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<mods:url id="DB822EC5CD3E9C4B85D4C6B3A8397EFA">http://dx.doi.org/10.3897/phytokeys.240.101716</mods:url>
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<mods:classification id="317951E836CFF4F19B4C49FC32B2CB1E">journal article</mods:classification>
<mods:identifier id="7179CE0E947EF7A2C19EA4BA801BC653" type="DOI">http://dx.doi.org/10.3897/phytokeys.240.101716</mods:identifier>
<mods:identifier id="5D5E7C79E96139694C8D519EDAB9299C" type="Pensoft-Pub">1314-2003-240-1</mods:identifier>
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<treatment id="C3D268747A3F3439CBB7B434BADBBD0E" LSID="urn:lsid:plazi:treatment:C3D268747A3F3439CBB7B434BADBBD0E" httpUri="http://treatment.plazi.org/id/C3D268747A3F3439CBB7B434BADBBD0E" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="7E7543EC845F2D7188579F6F970DFF60" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="1AE1C5611C83CACBFDABE9469574D492" pageId="0" pageNumber="1">
<taxonomicName id="2E48B4C4DC9C1B54B5C10B38A85DCFBB" ID-CoL="9CSK5" authority="Gagnon &amp; G. P. Lewis, PhytoKeys 71: 58. 2016." authorityName="Gagnon &amp; G. P. Lewis, PhytoKeys 71: 58. 2016." class="Magnoliopsida" family="Fabaceae" genus="Hultholia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hultholia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Hultholia Gagnon &amp; G. P. Lewis, PhytoKeys 71: 58. 2016.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="72E7A0198BD9DFBC4989EA57014344A2" pageId="0" pageNumber="1" type="description">
<paragraph id="6EE2C2CD416DC455DB2C2ADDCAD68DEC" pageId="0" pageNumber="1">
<figureCitation id="F0A49569F6BE21D2FEC55DAF0DAF51B1" captionStart="Figure 35" captionStartId="F42" captionText="Figure 35. Flowers of Caesalpinieae A Tara spinosa Britton &amp; Rose, Peru, Ancash (Hughes et al. 3043) B Cenostigma pluviosum (DC.) Gagnon &amp; G. P. Lewis var. pluviosum, Bolivia, Santa Cruz (Wood et al. 26552) C Caesalpinia bahamensis Lam., Cuba (Lewis 1853) D Hultholia mimosoides (Lam.) Gagnon &amp; G. P. Lewis, India E Pomaria burchellii (DC.) B. B. Simpson &amp; G. P. Lewis, Botswana Ghanzi district F Erythrostemon coccineus (G. P. Lewis &amp; J. L. Contr.) Gagnon &amp; G. P. Lewis, Mexico, Oaxaca (Lewis et al. 1802) G Caesalpinia pulcherrima (L.) Sw., Honduras H Erythrostemon melanadenius (Rose) Gagnon &amp; G. P. Lewis, Mexico, Oaxaca (Hughes et al. 2091) I Pterolobium stellatum (Forssk.) Brenan, Africa J Guilandina bonduc L., India K Gelrebia trothaei (Harms) Gagnon &amp; G. P. Lewis, Tanzania. Photo credits A E Gagnon B, F-H CE Hughes C GP Lewis D VR Vinayaraj, India Biodiversity Portal (https: // indiabiodiversity. org / group / wild _ orchids _ of _ india / observation / show / 335155), the basionym of Hultholia mimosoides E O Bourquin, Flora of Zimbabwe (https: // www. zimbabweflora. co. zw / speciesdata / image-display. php? species _ id = 127200 &amp; image _ id = 6) I P Van Wyk J M Sanjappa K PJ Cribb." figureDoi="10.3897/phytokeys.240.101716.figure35" httpUri="https://binary.pensoft.net/fig/1014708" pageId="0" pageNumber="1">Figs 35</figureCitation>
<figureCitation id="4C351531D3DFBA7BC53FEF858019AD2A" captionStart="Figure 37" captionStartId="F44" captionText="Figure 37. Fruits of the Caesalpinieae A Gelrebia rubra (Engl.) Gagnon &amp; G. P. Lewis, Namibia B Moullava spicata (Dalzell ex Wight) Nicolson, India, Maharashtra C Hultholia mimosoides (Lam.) Gagnon &amp; G. P. Lewis, India D Guilandina bonduc L., Madagascar E Pterolobium stellatum (Forssk.) Brenan, Zimbabwe F Ticanto sinensis (Hemsley) R. Clark &amp; Gagnon, China (Clark 415) G Biancaea decapetala (Roth) O. Deg., Peru, Ancash (Hughes et al. 2227) H Mezoneuron kauaiense (H. Mann) Hillebr. I Mezoneuron andamanicum Prain, Thailand (Clark 251). Photo credits A Dave U, iNaturalist (https: // www. inaturalist. org / photos / 41085094) B P Awale, Flowers of India (http: // www. flowersofindia. net /) C VR Vinayaraj, India Biodiversity Portal (https: // indiabiodiversity. org / observation / show / 335158), the basionym of Hultholia mimosoides D GP Lewis E BT Wursten, Flora of Zimbabwe (https: // www. zimbabweflora. co. zw / speciesdata / image-display. php? species _ id = 127190 &amp; image _ id = 1) F, I P Suksathan G CE Hughes H D Eickhoff, https: // www. flickr. com / photos / dweickhoff / 4822012867 / in / photostream /." figureDoi="10.3897/phytokeys.240.101716.figure37" httpUri="https://binary.pensoft.net/fig/1014710" pageId="0" pageNumber="1">, 37</figureCitation>
<figureCitation id="74D63995349217264CFA787CB1803A48" captionStart="Figure 60" captionStartId="F67" captionText="Figure 60. Distribution of Hultholia based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure60" httpUri="https://binary.pensoft.net/fig/1014733" pageId="0" pageNumber="1">, 60</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9B7A5C3F232AEA5DDF4DC3AFB6616E39" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="1F2AC83F3AB44944B53C59AF43479929" pageId="0" pageNumber="1">
<typeStatus id="1A59A9DAAC149DF7EC5EA46340968254">Type</typeStatus>
.
</paragraph>
<paragraph id="3070EC05B3E7D64DD9AF94D45092A574" pageId="0" pageNumber="1">
<taxonomicName id="F9AB44CD1CFBEF431AAEB0708C577D4B" class="Magnoliopsida" family="Caesalpiniaceae" genus="Hultholia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hultholia mimosoides" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="mimosoides">
<emphasis id="3531497EE7AAC9078D7DBD29CB93EA09" italics="true" pageId="0" pageNumber="1">Hultholia mimosoides</emphasis>
</taxonomicName>
(Lam.) Gagnon &amp; G. P. Lewis [≡ 
<taxonomicName id="E7C23C58DFE077C87DC3B48838587055" class="Magnoliopsida" family="Leguminosae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia mimosoides" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="mimosoides">
<emphasis id="24C56BC38392A2E5B2048A3DF08EF56B" italics="true" pageId="0" pageNumber="1">Caesalpinia mimosoides</emphasis>
</taxonomicName>
Lam.]
</paragraph>
</subSubSection>
<subSubSection id="0E76AC7B552F82E87FC9B90A5579D684" pageId="0" pageNumber="1" type="description">
<paragraph id="26260CA872778F8306890474C6D93AEC" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="9943D1DB8A2C5767FB27BD57B41A1DE7" pageId="0" pageNumber="1">
Climbing woody shrub; branches densely armed with short, robust, needle-like trichomes; young stems pubescent, with rust-coloured, hyaline hairs, and dome-shaped glands, topped with a few hairs. 
<emphasis id="D01F163812F3BEB0C69A7A486CF36312" bold="true" pageId="0" pageNumber="1">Stipules</emphasis>
subulate, caducous. 
<emphasis id="A8FFCB154CB3D22E31995026CA98E397" bold="true" pageId="0" pageNumber="1">Leaves</emphasis>
bipinnate; pinnae in 10-30 opposite pairs, with a pair of deflexed prickles at the insertion of the pinnae on the leaf rachis and at the insertion of leaflets on the pinnae rachides; leaflets 7-20 pairs per pinna, opposite, glabrous, eglandular. 
<emphasis id="094A2F62745E62B5116D7600EA299956" bold="true" pageId="0" pageNumber="1">Inflorescence</emphasis>
a terminal or leaf-opposed, lax raceme, with 50 or more flowers. 
<emphasis id="202F6E53C34D1B0FEC6360613D225669" bold="true" pageId="0" pageNumber="1">Flowers</emphasis>
bisexual, zygomorphic; hypanthium persisting as a wide shallow cup at the pedicel apex as fruit matures; sepals 5, caducous, the lower sepal strongly cucullate; petals 5, free, bright yellow, dark glands present on the blade, median petal smaller than the 4 lateral petals; stamens 10, free, filaments pubescent at least on the lower half; ovary densely pubescent and with glandular dots (often obscured by the dense pubescence). 
<emphasis id="FD012E3B615FE03EF0BE782510A7E1FC" bold="true" pageId="0" pageNumber="1">Fruit</emphasis>
an obovoid, falcate, vesicular, unarmed, dehiscent legume, sparsely pubescent, particularly along the margin, and with a few obscure stellate hairs, and covered in gland dots, 1-3-seeded. 
<emphasis id="769D31BE5E56CFFF68244EF42DC8DB6C" bold="true" pageId="0" pageNumber="1">Seeds</emphasis>
sub-globose, grey.
</paragraph>
</subSubSection>
<subSubSection id="E4928FA61C33FDBF48D87A099E0F1512" pageId="0" pageNumber="1" type="chromosome number">
<paragraph id="D5F170195184227CA54B09E7F0FC303F" pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph id="C4ECAFFFEFE1D03A81C6EC17983F2445" pageId="0" pageNumber="1">
2 
<emphasis id="C1B645BDC80F5DFE0743622011D7029D" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 24 (
<bibRefCitation id="3241C4AFBA88AD9121B81F42A062BA74" author="Fedorov, A [Ed.]" journalOrPublisher="Komorov Botanical Institute, Leningrad" pageId="0" pageNumber="1" refId="B270" refString="Fedorov, A [Ed.], 1969. Chromosome numbers of flowering plants. Academy of Sciences of USSR, V. L. Komorov Botanical Institute, Leningrad" title="Chromosome numbers of flowering plants. Academy of Sciences of USSR, V. L." year="1969">Fedorov 1969</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0FD5E6F9A05E206257588CBD4340D63D" pageId="0" pageNumber="1" type="distribution">
<paragraph id="7E9A979ADDB78D550DA42A9579622E0F" pageId="0" pageNumber="1">Included species and geographic distribution.</paragraph>
<paragraph id="368790DABC52CD5E804501231ADE51EA" pageId="0" pageNumber="1">
Monospecific (
<taxonomicName id="34B47550332105FB978024A3B2E14CE5" lsidName="H. mimosoides" pageId="0" pageNumber="1" rank="species" species="mimosoides">
<emphasis id="F862457B41E2811D650A9DF23EA2FF30" italics="true" pageId="0" pageNumber="1">H. mimosoides</emphasis>
</taxonomicName>
), distributed across Asia, in China (Yunnan), Bangladesh, India, Laos, Myanmar (Burma), Thailand and Vietnam (Fig. 
<figureCitation id="EAB1156753B16BCC4664ECB8F89459F1" captionStart="Figure 60" captionStartId="F67" captionText="Figure 60. Distribution of Hultholia based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure60" httpUri="https://binary.pensoft.net/fig/1014733" pageId="0" pageNumber="1">60</figureCitation>
).
</paragraph>
<caption id="A41EBC92E2CC80334F42D876F3BDFBF5" doi="10.3897/phytokeys.240.101716.figure60" httpUri="https://binary.pensoft.net/fig/1014733" pageId="0" pageNumber="1" start="Figure 60" startId="F67">
<paragraph id="72172EEA970441BBFBFA214B6601AA3C" pageId="0" pageNumber="1">
<emphasis id="1C6C915497FF98F4412B46A99FD35D85" bold="true" pageId="0" pageNumber="1">Figure 60.</emphasis>
Distribution of 
<taxonomicName id="4E6990EBFA683F02670A3A5BBAD19E96" class="Magnoliopsida" family="Caesalpiniaceae" genus="Hultholia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hultholia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="C98569D7524788E9877DDE4F77DB0AE4" italics="true" pageId="0" pageNumber="1">Hultholia</emphasis>
</taxonomicName>
based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="CC1C632E313B0A14C18A99B5DF981EF4" pageId="0" pageNumber="1" type="ecology">
<paragraph id="7DBDC8D1F7D7001C28BB484C5C919554" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="981B12ABE915A6CA7ECD0C6A4D68CE70" pageId="0" pageNumber="1">In secondary thickets and clearings, often on roadsides.</paragraph>
</subSubSection>
<subSubSection id="3C09BC17802C189BCCFC23D9EA9A11E2" pageId="0" pageNumber="1" type="etymology">
<paragraph id="A114E6E3D11A27CC1C59EBAF77B8E666" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="71A620A5F26D0AD326BCB2E25EE46A32" pageId="0" pageNumber="1">
The name 
<taxonomicName id="5DCDBDE91EFEA8A98B50C646A9CCCD2B" class="Magnoliopsida" family="Caesalpiniaceae" genus="Hultholia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hultholia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="31D0CF99AD72E15FD2013F07F29E7D3C" italics="true" pageId="0" pageNumber="1">Hultholia</emphasis>
</taxonomicName>
honours the Cambodian botanist Dr. Sovanmoly Hul Thol (born 1946), whose doctoral thesis, &quot;Contribution 
<normalizedToken id="306F70AA55D234EF5213EB9F57548E34" originalValue="à">a</normalizedToken>
la 
<normalizedToken id="623C546861AB6353AF84CC26CCEE7CF8" originalValue="révision">revision</normalizedToken>
de quelques genres de 
<taxonomicName id="A6E6588B04E79ABBC4DC11A445AC43AF" family="Caesalpiniaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="family">Caesalpiniaceae</taxonomicName>
, 
<normalizedToken id="2F8B01D63A5B7EFF783F74D9C146D30E" originalValue="representés">representes</normalizedToken>
en Asie&quot; (Hul Thol 1976), is an important revision of the Asian species and genera of the 
<taxonomicName id="5285D28AC120ED5F4354109FBB2F6225" family="Caesalpiniaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Caesalpinieae">Caesalpinieae</taxonomicName>
, and particularly the genus 
<taxonomicName id="18CE4789644FFDCA88951E067821B830" class="Magnoliopsida" family="Fabaceae" genus="Pterolobium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pterolobium" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="0969DA482CF1212EE4C305473208B1C8" italics="true" pageId="0" pageNumber="1">Pterolobium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BA03B68F52CB924258C2730064183A74" pageId="0" pageNumber="1" type="human uses">
<paragraph id="7F4794CE4DFCF7A6EAF02664BFF957DB" pageId="0" pageNumber="1">Human uses.</paragraph>
<paragraph id="D3D2C4D615C38A0155B2E468558A5572" pageId="0" pageNumber="1">
Although 
<taxonomicName id="5E33E699D8915C81D9CA2FC9C7F1853B" class="Magnoliopsida" family="Caesalpiniaceae" genus="Hultholia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hultholia mimosoides" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="mimosoides">
<emphasis id="EEA66DF4CBEE9AA9652A1CB688447D61" italics="true" pageId="0" pageNumber="1">Hultholia mimosoides</emphasis>
</taxonomicName>
is not known to be cultivated, the young, pungent, flowering shoots are sold as a vegetable in markets in Vientiane (Laos) (Vidal and Hul Thol 1976).
</paragraph>
</subSubSection>
<subSubSection id="28E9EF691A4051F5B6587C23B58EA42B" pageId="0" pageNumber="1" type="notes">
<paragraph id="8A4A53487D492F03329CDD9920F59B6C" pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph id="AAD668AD2A9C5CE000732A712D8D82BA" pageId="0" pageNumber="1">
This monospecific genus was described by 
<bibRefCitation id="2B62B63588B9198213222361535C8CE6" DOI="https://doi.org/10.3897/phytokeys.71.9203" author="Gagnon, E" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 160" refId="B299" refString="Gagnon, E, Bruneau, A, Hughes, CE, Queiroz, LP, Lewis, GP, 2016. A new generic system for the pantropical Caesalpinia group (Leguminosae). PhytoKeys 71: 1 - 160, DOI: https://doi.org/10.3897/phytokeys.71.9203" title="A new generic system for the pantropical Caesalpinia group (Leguminosae)." url="https://doi.org/10.3897/phytokeys.71.9203" volume="71" year="2016">Gagnon et al. (2016)</bibRefCitation>
and has an unresolved phylogenetic position within a clade of lianescent Old World genera.
</paragraph>
</subSubSection>
<subSubSection id="0FAA8249411DD0376D00985C4D6F44D5" pageId="0" pageNumber="1" type="taxonomic references">
<paragraph id="2C5ED369FA2428417CFBF2DDA3868EE2" pageId="0" pageNumber="1">Taxonomic references.</paragraph>
<paragraph id="F4D7AF8272C1A13A25C734D776FA8746" pageId="0" pageNumber="1">
<bibRefCitation id="B04734FA63097DCBCB49CEF5EB625321" author="Chen, D" editor="Wu, CY" journalOrPublisher="10. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis, USA" pageId="0" pageNumber="1" pagination="41 - 47" refId="B178" refString="Chen, D, Zhang, D, Hou, D, 2010a. 19. Caesalpinia. In: Wu, CY, Raven, P, Eds., Flora of China, vol. 10. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis, USA: 41 - 47" title="19. Caesalpinia." volumeTitle="Flora of China, vol." year="2010 a">Chen et al. (2010a)</bibRefCitation>
; 
<bibRefCitation id="71A31A5229CF994F09F787403CE259DF" DOI="https://doi.org/10.3897/phytokeys.71.9203" author="Gagnon, E" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 160" refId="B299" refString="Gagnon, E, Bruneau, A, Hughes, CE, Queiroz, LP, Lewis, GP, 2016. A new generic system for the pantropical Caesalpinia group (Leguminosae). PhytoKeys 71: 1 - 160, DOI: https://doi.org/10.3897/phytokeys.71.9203" title="A new generic system for the pantropical Caesalpinia group (Leguminosae)." url="https://doi.org/10.3897/phytokeys.71.9203" volume="71" year="2016">Gagnon et al. (2016)</bibRefCitation>
; Vidal and Hul Thol (1976).
</paragraph>
</subSubSection>
</treatment>
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