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<mods:title id="E376C6B55061A51EFC0AFD0D69E612A5">Taxonomic review of the genus Sicciaemorpha van Eecke (Lepidoptera Erebidae: Arctiinae), with descriptions of six new species from Malaysia and the Philippines</mods:title>
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<paragraph id="357C3693FF9A9D15FF74FF602F74ADA0" blockId="1.[151,628,223,250]" box="[151,628,223,250]" pageId="1" pageNumber="70">
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Genus
<taxonomicName id="F2C34D10FF9A9D15FF0DFF602F74ADA0" authority="van Eecke, 1920" authorityName="van Eecke" authorityYear="1920" box="[238,628,223,250]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF0DFF602CA3ADA3" bold="true" box="[238,419,223,250]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
<bibRefCitation id="51524B62FF9A9D15FE48FF602F74ADA0" author="van Eecke, R." box="[427,628,223,249]" pageId="1" pageNumber="70" pagination="112 - 138" refId="ref6757" refString="van Eecke, R. (1920) Studien over indo-australische Lepidoptera. IV. Bijdrage tot de kennis der Heterocera-fauna der oost-in- dische Kolonien. Zoologische mededeelingen, 5 (13), 112 - 138. [in Dutch]" type="journal article" year="1920">van Eecke, 1920</bibRefCitation>
</taxonomicName>
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<taxonomicName id="F2C34D10FF9A9D15FF74FE9B2CE0AC65" authority="van Eecke, 1920" authorityName="van Eecke" authorityYear="1920" box="[151,480,292,316]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF74FE9B2C2FAC65" box="[151,303,292,316]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
<bibRefCitation id="51524B62FF9A9D15FED6FE9B2CE0AC65" author="van Eecke, R." box="[309,480,292,316]" pageId="1" pageNumber="70" pagination="112 - 138" refId="ref6757" refString="van Eecke, R. (1920) Studien over indo-australische Lepidoptera. IV. Bijdrage tot de kennis der Heterocera-fauna der oost-in- dische Kolonien. Zoologische mededeelingen, 5 (13), 112 - 138. [in Dutch]" type="journal article" year="1920">van Eecke, 1920</bibRefCitation>
</taxonomicName>
,
<emphasis id="07B7EA81FF9A9D15FE09FE9B2E04AC65" box="[490,772,292,316]" italics="true" pageId="1" pageNumber="70">Zoologische mededeelingen</emphasis>
, 5, 129.
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<paragraph id="357C3693FF9A9D15FF74FEFC2E56AC02" blockId="1.[151,855,291,347]" box="[151,854,322,347]" pageId="1" pageNumber="70">
<typeStatus id="EA788831FF9A9D15FF74FEFC2DC9AC02" box="[151,201,323,347]" pageId="1" pageNumber="70">Type</typeStatus>
species:
<taxonomicName id="F2C34D10FF9A9D15FEC5FEFD2FC0AC03" authority="van Eecke, 1920" authorityName="van Eecke" authorityYear="1920" box="[294,704,322,346]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="species" species="ivyalba">
<emphasis id="07B7EA81FF9A9D15FEC5FEFD2F10AC03" box="[294,528,322,346]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha ivyalba</emphasis>
<bibRefCitation id="51524B62FF9A9D15FDF5FEFC2FC0AC03" author="van Eecke, R." box="[534,704,322,346]" pageId="1" pageNumber="70" pagination="112 - 138" refId="ref6757" refString="van Eecke, R. (1920) Studien over indo-australische Lepidoptera. IV. Bijdrage tot de kennis der Heterocera-fauna der oost-in- dische Kolonien. Zoologische mededeelingen, 5 (13), 112 - 138. [in Dutch]" type="journal article" year="1920">van Eecke, 1920</bibRefCitation>
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, by monotypy.
</paragraph>
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<paragraph id="357C3693FF9A9D15FF74FE372CA4A837" blockId="1.[151,1437,392,2039]" pageId="1" pageNumber="70">
<emphasis id="07B7EA81FF9A9D15FF74FE372C0AACFB" bold="true" box="[151,266,392,418]" pageId="1" pageNumber="70">Diagnosis</emphasis>
. Superficially, members of the genus (
<figureCitation id="ADF82A16FF9A9D15FD5DFE372E30ACFA" box="[702,816,392,419]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 914" captionStartId-0="5.[151,264,1950,1974]" captionStartId-1="6.[151,264,1965,1989]" captionTargetBox-0="[173,1413,191,1919]" captionTargetBox-1="[163,1424,527,1934]" captionTargetId-0="figure-21@5.[166,1421,181,1926]" captionTargetId-1="figure-199@6.[151,1437,525,1942]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 18. Sicciaemorpha spp.: adults. Depositories of the specimens: 1 and 2 in RMNH; 3 and 58 in MWM/ZSM; 4 in CKC." captionText-1="FIGURES 914. Sicciaemorpha spp.: adults. Depositories of the specimens: 9 and 1113 in MWM/ZSM; 10 and 14 in CKC." figureDoi-0="http://doi.org/10.5281/zenodo.4929825" figureDoi-1="http://doi.org/10.5281/zenodo.4929827" httpUri-0="https://zenodo.org/record/4929825/files/figure.png" httpUri-1="https://zenodo.org/record/4929827/files/figure.png" pageId="1" pageNumber="70">Figs 114</figureCitation>
) are reminiscent of some narrow winged groups of the genera
<taxonomicName id="F2C34D10FF9A9D15FF08FE122C44AC9E" authorityName="Walker" authorityYear="1854" box="[235,324,429,455]" class="Insecta" family="Arctiidae" genus="Aemene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF08FE122C44AC9E" box="[235,324,429,455]" italics="true" pageId="1" pageNumber="70">Aemene</emphasis>
</taxonomicName>
and
<taxonomicName id="F2C34D10FF9A9D15FE9DFE122CC2AC9E" authorityName="Walker" authorityYear="1854" box="[382,450,429,455]" class="Insecta" family="Arctiidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FE9DFE122CC2AC9E" box="[382,450,429,455]" italics="true" pageId="1" pageNumber="70">Siccia</emphasis>
</taxonomicName>
. However, they can easily be distinguished by the bipectinate antennae of males. The male genitalia structure of
<taxonomicName id="F2C34D10FF9A9D15FE26FE6F2F6DACB3" box="[453,621,464,490]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FE26FE6F2F6DACB3" box="[453,621,464,490]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
(
<figureCitation id="ADF82A16FF9A9D15FD9EFE6F2FFDACB2" box="[637,765,464,491]" captionStart-0="FIGURES 1518" captionStart-1="FIGURES 1921" captionStartId-0="8.[151,264,1946,1970]" captionStartId-1="9.[151,264,1984,2008]" captionTargetBox-0="[171,1419,191,1914]" captionTargetBox-1="[247,1340,181,1951]" captionTargetId-0="figure-27@8.[164,1424,181,1923]" captionTargetId-1="figure-21@9.[234,1354,181,1960]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 1518. Sicciaemorpha spp.: male genitalia. Depositories of the specimens dissected: 15 in RMNH; 1618 in MWM/ ZSM." captionText-1="FIGURES 1921. Sicciaemorpha spp.: male genitalia. The specimens dissected are deposited in MWM/ZSM." figureDoi-0="http://doi.org/10.5281/zenodo.4929833" figureDoi-1="http://doi.org/10.5281/zenodo.4929835" httpUri-0="https://zenodo.org/record/4929833/files/figure.png" httpUri-1="https://zenodo.org/record/4929835/files/figure.png" pageId="1" pageNumber="70">Figs 1521</figureCitation>
) is most similar to that of the superficially dissimilar genus
<taxonomicName id="F2C34D10FF9A9D15FF74FE4B2C8FAF56" authority="Moore, 1882" authorityName="Moore" authorityYear="1882" box="[151,399,500,527]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF74FE4B2DF4AF57" box="[151,244,500,526]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
<bibRefCitation id="51524B62FF9A9D15FF1EFE4A2C8FAF56" author="Moore, F." box="[253,399,501,527]" pageId="1" pageNumber="70" pagination="72 - 107" refId="ref6632" refString="Moore, F. (1882) The Lepidoptera of Ceylon. Vol. 2. Part 1. L. Reeve &amp; Co., London, 72 pp., pls. 72 - 107." type="book chapter" year="1882">Moore, 1882</bibRefCitation>
</taxonomicName>
(illustrated by
<bibRefCitation id="51524B62FF9A9D15FDDEFE4B2FF9AF56" author="Holloway, J. D." box="[573,761,500,527]" pageId="1" pageNumber="70" pagination="279 - 486" refId="ref6564" refString="Holloway, J. D. (2001) The Moths of Borneo. Part 7. Family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 55, 279 - 486." type="journal article" year="2001">Holloway (2001</bibRefCitation>
: pl. 6, figs 18, 19)) due to the long membranous distal lobe taking up to 1/2 of the total length of the valva and separated from the costa, and the short costa terminating at the base of the membranous distal lobe and bearing a robust process directed dorsally. However, in the genital capsule of
<taxonomicName id="F2C34D10FF9A9D15FF5AFDDF2C61AF23" box="[185,353,608,634]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF5AFDDF2C61AF23" box="[185,353,608,634]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
, the tegumen and the vinculum are markedly longer (in comparison to the length of valvae) than in
<taxonomicName id="F2C34D10FF9A9D15FF0EFD3B2C4AAFC7" authorityName="Moore" authorityYear="1882" box="[237,330,644,670]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF0EFD3B2C4AAFC7" box="[237,330,644,670]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="51524B62FF9A9D15FE1BFD3B2FB4AFC6" author="Holloway, J. D." box="[504,692,644,671]" pageId="1" pageNumber="70" pagination="279 - 486" refId="ref6564" refString="Holloway, J. D. (2001) The Moths of Borneo. Part 7. Family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 55, 279 - 486." type="journal article" year="2001">Holloway (2001</bibRefCitation>
: gen. figs 319, 320)), the costal process is unilobate, triangular or stick-shaped (whereas that of
<taxonomicName id="F2C34D10FF9A9D15FE08FD172F48AF9B" authorityName="Moore" authorityYear="1882" box="[491,584,680,706]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FE08FD172F48AF9B" box="[491,584,680,706]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
is complex and usually bearing several various shaped processes), and the sclerotized ventro-distal lobe of the valva is present (an autapomorphic feature, absent in
<taxonomicName id="F2C34D10FF9A9D15FB93FD7329CDAFBF" authorityName="Moore" authorityYear="1882" box="[1136,1229,716,742]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FB93FD7329CDAFBF" box="[1136,1229,716,742]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
). Additionally, the sacculus of
<taxonomicName id="F2C34D10FF9A9D15FEF9FD4F2CC2AE53" box="[282,450,752,778]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FEF9FD4F2CC2AE53" box="[282,450,752,778]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
is long (ca. 1/2 of the valva length) and bearing a distal process, whereas in
<taxonomicName id="F2C34D10FF9A9D15FAF3FD4F286DAE53" authorityName="Moore" authorityYear="1882" box="[1296,1389,752,778]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FAF3FD4F286DAE53" box="[1296,1389,752,778]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
it is conspicuously shorter (ca. 1/3 of the valva length) and bearing a subbasal process directed distally. The aedeagus of
<taxonomicName id="F2C34D10FF9A9D15FF74FC872C3FAE0B" box="[151,319,824,850]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF74FC872C3FAE0B" box="[151,319,824,850]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
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is long (approximately the same length as the valva or somewhat longer in certain species) and thin (similar to those of
<taxonomicName id="F2C34D10FF9A9D15FE92FCE32FB1AE2E" authority="Rosenstock, 1885" authorityName="Rosenstock" authorityYear="1885" box="[369,689,860,887]" class="Insecta" family="Arctiidae" genus="Philenora" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FE92FCE32CE1AE2F" box="[369,481,860,886]" italics="true" pageId="1" pageNumber="70">Philenora</emphasis>
<bibRefCitation id="51524B62FF9A9D15FE04FCE32FB1AE2E" author="Rosenstock, R." box="[487,689,860,887]" pageId="1" pageNumber="70" pagination="376 - 385" refId="ref6670" refString="Rosenstock, R. (1885) Notes on Australian Lepidoptera, with descriptions of new species. The Annals and magazine of natural history, Series 5, 16, 376 - 385. https: // doi. org / 10.1080 / 00222938509459894" type="journal article" year="1885">Rosenstock, 1885</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F2C34D10FF9A9D15FD5FFCE32EC0AE2E" authority="Toulgoet, 1958" authorityName="Toulgoet" authorityYear="1958" box="[700,960,860,887]" class="Insecta" family="Arctiidae" genus="Exilisia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FD5FFCE32E12AE2F" box="[700,786,860,886]" italics="true" pageId="1" pageNumber="70">Exilisia</emphasis>
<bibRefCitation id="51524B62FF9A9D15FCF4FCE32EC0AE2E" author="Toulgoet, H. de" box="[791,960,860,887]" pageId="1" pageNumber="70" pagination="177 - 214" refId="ref6715" refString="Toulgoet, H. de (1958) Description d'Arctiidae nouveaux de Madagascar (Lepidoptera) (8 e note). Memoires de l'Institut scientifique de Madagascar, Serie E, 9, 177 - 214. [in French]" type="journal article" year="1958">Toulgoët, 1958</bibRefCitation>
</taxonomicName>
and related genera), and has a short vesica (ca. 1/3 of the aedeagus length) bearing a cluster of spinules, while in
<taxonomicName id="F2C34D10FF9A9D15FC7FFC3F2EF9AEC3" authorityName="Moore" authorityYear="1882" box="[924,1017,896,922]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FC7FFC3F2EF9AEC3" box="[924,1017,896,922]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
, the aedeagus is shorter and wider (in comparison to the genital capsule) and the vesica is much broader and having diverticula and fields of granulation. The female genitalia of
<taxonomicName id="F2C34D10FF9A9D15FE41FC772F4AAEBB" box="[418,586,968,994]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FE41FC772F4AAEBB" box="[418,586,968,994]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
(
<figureCitation id="ADF82A16FF9A9D15FDBAFC772FDAAEBA" box="[601,730,968,995]" captionStart="FIGURES 2227" captionStartId="10.[151,264,1836,1860]" captionTargetBox="[152,1429,190,1804]" captionTargetId="figure-27@10.[151,1435,181,1812]" captionTargetPageId="10" captionText="FIGURES 2227. Sicciaemorpha spp.: female genitalia. Depositories of the specimens dissected: 22 in RMNH; 23, 25 and 27 in CKC; 24 and 26 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.4929839" httpUri="https://zenodo.org/record/4929839/files/figure.png" pageId="1" pageNumber="70">Figs 2227</figureCitation>
) are fundamentally different from those of
<taxonomicName id="F2C34D10FF9A9D15FB5CFC77281CAEBB" authorityName="Moore" authorityYear="1882" box="[1215,1308,968,994]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FB5CFC77281CAEBB" box="[1215,1308,968,994]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="51524B62FF9A9D15FF59FC532C75A95E" author="Holloway, J. D." box="[186,373,1004,1031]" pageId="1" pageNumber="70" pagination="279 - 486" refId="ref6564" refString="Holloway, J. D. (2001) The Moths of Borneo. Part 7. Family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 55, 279 - 486." type="journal article" year="2001">Holloway (2001</bibRefCitation>
: gen. fig. 328)) in the following aspects. The antrum of
<taxonomicName id="F2C34D10FF9A9D15FC0BFC532990A95F" box="[1000,1168,1004,1030]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FC0BFC532990A95F" box="[1000,1168,1004,1030]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
is long (from 1/2 to 3/4 of the ductus bursae length) and more or less caliciform, having a deep ventral incision and is fully fused with 7
<superScript id="C2B69BDBFF9A9D15FA73FBAF289DA947" attach="none" box="[1424,1437,1040,1054]" fontSize="6" pageId="1" pageNumber="70">th</superScript>
abdominal segment which are well-sclerotized and bearing ventro-lateral pockets, whereas the antrum of
<taxonomicName id="F2C34D10FF9A9D15FADCFB8B289CA917" authorityName="Moore" authorityYear="1882" box="[1343,1436,1076,1102]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FADCFB8B289CA917" box="[1343,1436,1076,1102]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
is short (ca. 1/3 of the ductus bursae length) and trapezoid, and the 7
<superScript id="C2B69BDBFF9A9D15FC7DFBE72EABA93F" attach="left" box="[926,939,1112,1126]" fontSize="6" pageId="1" pageNumber="70">th</superScript>
abdominal segments are weakly sclerotized and lack pockets. The anterior section of the ductus bursae of
<taxonomicName id="F2C34D10FF9A9D15FCA1FBC32EEAA9CF" box="[834,1002,1148,1174]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FCA1FBC32EEAA9CF" box="[834,1002,1148,1174]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
is narrow (ca. 1/2 of the antrum width) and membranous, whereas it is wide (almost wide as the antrum width) and bearing elongate clusters of spines in
<taxonomicName id="F2C34D10FF9A9D15FF74FB7B2DF4A987" authorityName="Moore" authorityYear="1882" box="[151,244,1220,1246]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FF74FB7B2DF4A987" box="[151,244,1220,1246]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
. The corpus bursae of
<taxonomicName id="F2C34D10FF9A9D15FE1EFB7B2FA5A987" box="[509,677,1220,1246]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FE1EFB7B2FA5A987" box="[509,677,1220,1246]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
is globular and lacking spines, whereas it is sack-like and bearing wide clusters of spines in
<taxonomicName id="F2C34D10FF9A9D15FE5FFB572F19A85B" authorityName="Moore" authorityYear="1882" box="[444,537,1256,1282]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FE5FFB572F19A85B" box="[444,537,1256,1282]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
. Additionally, the appendix bursae of
<taxonomicName id="F2C34D10FF9A9D15FC2BFB572970A85B" box="[968,1136,1256,1282]" class="Insecta" family="Arctiidae" genus="Sicciaemorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FC2BFB572970A85B" box="[968,1136,1256,1282]" italics="true" pageId="1" pageNumber="70">Sicciaemorpha</emphasis>
</taxonomicName>
is small (in comparison to the corpus bursae), gelatinous and positioned ventro-posteriorly at the junction of the ductus and the corpus bursae, whereas in
<taxonomicName id="F2C34D10FF9A9D15FEF4FA8F2C74A813" authorityName="Moore" authorityYear="1882" box="[279,372,1328,1354]" class="Insecta" family="Arctiidae" genus="Padenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="07B7EA81FF9A9D15FEF4FA8F2C74A813" box="[279,372,1328,1354]" italics="true" pageId="1" pageNumber="70">Padenia</emphasis>
</taxonomicName>
the appendix bursae is large (about 1/4 of the size of the corpus bursae), conical, membranous, and positioned laterally.
</paragraph>
</subSubSection>
<subSubSection id="7DD96518FF9A9D16FF24FAC72FE1ACD0" lastPageId="2" lastPageNumber="71" pageId="1" pageNumber="70" type="description">
<paragraph id="357C3693FF9A9D16FF24FAC72FE1ACD0" blockId="1.[151,1437,392,2039]" lastBlockId="2.[151,1437,151,465]" lastPageId="2" lastPageNumber="71" pageId="1" pageNumber="70">
<emphasis id="07B7EA81FF9A9D15FF24FAC72C51A8CB" bold="true" box="[199,337,1400,1426]" pageId="1" pageNumber="70">Description</emphasis>
.
<emphasis id="07B7EA81FF9A9D15FE81FAC62F65A8CA" bold="true" box="[354,613,1401,1427]" pageId="1" pageNumber="70">External morphology</emphasis>
(
<figureCitation id="ADF82A16FF9A9D15FD95FAC72FEDA8CA" box="[630,749,1400,1427]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 914" captionStartId-0="5.[151,264,1950,1974]" captionStartId-1="6.[151,264,1965,1989]" captionTargetBox-0="[173,1413,191,1919]" captionTargetBox-1="[163,1424,527,1934]" captionTargetId-0="figure-21@5.[166,1421,181,1926]" captionTargetId-1="figure-199@6.[151,1437,525,1942]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 18. Sicciaemorpha spp.: adults. Depositories of the specimens: 1 and 2 in RMNH; 3 and 58 in MWM/ZSM; 4 in CKC." captionText-1="FIGURES 914. Sicciaemorpha spp.: adults. Depositories of the specimens: 9 and 1113 in MWM/ZSM; 10 and 14 in CKC." figureDoi-0="http://doi.org/10.5281/zenodo.4929825" figureDoi-1="http://doi.org/10.5281/zenodo.4929827" httpUri-0="https://zenodo.org/record/4929825/files/figure.png" httpUri-1="https://zenodo.org/record/4929827/files/figure.png" pageId="1" pageNumber="70">Figs 114</figureCitation>
). Forewing length 8.5
<quantity id="F23B9B76FF9A9D15FC18FAC7294AA8CA" box="[1019,1098,1400,1427]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="70" unit="mm" value="11.0">11 mm</quantity>
in both sexes. Sexual dimorphism limited: males have bipectinate antennae (ciliate in females) and less elongate forewing in comparison to females. Body coloration varies from white to brown in different species, thorax with black spots. Forewing elongate and narrow, with almost parallel costal and anal margins. Forewing pattern consists of numerous small black or dark brown spots. Hindwing without markings.
<emphasis id="07B7EA81FF9A9D15FD93F9B62E1AAB7B" bold="true" box="[624,794,1544,1571]" pageId="1" pageNumber="70">Male genitalia</emphasis>
(
<figureCitation id="ADF82A16FF9A9D15FCC4F9B72EA8AB7A" box="[807,936,1544,1571]" captionStart-0="FIGURES 1518" captionStart-1="FIGURES 1921" captionStartId-0="8.[151,264,1946,1970]" captionStartId-1="9.[151,264,1984,2008]" captionTargetBox-0="[171,1419,191,1914]" captionTargetBox-1="[247,1340,181,1951]" captionTargetId-0="figure-27@8.[164,1424,181,1923]" captionTargetId-1="figure-21@9.[234,1354,181,1960]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 1518. Sicciaemorpha spp.: male genitalia. Depositories of the specimens dissected: 15 in RMNH; 1618 in MWM/ ZSM." captionText-1="FIGURES 1921. Sicciaemorpha spp.: male genitalia. The specimens dissected are deposited in MWM/ZSM." figureDoi-0="http://doi.org/10.5281/zenodo.4929833" figureDoi-1="http://doi.org/10.5281/zenodo.4929835" httpUri-0="https://zenodo.org/record/4929833/files/figure.png" httpUri-1="https://zenodo.org/record/4929835/files/figure.png" pageId="1" pageNumber="70">Figs 1521</figureCitation>
). Uncus elongate and thin, curved, somewhat dilated subbasally or medially. Tuba analis relatively narrow (in comparison to its length), scaphium thin but heavily sclerotized. Tegumen moderately long (ca. 2/3 of valva length) with relatively narrow arms. Vinculum somewhat shorter than tegumen, with short (ca. 1/4 of its length) and moderately sclerotized, U-shaped or rectangular saccus. Juxta elongate shield-like. Costa of valva short (terminating at base of membranous distal lobe), with robust and apically pointed process directed dorsally and ventral protrusion almost reaching base of ventro-distal process of valva. Distal lobe of valva large (ca. 1/2 of valva length), elongate, membranous. Ventro-distal process of valva large (from 1/4 to 3/4 of valva length), heavily sclerotized, fused basally with basal section of distal saccular process, directed distally or ventro-distally. Sacculus narrow (from 1/3 to 1/2 of valva width), with heavily sclerotized and apically pointed distal process directed distally or somewhat curved dorsally. Aedeagus long (approximately same length as valva or somewhat longer) and thin, in certain species somewhat curved distally, with well-developed coecum and elongate, thin and apically pointed carina. Vesica short (ca. 1/3 of the aedeagus length), narrow (as aedeagus width of somewhat broader), curved laterally, and bearing dense cluster of small spinules.
<emphasis id="07B7EA81FF9A9D15FB76F806285BAA8B" bold="true" box="[1173,1371,1976,2003]" pageId="1" pageNumber="70">Female genitalia</emphasis>
(
<figureCitation id="ADF82A16FF9A9D15FA89F8072DE2AAAE" captionStart="FIGURES 2227" captionStartId="10.[151,264,1836,1860]" captionTargetBox="[152,1429,190,1804]" captionTargetId="figure-27@10.[151,1435,181,1812]" captionTargetPageId="10" captionText="FIGURES 2227. Sicciaemorpha spp.: female genitalia. Depositories of the specimens dissected: 22 in RMNH; 23, 25 and 27 in CKC; 24 and 26 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.4929839" httpUri="https://zenodo.org/record/4929839/files/figure.png" pageId="1" pageNumber="70">Figs 2227</figureCitation>
). Papillae anales rectangular with rounded corners, weakly setose. Apophyses posteriores elongate and thin, apophyses anteriores thin, approximately twice or three times shorter than posteriores ones. Ductus bursae elongate and relatively narrow (length with width ratio from 3:1 to 4:1), its anterior section membranous. Antrum wellsclerotized, long (ca. 1/2 to 3/4 of ductus bursae length), round or caliciform, with deep incision ventrally. Margins of antrum fused with sclerotized 7
<superScript id="C2B69BDBFF999D16FDC2FEBD2F2EAC49" attach="left" box="[545,558,258,272]" fontSize="6" pageId="2" pageNumber="71">th</superScript>
sternite forming entire structure with strongly elongate postvaginal area and shallow ventro-lateral pockets on the sides of ostium bursae. Corpus bursae globular, membranous with very weak granulation. Appendix bursae short and narrow (in comparison to corpus bursae), gelatinous, positioned ventrally or laterally at the junction of ductus and corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="7DD96518FF999D16FF24FE2D2F35AC88" pageId="2" pageNumber="71" type="distribution">
<paragraph id="357C3693FF999D16FF24FE2D2F35AC88" blockId="2.[151,1437,151,465]" pageId="2" pageNumber="71">
<emphasis id="07B7EA81FF999D16FF24FE2D2C59ACF5" bold="true" box="[199,345,402,428]" pageId="2" pageNumber="71">Distribution</emphasis>
. Known from the islands of
<collectingRegion id="F707F871FF999D16FD7EFE2C2FCFACF4" box="[669,719,403,429]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="71">Java</collectingRegion>
(
<collectingCountry id="4DD47603FF999D16FD3DFE2C2E51ACF7" box="[734,849,403,430]" name="Indonesia" pageId="2" pageNumber="71">Indonesia</collectingCountry>
), Borneo (
<collectingCountry id="4DD47603FF999D16FC25FE2C2930ACF7" box="[966,1072,403,430]" name="Malaysia" pageId="2" pageNumber="71">Malaysia</collectingCountry>
) and the Philippine Archipelago where it reaches its highest diversity.
</paragraph>
</subSubSection>
</treatment>
</document>