10.3897/zookeys.969.56214.figure4" httpUri="https://binary.pensoft.net/fig/453779" pageId="0" pageNumber="43">4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="etymology">
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<paragraph pageId="0" pageNumber="43">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="43">Patronym. This species is dedicated to Mr. Tetsuo Miyashita (Japan). Miyashita is a major private collector who has amassed one of the largest insect collections in the world. Miyashita and the specimens from his collection have allowed the description of several new beetle taxa over the years with this being the first phasmid described from his collection.</paragraph>
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</subSubSection>
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</treatment>
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</document> |