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<document id="9ADC068429350CAC9078145EF012DFC1" ID-DOI="10.11646/zootaxa.3941.4.7" ID-GBIF-Dataset="68e478d2-0f12-4797-9516-c6ceeec684f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="234951" ID-ZooBank="B6D8072E-C183-4467-BF62-2D6DB1698968" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461028833841" checkinUser="plazi" docAuthor="Zettel, Herbert &amp; Pangantihon, Clister V." docDate="2015" docId="795F824C514A966E5393FAFFFD78FA4C" docLanguage="en" docName="zt03941p588.pdf" docOrigin="Zootaxa 3941 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Myriochila philippinensis Mandl 1956" docType="treatment" docVersion="8" lastPageNumber="586" masterDocId="8566FA34514A966F5304FFE1FFD7FF98" masterDocTitle="On the identity of the tiger beetle Cicindela philippinensis Mandl, 1956 (Coleoptera: Carabidae: Cicindelinae)" masterLastPageNumber="588" masterPageNumber="585" pageNumber="585" updateTime="1698573008746" updateUser="plazi">
<mods:mods id="F600DBA871CBC3769AC77DE83638BA0C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CEB978D712D0D47901736315FD8A477B">
<mods:title id="5238F400F47B26986FC52E6FD470E81A">On the identity of the tiger beetle Cicindela philippinensis Mandl, 1956 (Coleoptera: Carabidae: Cicindelinae)</mods:title>
</mods:titleInfo>
<mods:name id="708A58BE3A2E3D03CBE1A69EA8847FB8" type="personal">
<mods:role id="C6A9ACA7EF20801AE04B593D0BB3C46B">
<mods:roleTerm id="B443DC15A5DB37B066799B8697F7787E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D1E0B7BC485F9AE507E42D5790F6C52D">Zettel, Herbert</mods:namePart>
</mods:name>
<mods:name id="274A15B293D5EE0489FCD95BB43983B6" type="personal">
<mods:role id="D56F1ACE600E8317A0E7434DE1B6A292">
<mods:roleTerm id="6219AB80E04F962ED4037394E7E8FCE9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="772D0582801E6A73767C95D147AFEA92">Pangantihon, Clister V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B25C8E02DC06C19AF580CE0735AD8736">text</mods:typeOfResource>
<mods:relatedItem id="156AF22D9B9AC34CA4D8FE6B77E82F5E" type="host">
<mods:titleInfo id="C27ED2389F22AE80C1F2D61CFFD0FA8B">
<mods:title id="F31A514908B73B029CA8874B4A0E099F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="83AF0EE725BC6581B44A118B9E9E592B">
<mods:date id="00F240CD22FE7CF71EB7234C452C677D">2015</mods:date>
<mods:detail id="DF5BDE47C2E5AC3DBE8D33D2A2AC6671" type="volume">
<mods:number id="687A1073027FE70FA4C29D10FEAFE163">3941</mods:number>
</mods:detail>
<mods:detail id="F5126515B74E15F579202B455FE3B7D3" type="issue">
<mods:number id="3E1193D3B52598B0EC83CE4688C00D87">4</mods:number>
</mods:detail>
<mods:extent id="E9EB0004F87FBB3206CDD6C39E645939" unit="page">
<mods:start id="FEE6836EA715854799EF700F6CF70B3C">585</mods:start>
<mods:end id="8E56789B75A36165427DD5128E465D85">588</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BFE76D9C2723735325A9158FF056BF06">journal article</mods:classification>
<mods:identifier id="5F64627066E81B9A0BA31ED95B1EF469" type="DOI">10.11646/zootaxa.3941.4.7</mods:identifier>
<mods:identifier id="32A2F856346010F48CA921999DAA786C" type="GBIF-Dataset">68e478d2-0f12-4797-9516-c6ceeec684f5</mods:identifier>
<mods:identifier id="A88081C87471AD24C65C5C48FC62DE84" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E51974E901B186F77614C884532CFA93" type="Zenodo-Dep">234951</mods:identifier>
<mods:identifier id="CAA25C9DC1E2374F6446275A1F9D58BA" type="ZooBank">B6D8072E-C183-4467-BF62-2D6DB1698968</mods:identifier>
</mods:mods>
<treatment id="795F824C514A966E5393FAFFFD78FA4C" ID-DOI="http://doi.org/10.5281/zenodo.6120898" ID-GBIF-Taxon="119607164" ID-Zenodo-Dep="6120898" LSID="urn:lsid:plazi:treatment:795F824C514A966E5393FAFFFD78FA4C" httpUri="http://treatment.plazi.org/id/795F824C514A966E5393FAFFFD78FA4C" lastPageId="1" lastPageNumber="586" pageId="0" pageNumber="585">
<subSubSection id="B9EC60D1514A966F5393FAFFFEDBFAC0" pageId="0" pageNumber="585" type="nomenclature">
<paragraph id="F149335A514A966F5393FAFFFD53FAA1" blockId="0.[151,644,1310,1368]" box="[151,644,1310,1337]" pageId="0" pageNumber="585">
<heading id="AA018436514A966F5393FAFFFD53FAA1" bold="true" box="[151,644,1310,1337]" fontSize="11" level="1" pageId="0" pageNumber="585" reason="1">
<taxonomicName id="36F648D9514A966F5393FAFFFD53FAA1" ID-CoL="458R6" authority="Mandl, 1956" authorityName="Mandl" authorityYear="1956" box="[151,644,1310,1337]" class="Insecta" family="Carabidae" genus="Myriochila" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514A966F5393FAFFFD53FAA1" bold="true" box="[151,644,1310,1337]" pageId="0" pageNumber="585">
<emphasis id="C382EF48514A966F5393FAFFFE1AFAA0" bold="true" box="[151,461,1310,1336]" italics="true" pageId="0" pageNumber="585">Myriochila philippinensis</emphasis>
(
<bibRefCitation id="95674EAB514A966F52D8FAFEFDABFAA1" author="Mandl" box="[476,636,1311,1337]" pageId="0" pageNumber="585" refString="Mandl, K. (1956) Neun neue Cicindelidenformen aus tropischen Landern. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 7 (1956), 378 - 397." type="journal article" year="1956">Mandl, 1956</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F149335A514A966F5393FAA0FEDBFAC0" blockId="0.[151,644,1310,1368]" box="[151,268,1345,1368]" pageId="0" pageNumber="585">
(
<figureCitation id="69CD2FDF514A966F539BFAA0FED4FAC0" box="[159,259,1345,1368]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="0" pageNumber="585">Figs. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B9EC60D1514A966F5393FA66FB13FA7B" pageId="0" pageNumber="585" type="reference_group">
<paragraph id="F149335A514A966F5393FA66FCFCFA06" blockId="0.[151,1220,1415,1507]" box="[151,811,1415,1438]" pageId="0" pageNumber="585">
<treatmentCitationGroup id="D1E61474514A966F5393FA66FCFCFA06" box="[151,811,1415,1438]" pageId="0" pageNumber="585">
<taxonomicName id="36F648D9514A966F5393FA66FDB7FA06" ID-CoL="5Z6L6" authority="Mandl, 1956: 396" authorityName="Mandl" authorityPageNumber="396" authorityYear="1956" box="[151,608,1415,1438]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514A966F5393FA66FE4DFA06" box="[151,410,1415,1438]" italics="true" pageId="0" pageNumber="585">Cicindela philippinensis</emphasis>
<treatmentCitation id="7057154B514A966F52A5FA66FDB7FA06" author="Mandl" box="[417,608,1415,1438]" page="396" pageId="0" pageNumber="585" year="1956">
<bibRefCitation id="95674EAB514A966F52A5FA66FDFAFA06" author="Mandl" box="[417,557,1415,1438]" pageId="0" pageNumber="585" refString="Mandl, K. (1956) Neun neue Cicindelidenformen aus tropischen Landern. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 7 (1956), 378 - 397." type="journal article" year="1956">Mandl, 1956</bibRefCitation>
: 396
</treatmentCitation>
</taxonomicName>
–397 (description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F149335A514A966F5393FA48FB13FA7B" blockId="0.[151,1220,1415,1507]" pageId="0" pageNumber="585">
<treatmentCitationGroup id="D1E61474514A966F5393FA48FB13FA7B" pageId="0" pageNumber="585">
<taxonomicName id="36F648D9514A966F5393FA48FD9BFA58" authority="Mandl, 1956" authorityName="Mandl" authorityYear="1956" box="[151,588,1449,1472]" class="Insecta" family="Carabidae" genus="Myriochile" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514A966F5393FA48FE70FA58" box="[151,423,1449,1472]" italics="true" pageId="0" pageNumber="585">Myriochile philippinensis</emphasis>
(
<bibRefCitation id="95674EAB514A966F52B2FA48FD92FA58" author="Mandl" box="[438,581,1449,1472]" pageId="0" pageNumber="585" refString="Mandl, K. (1956) Neun neue Cicindelidenformen aus tropischen Landern. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 7 (1956), 378 - 397." type="journal article" year="1956">Mandl, 1956</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="7057154B514A966F5153FA48FCF1FA58" author="Wiesner" box="[599,806,1449,1472]" page="126" pageId="0" pageNumber="585" year="1980">
<bibRefCitation id="95674EAB514A966F5153FA48FD24FA58" author="Wiesner" box="[599,755,1449,1472]" pageId="0" pageNumber="585" refString="Wiesner, J. (1980) Beitrage zur Kenntnis der philippinischen Cicindelidae (Coleoptera). Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 119 - 127." type="journal article" year="1980">Wiesner, 1980</bibRefCitation>
: 126
</treatmentCitation>
(in list of
<collectingCountry id="89E173CA514A966F5091FA48FBD5FA58" box="[917,1026,1449,1472]" name="Philippines" pageId="0" pageNumber="585">Philippine</collectingCountry>
tiger beetles).
<taxonomicName id="36F648D9514A966F5393FA2DFEDBFA7B" ID-CoL="8LS9S" box="[151,268,1484,1507]" class="Insecta" family="Carabidae" genus="Myriochile" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="C382EF48514A966F5393FA2DFEDBFA7B" box="[151,268,1484,1507]" italics="true" pageId="0" pageNumber="585">Myriochile</emphasis>
</taxonomicName>
?
<taxonomicName id="36F648D9514A966F5221FA2DFD88FA7B" authority="Mandl, 1956" authorityName="Mandl" authorityYear="1956" box="[293,607,1484,1507]" class="Insecta" family="Carabidae" genus="Myriochile" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514A966F5221FA2DFE6DFA7B" box="[293,442,1484,1507]" italics="true" pageId="0" pageNumber="585">philippinensis</emphasis>
(
<bibRefCitation id="95674EAB514A966F52CDFA2DFD80FA7B" author="Mandl" box="[457,599,1484,1507]" pageId="0" pageNumber="585" refString="Mandl, K. (1956) Neun neue Cicindelidenformen aus tropischen Landern. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 7 (1956), 378 - 397." type="journal article" year="1956">Mandl, 1956</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="7057154B514A966F516DFA2DFCEFFA7B" author="Wiesner" box="[617,824,1484,1507]" page="210" pageId="0" pageNumber="585" year="1992">
<bibRefCitation id="95674EAB514A966F516DFA2DFCD1FA7B" author="Wiesner" box="[617,774,1484,1507]" pageId="0" pageNumber="585" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt. Checklist of the tiger beetles of the world. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">Wiesner, 1992</bibRefCitation>
: 210
</treatmentCitation>
(in world catalogue of tiger beetles).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B9EC60D1514A966F5393F9F0FD52F961" pageId="0" pageNumber="585" type="materials_examined">
<paragraph id="F149335A514A966F5393F9F0FD07F9D3" blockId="0.[151,1436,1553,2027]" pageId="0" pageNumber="585">
<emphasis id="C382EF48514A966F5393F9F0FE60F9B0" bold="true" box="[151,439,1553,1576]" pageId="0" pageNumber="585">
<typeStatus id="2E4D8DF8514A966F5393F9F0FF18F9B0" box="[151,207,1553,1576]" pageId="0" pageNumber="585">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="2E4D8DF8514A966F52C4F9F0FDF4F9B0" box="[448,547,1553,1576]" pageId="0" pageNumber="585" type="holotype">Holotype</typeStatus>
(male, Catalogue Number 63, Naturhistorisches Museum Basel), labelled &quot;Pilar. Abra
<collectingCountry id="89E173CA514A966F53D7F9D5FE99F9D3" box="[211,334,1588,1611]" name="Philippines" pageId="0" pageNumber="585">Philippinen</collectingCountry>
R.P. Niedurny&quot;, labels see
<figureCitation id="69CD2FDF514A966F5174F9D5FD1BF9D3" box="[624,716,1588,1611]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="0" pageNumber="585">Figure 7</figureCitation>
.
</paragraph>
<paragraph id="F149335A514A966F53C3F9B6FB14F92B" blockId="0.[151,1436,1553,2027]" pageId="0" pageNumber="585">
<emphasis id="C382EF48514A966F53C3F9B6FE11F9F6" bold="true" box="[199,454,1623,1646]" pageId="0" pageNumber="585">
Notes on the
<typeStatus id="2E4D8DF8514A966F5259F9B6FE16F9F6" box="[349,449,1623,1646]" pageId="0" pageNumber="585" type="holotype">holotype</typeStatus>
.
</emphasis>
The specimen is in a regular state of conservation. It seems to have been originally pinned and later glued onto a cardboard. Left antenna, left foreleg, left hind leg, and distal part (tibia and tarsus) of right hind leg are missing. The right elytron was broken and glued to the abdomen; its apical margin is broken off.
</paragraph>
<paragraph id="F149335A514A966F53C3F95EFD52F961" blockId="0.[151,1436,1553,2027]" pageId="0" pageNumber="585">
<emphasis id="C382EF48514A966F53C3F95EFE3CF94E" bold="true" box="[199,491,1727,1750]" pageId="0" pageNumber="585">
Notes on the
<typeStatus id="2E4D8DF8514A966F525FF95EFE5BF94E" box="[347,396,1727,1750]" pageId="0" pageNumber="585">type</typeStatus>
locality.
</emphasis>
Pilar (ca.
<geoCoordinate id="94C2559D514A966F515EF95EFD6DF94E" box="[602,698,1727,1750]" direction="north" orientation="latitude" pageId="0" pageNumber="585" precision="925" value="17.416666">N 17°25'</geoCoordinate>
<geoCoordinate id="94C2559D514A966F51C4F95EFCFBF94E" box="[704,812,1727,1750]" direction="east" orientation="longitude" pageId="0" pageNumber="585" precision="925" value="120.6">E 120°36'</geoCoordinate>
, ca.
<quantity id="360E9EBF514A966F5058F95EFC77F94E" box="[860,928,1727,1750]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="585" unit="m" value="300.0">300 m</quantity>
a.s.l.) is a municipality in Abra Province in the northern part of Luzon Island, the
<collectingCountry id="89E173CA514A966F5101F903FD56F961" box="[517,641,1762,1785]" name="Philippines" pageId="0" pageNumber="585">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B9EC60D1514A966E53C3F8E5FD78FA4C" lastPageId="1" lastPageNumber="586" pageId="0" pageNumber="585" type="description">
<paragraph id="F149335A514A966F53C3F8E5FD20F8A6" blockId="0.[151,1436,1553,2027]" pageId="0" pageNumber="585">
<emphasis id="C382EF48514A966F53C3F8E5FEBBF883" bold="true" box="[199,364,1796,1819]" pageId="0" pageNumber="585">Redescription.</emphasis>
Measurements: Total length ca.
<quantity id="360E9EBF514A966F51CFF8E5FCF6F883" box="[715,801,1796,1819]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="0" pageNumber="585" unit="mm" value="8.2">8.2 mm</quantity>
. Head width
<quantity id="360E9EBF514A966F50ABF8E5FBC5F883" box="[943,1042,1796,1819]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="0" pageNumber="585" unit="mm" value="2.25">2.25 mm</quantity>
. Pronotum width
<quantity id="360E9EBF514A966F57D5F8E5FAE4F883" box="[1233,1331,1796,1819]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="0" pageNumber="585" unit="mm" value="1.6">1.60 mm</quantity>
, length at midline
<quantity id="360E9EBF514A966F53EBF8C6FE86F8A6" box="[239,337,1831,1854]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="0" pageNumber="585" unit="mm" value="1.54">1.54 mm</quantity>
. Sutural length of left elytron
<quantity id="360E9EBF514A966F5195F8C6FD24F8A6" box="[657,755,1831,1854]" metricMagnitude="-3" metricUnit="m" metricValue="4.81" pageId="0" pageNumber="585" unit="mm" value="4.81">4.81 mm</quantity>
.
</paragraph>
<paragraph id="F149335A514A966E53C3F8ABFA4BFE7F" blockId="0.[151,1436,1553,2027]" lastBlockId="1.[151,1437,152,1492]" lastPageId="1" lastPageNumber="586" pageId="0" pageNumber="585">
Colour and pilosity (
<figureCitation id="69CD2FDF514A966F52AFF8ABFDC2F8F9" box="[427,533,1866,1889]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="0" pageNumber="585">
Figs. 1–
<docIssue id="B5954CB5514A966F5101F8ABFDC2F8F9" box="[517,533,1866,1889]" pageId="0" pageNumber="585">4</docIssue>
</figureCitation>
): Head dorsally bronze black with metallic purple to greenish reflection, ventrally with strong blue reflection; antennal plate and suborbital declivity to genae with strong bluish green reflection; clypeus metallic bronze with green reflection. Labrum pale yellow to testaceous, with black apical margin. Eyes dark brown to grey. Mandibles basally testaceous, apically and on teeth dark rufous to black, with strong green reflection at mid-length. Labial and maxillary palpi testaceous except all terminal palpomeres metallic green, and penultimate segment of maxillary palpi dark brown. Antenna: scape to antennomere 4 bronze black with greenish and purple reflection; antennomeres 5–11 brown, entirely covered with white pilosity. Pronotum bronze with metallic green to bluish reflection; posterior lateral tubercles metallic green with strong reflection of bronze on both sides; sides of front and hind lobe black with bronze to green reflection. Scutellum bronze to purple. Elytra bronze black, with basally greenish reflection, with purple shimmer (more on sides); grooves green, their centres often blue; white to yellowish markings strongly reduced; epipleura dark testaceous, with strong purple reflection. Sides and venter (as far as visible) bronze black. Legs almost entirely with metallic reflection, except trochanters dark testaceous; coxae and femora chiefly green, tibiae chiefly bronze, tarsi basally green tending to violet and blue towards apex; apices of femora and tibiae polychromous. Maxilla and labial palps with some long white hairs, especially penultimate segment of labial palp with dense brush. Sides of prothorax with some white hairs ventrally (
<figureCitation id="69CD2FDF514B966E5043FE31FC50FE7F" box="[839,903,464,487]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 3</figureCitation>
). Venter with numerous long, slender, white hairs.
</paragraph>
<paragraph id="F149335A514B966E53C3FE13FAB3FD64" blockId="1.[151,1437,152,1492]" pageId="1" pageNumber="586">
Structures: Head fully glabrous, except two pairs of short orbital setae (anteriorly and at mid-length of eye). Frons finely shagreened, vertex finely, irregularly wrinkled, orbitals longitudinally creased (
<figureCitation id="69CD2FDF514B966E575AFDF4FB71FDB4" box="[1118,1190,533,556]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 2</figureCitation>
). Genae and tempora longitudinally striate and somewhat shiny (
<figureCitation id="69CD2FDF514B966E5166FDD9FD75FDD7" box="[610,674,568,591]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 3</figureCitation>
). Orbitals and frons almost in the same plane. Eyes large, their smallest distance 0.39 times head width. Clypeus (
<figureCitation id="69CD2FDF514B966E5164FDBBFD74FDE9" box="[608,675,602,625]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 4</figureCitation>
) narrow, finely shagreened, without setae. Labrum (
<figureCitation id="69CD2FDF514B966E57DBFDBBFAF6FDE9" box="[1247,1313,602,625]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 4</figureCitation>
) shiny and almost smooth, with hardly discernable reticulate microstructure, medially protruded, 2.2 times as wide as long, with two pairs of setae, one laterally and one near midline; foremargin with one distinct median tooth and two blunt lobes laterally of it. Mandible long, with four slender teeth (including apex). Antenna (
<figureCitation id="69CD2FDF514B966E50A0FD23FBDBFD41" box="[932,1036,706,729]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Figs. 1, 5</figureCitation>
) long, in male reaching to middle of elytra, antennal segments cylindrical, segments 1–4 shiny, with extremely fine microstructure, segments 5–11 matt.
</paragraph>
<paragraph id="F149335A514B966E53C3FCE9FE7BFC1F" blockId="1.[151,1437,152,1492]" pageId="1" pageNumber="586">
Pronotum (
<figureCitation id="69CD2FDF514B966E5245FCE9FE53FC87" box="[321,388,776,799]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 2</figureCitation>
) minimally wider than long, with moderately convex sides, hardly constricted before apex and base; transverse furrows near anterior and posterior margins shallow, but distinct, sinuate; midline hardly impressed; disk with coarse wrinkles; sides finely shagreened and with some superficial furrows. Scutellum finely shagreened and with some transverse wrinkles.
</paragraph>
<paragraph id="F149335A514B966E53C3FC73FC5AFB89" blockId="1.[151,1437,152,1492]" pageId="1" pageNumber="586">
Elytra (
<figureCitation id="69CD2FDF514B966E521DFC73FE8EFC31" box="[281,345,914,937]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 1</figureCitation>
) with numerous small shallow grooves and several deeper pits (some of them bearing short setae); deep and rather large pits distributed along lateral margins, and along medial margin next to scutellum, in a row in some distance from suture, and in a double row medially of humerus. Apex (
<figureCitation id="69CD2FDF514B966E5087FC39FC15FC77" box="[899,962,984,1007]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 6</figureCitation>
) almost rectangular, with an extremely short sutural spine. Posterior margin with a minute, hardly discernable tooth.
</paragraph>
<paragraph id="F149335A514B966E53C3FBFCFCFEFBCF" blockId="1.[151,1437,152,1492]" pageId="1" pageNumber="586">
Legs very slender, moderately long. Tarsomeres 1–3 of foreleg (
<figureCitation id="69CD2FDF514B966E506AFBFCFC79FBAC" box="[878,942,1053,1076]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 5</figureCitation>
) moderately widened, almost twice as wide as tarsomeres 4 and 5; third tarsomere 2.9 times as long as wide.
</paragraph>
<paragraph id="F149335A514B966E53C3FB83FAB8FBE1" blockId="1.[151,1437,152,1492]" box="[199,1391,1122,1145]" pageId="1" pageNumber="586">Venter (as far as visible in the glued specimen) matt, densely shagreened and with strong punctures at hair base.</paragraph>
<paragraph id="F149335A514B966E53C3FB64FA45FB04" blockId="1.[151,1437,152,1492]" box="[199,1426,1157,1180]" pageId="1" pageNumber="586">
Aedeagus: Not dissected due to the condition of the unique specimen. Apex of median lobe externally visible (
<figureCitation id="69CD2FDF514B966E5642FB64FA52FB04" box="[1350,1413,1157,1180]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="F149335A514B966E53C3FB49FD78FA4C" blockId="1.[151,1437,152,1492]" pageId="1" pageNumber="586">
<emphasis id="C382EF48514B966E53C3FB49FE70FB27" bold="true" box="[199,423,1192,1215]" pageId="1" pageNumber="586">Comparative notes:</emphasis>
<taxonomicName id="36F648D9514B966E52AAFB49FD17FB27" box="[430,704,1192,1215]" class="Insecta" family="Carabidae" genus="Myriochila" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514B966E52AAFB49FD17FB27" box="[430,704,1192,1215]" italics="true" pageId="1" pageNumber="586">Myriochila philippinensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="36F648D9514B966E5048FB49FC78FB27" box="[844,943,1192,1215]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="C382EF48514B966E5048FB49FC78FB27" box="[844,943,1192,1215]" italics="true" pageId="1" pageNumber="586">M. sinica</emphasis>
</taxonomicName>
.
<bibRefCitation id="95674EAB514B966E50B8FB49FB87FB27" author="Mandl" box="[956,1104,1192,1215]" pageId="1" pageNumber="586" refString="Mandl, K. (1956) Neun neue Cicindelidenformen aus tropischen Landern. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 7 (1956), 378 - 397." type="journal article" year="1956">Mandl (1956)</bibRefCitation>
stated that
<taxonomicName id="36F648D9514B966E57CEFB49FA54FB27" box="[1226,1411,1192,1215]" class="Insecta" family="Carabidae" genus="Myriochila" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514B966E57CEFB49FA54FB27" box="[1226,1411,1192,1215]" italics="true" pageId="1" pageNumber="586">M. philippinensis</emphasis>
</taxonomicName>
is smaller than
<taxonomicName id="36F648D9514B966E5220FB2BFE5DFB79" box="[292,394,1226,1249]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="C382EF48514B966E5220FB2BFE5DFB79" box="[292,394,1226,1249]" italics="true" pageId="1" pageNumber="586">M. sinica</emphasis>
</taxonomicName>
, but his measurement (&quot;Länge:
<quantity id="360E9EBF514B966E51ECFB2BFCFCFB79" box="[744,811,1226,1249]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="586" unit="mm" value="7.0">7 mm</quantity>
&quot;) was incorrect. We have studied specimens of
<taxonomicName id="36F648D9514B966E5633FB2BFA4BFB79" box="[1335,1436,1226,1249]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="C382EF48514B966E5633FB2BFA4BFB79" box="[1335,1436,1226,1249]" italics="true" pageId="1" pageNumber="586">M. sinica</emphasis>
</taxonomicName>
from
<collectingCountry id="89E173CA514B966E53D6FB0CFEE4FA9C" box="[210,307,1261,1284]" name="Thailand" pageId="1" pageNumber="586">Thailand</collectingCountry>
,
<collectingCountry id="89E173CA514B966E523AFB0CFEA3FA9C" box="[318,372,1261,1284]" name="Laos" pageId="1" pageNumber="586">Laos</collectingCountry>
, and
<collectingCountry id="89E173CA514B966E52AAFB0CFDDFFA9C" box="[430,520,1261,1284]" name="Vietnam" pageId="1" pageNumber="586">Vietnam</collectingCountry>
in the Natural History Museum Vienna, and two males have a similarly small size as the
<typeStatus id="2E4D8DF8514B966E53BAFAF1FECCFABF" box="[190,283,1296,1319]" pageId="1" pageNumber="586" type="holotype">holotype</typeStatus>
of
<taxonomicName id="36F648D9514B966E523BFAF1FE2FFABF" box="[319,504,1296,1319]" class="Insecta" family="Carabidae" genus="Myriochila" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514B966E523BFAF1FE2FFABF" box="[319,504,1296,1319]" italics="true" pageId="1" pageNumber="586">M. philippinensis</emphasis>
</taxonomicName>
. The elytral colour pattern of
<taxonomicName id="36F648D9514B966E503EFAF1FC49FABF" box="[826,926,1296,1319]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="C382EF48514B966E503EFAF1FC49FABF" box="[826,926,1296,1319]" italics="true" pageId="1" pageNumber="586">M. sinica</emphasis>
</taxonomicName>
is rather variable. We found differences only in the structure of the labrum and the antenna:
<taxonomicName id="36F648D9514B966E517CFAD3FC5BFAD1" box="[632,908,1330,1353]" class="Insecta" family="Carabidae" genus="Myriochila" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514B966E517CFAD3FC5BFAD1" box="[632,908,1330,1353]" italics="true" pageId="1" pageNumber="586">Myriochila philippinensis</emphasis>
</taxonomicName>
has a median labrum tooth accompanied by two short blunt lobes (
<figureCitation id="69CD2FDF514B966E525CFAB4FE4EFAF4" box="[344,409,1365,1388]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 4</figureCitation>
), whereas
<taxonomicName id="36F648D9514B966E5108FAB4FDA7FAF4" box="[524,624,1365,1388]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="C382EF48514B966E5108FAB4FDA7FAF4" box="[524,624,1365,1388]" italics="true" pageId="1" pageNumber="586">M. sinica</emphasis>
</taxonomicName>
has three subequal teeth.
<taxonomicName id="36F648D9514B966E5080FAB4FB40FAF4" box="[900,1175,1365,1388]" class="Insecta" family="Carabidae" genus="Myriochila" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514B966E5080FAB4FB40FAF4" box="[900,1175,1365,1388]" italics="true" pageId="1" pageNumber="586">Myriochila philippinensis</emphasis>
</taxonomicName>
has cylindrical antennal segments 3 and 4 (
<figureCitation id="69CD2FDF514B966E525AFA99FE48FA17" box="[350,415,1400,1423]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1948,1971]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@2.[151,1436,184,1924]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Holotype of Cicindela philippinensis: (1) Habitus, dorsal view; (2) head and pronotum, dorsal view; (3) head and pronotum, lateral view; (4) head, frontal view; (5) basal antennal segments and right foretarsus; (6) apices of elytra and abdomen with protruded tip of aedeagus; (7) labels." httpUri="https://zenodo.org/record/234952/files/figure.png" pageId="1" pageNumber="586">Fig. 5</figureCitation>
), whereas they are compressed in
<taxonomicName id="36F648D9514B966E500AFA99FCA6FA17" box="[782,881,1400,1423]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="C382EF48514B966E500AFA99FCA6FA17" box="[782,881,1400,1423]" italics="true" pageId="1" pageNumber="586">M. sinica</emphasis>
</taxonomicName>
. Whether these differences are sufficient to hold
<taxonomicName id="36F648D9514B966E567BFA99FEFAFA29" class="Insecta" family="Carabidae" genus="Myriochila" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="C382EF48514B966E567BFA99FEFAFA29" italics="true" pageId="1" pageNumber="586">M. philippinensis</emphasis>
</taxonomicName>
as a distinct species, or are a matter of intraspecific variation or individual aberration must be decided after the study of more specimens from the
<collectingCountry id="89E173CA514B966E5134FA5CFD7BFA4C" box="[560,684,1469,1492]" name="Philippines" pageId="1" pageNumber="586">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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