448 lines
44 KiB
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448 lines
44 KiB
XML
<document id="9F76773D654A4CF65516062735F78259" ID-DOI="http://dx.doi.org/10.3897/zookeys.689.12021" ID-GBIF-Dataset="907713bd-c7f2-4f8e-a75e-b7f9c958865f" ID-PMC="PMC5672588" ID-Pensoft-Pub="1313-2970-689-1" ID-PubMed="29200920" ID-ZooBank="2F40BF4AD35F4CC697D5976EC201E652" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-689-1" ModsDocOrigin="ZooKeys 689" ModsDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" checkinTime="1502744367222" checkinUser="pensoft" docAuthor="Lackner, Tomas & Leschen, Richard A. B." docDate="2017" docId="6E4C24EA984D9B49E4DA9BE6201B0934" docLanguage="en" docName="ZooKeys 689: 1-263" docOrigin="ZooKeys 689" docSource="http://dx.doi.org/10.3897/zookeys.689.12021" docTitle="Saprinus Erichson 1834" docType="treatment" docVersion="6" lastPageNumber="70" masterDocId="D302FFB6FFA66345FFBAFF89311EFFDE" masterDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" masterLastPageNumber="263" masterPageNumber="1" pageNumber="69" updateTime="1701316739810" updateUser="plazi">
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<mods:title id="FF0AD11A297B2BD686D351166A19C8A2">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:namePart id="290CD937E87E8ED9D43459C18459FB50">Lackner, Tomas</mods:namePart>
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<mods:namePart id="CCCFFACC8825058DAB0AB6EC8EB72E5D">Leschen, Richard A. B.</mods:namePart>
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<mods:date id="DDA194A70F04955F5829C4D0A500FEA8">2017</mods:date>
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<treatment id="6E4C24EA984D9B49E4DA9BE6201B0934" ID-GBIF-Taxon="132648671" LSID="urn:lsid:plazi:treatment:6E4C24EA984D9B49E4DA9BE6201B0934" httpUri="http://treatment.plazi.org/id/6E4C24EA984D9B49E4DA9BE6201B0934" lastPageId="72" lastPageNumber="70" pageId="68" pageNumber="69">
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<subSubSection id="9A203AF511E1EE7B167779A34FC85924" pageId="68" pageNumber="69" type="nomenclature">
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<paragraph id="0AB372B7367E733F4D2A0D46DBBB8496" pageId="68" pageNumber="69">
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<taxonomicName id="A81018BAA9C312B18493CC4368D5E356" ID-CoL="7CBZ" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
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<pageBreakToken id="0883193EDC8D202C7DDF74BEC2CBE69F" pageId="68" pageNumber="69" start="start">Saprinus</pageBreakToken>
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Erichson, 1834
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</taxonomicName>
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Figs 301, 302-310, 311-319, 320, 321-329, 330-336, 337, 338-343, 344-352, 353, 354-362, 363-371, 372, 373-381, 382-388, 389, 390-398, 399-407, 408, 409-414, 415-421, 422, 423-431, 432-438, 439, 440-448, 449-457, 458, 459-467, 468-476, 477, 478-486, 487-493, 494, 495-503, 504-510, 511, 512-520, 521-527, 528, 529-537, 538-546, 547, 548-556, 557-564, 565, 566-577, 578-584, 585, 586-594, 595-602, 603, 604-611, 612-618, 753, 755, 758, 759, 760, 761, 762, 763, 764, 765
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</paragraph>
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</subSubSection>
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<subSubSection id="F34CC730D0130FF3D2EF0382D221FB41" pageId="68" pageNumber="69" type="reference_group">
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<paragraph id="AA4F15A1A9E7FDE81797F73DAEC9D20B" pageId="68" pageNumber="69">
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<taxonomicName id="88F1D97215057C44797FE600EF43D8C2" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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Erichson, 1834: 172. Type species
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<taxonomicName id="20B5A038C00CA552AB4760A7161A7F74" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister nitidulus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="nitidulus">Hister nitidulus</taxonomicName>
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Fabricius, 1801 (=
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<taxonomicName id="70CA2360235C65D8D61D1232D84378FD" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus semistriatus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="semistriatus">Saprinus semistriatus</taxonomicName>
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(L.G. Scriba, 1790)), designated by Westwood (1838): 22.
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</paragraph>
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</subSubSection>
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<subSubSection id="5528EE8EB4987DAE87A1077217E27EF4" pageId="68" pageNumber="69" type="diagnosis">
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<paragraph id="8AC2705A49895B4E2440A19152873B23" pageId="68" pageNumber="69">Diagnosis.</paragraph>
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<paragraph id="0C2BD3AEA248D04A6B8F0D4000D579B3" pageId="68" pageNumber="69">
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Some members of
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<taxonomicName id="DCB866B9805FC1AE7A17215BA71EAC91" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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are the largest of the Australian saprinines with specimens of S. (S.)
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<taxonomicName id="D3EDB54990FD44E9571B09A47D1D76D0" lsidName="viridanus" pageId="68" pageNumber="69" rank="species" species="viridanus">viridanus</taxonomicName>
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measuring up to 6.70 mm (PEL). The genus is also the most species-rich; it contains 18 species, including 4 newly described and 2 introduced species. All Australopacific species (with the exception of
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<taxonomicName id="5ED1316494A1A08790EC6552A2331666" lsidName="S. detritus" pageId="68" pageNumber="69" rank="species" species="detritus">S. detritus</taxonomicName>
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,
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<taxonomicName id="803FD87CFC8A52FEE92806B3D578DE21" lsidName="S. pseudodetritus" pageId="68" pageNumber="69" rank="species" species="pseudodetritus">S. pseudodetritus</taxonomicName>
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, and
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<taxonomicName id="2ABBD74277AC05127E46AB52BB32D14F" lsidName="S. chathamensis" pageId="68" pageNumber="69" rank="species" species="chathamensis">S. chathamensis</taxonomicName>
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from New Zealand/Chatham Islands, and S.
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<taxonomicName id="BBA24B6980D4078F44EA1C604ABDEE46" lsidName="nitiduloides" pageId="68" pageNumber="69" rank="species" species="nitiduloides">nitiduloides</taxonomicName>
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Fairmaire, 1883 from New Britain/Solomon Islands and
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<taxonomicName id="ACCAA556E16867BDD6E69936BAB32C51" lsidName="S. artensis" pageId="68" pageNumber="69" rank="species" species="artensis">S. artensis</taxonomicName>
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Marseul, 1862 from New Caledonia that are dark brown) are characterized by strong metallic luster on the dorsum of their bodies, especially on elytra, and absence of red, orange or yellow elytral patches (often present in Palaearctic, Oriental and African species). Likewise, all have complete frontal stria (with the exception of
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<taxonomicName id="C2CE19BB3616EC71065DCADD62782C8E" lsidName="S. grandiclava" pageId="68" pageNumber="69" rank="species" species="grandiclava">S. grandiclava</taxonomicName>
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from New Guinea and two newly described species from the Chatham Islands). The frontal stria is usually widely interrupted in the Palaearctic or African species and is used to diagnose the Palaearctic members of
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<taxonomicName id="FDAFC3F849DADC22D20835EC0DCC3C4D" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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. One species,
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<taxonomicName id="E0CE5EB9D0DC0ED8473DB3450D1AE658" lsidName="S. viridipennis" pageId="68" pageNumber="69" rank="species" species="viridipennis">S. viridipennis</taxonomicName>
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(Australia), lacks carinal prosternal striae and two species,
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<taxonomicName id="FBE2E5E8BB49622B5CC5E929FBC8EA26" lsidName="S. grandiclava" pageId="68" pageNumber="69" rank="species" species="grandiclava">S. grandiclava</taxonomicName>
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(New Guinea) and
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<taxonomicName id="7CB3A4C4F80C4C92D62BE0ABDBA34AB2" lsidName="S. viridanus" pageId="68" pageNumber="69" rank="species" species="viridanus">S. viridanus</taxonomicName>
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(Australia), share unusually large, circular antennal clubs.
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</paragraph>
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<caption id="FA62EBC843AAC5369EE283308C655EAA" pageId="68" pageNumber="69">
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<paragraph id="3BD33124F8E026244175DFABC1265236" pageId="68" pageNumber="69">
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Figure 301.
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<taxonomicName id="CFD845C28EB936652985FC5ED0E17130" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) amethystinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="amethystinus" subGenus="Saprinus">Saprinus (Saprinus) amethystinus</taxonomicName>
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Lewis, 1900 habitus, dorsal view.
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</paragraph>
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</caption>
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<paragraph id="429D9B5ADDBFCE6FF6496905769ADFB3" pageId="68" pageNumber="69">
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An aedeagal peculiarity among Australopacific
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<taxonomicName id="66F2BDD32261A37E07C41342A93162D8" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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is the presence of well-separated parameres (see also below; e.g. Fig. 475) in the Australian endemic species
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<taxonomicName id="0E7CCB91E68E4B7E2B9CB3EB72140605" lsidName="S. amethystinus" pageId="68" pageNumber="69" rank="species" species="amethystinus">S. amethystinus</taxonomicName>
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Lewis, 1900,
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<taxonomicName id="F2D504E0C707DFF3592CBB2C58110FC7" lsidName="S. australis" pageId="68" pageNumber="69" rank="species" species="australis">S. australis</taxonomicName>
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(Boisduval, 1835),
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<taxonomicName id="BAC69955EF5AF7C12314697B017DBE11" lsidName="S. laetus" pageId="68" pageNumber="69" rank="species" species="laetus">S. laetus</taxonomicName>
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Erichson, 1834, and
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<taxonomicName id="62A6BC2F6810F8101B5ED47949D4439F" lsidName="S. tyrrhenus" pageId="68" pageNumber="69" rank="species" species="tyrrhenus">S. tyrrhenus</taxonomicName>
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Blackburn, 1903. The three New Zealand endemics, as well as
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<taxonomicName id="48B71D95752F4A16ABBE9E6726B2B00F" lsidName="S. grandiclava" pageId="68" pageNumber="69" rank="species" species="grandiclava">S. grandiclava</taxonomicName>
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Kanaar, 1989 (endemic to New Guinea), have their parameres separated in the apical half and are more approximate than in the Australian species (Fig. 456).
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</paragraph>
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</subSubSection>
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<subSubSection id="938C3BA287588D95A99CD5C249ED5340" pageId="68" pageNumber="69" type="biology_ecology">
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<paragraph id="F54F75AD32135EB8B48255174792FE34" pageId="68" pageNumber="69">Biology.</paragraph>
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<paragraph id="DFF51286BED31AE27DBAD8038D9F00D0" pageId="68" pageNumber="69">
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Species of
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<taxonomicName id="793E8BDFD5EB45FE9F55D5BC092CCBDE" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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are typical volant predators with a preference for open grasslands and xerothermic habitats where they prey upon fly larvae found in decomposing organic matter, such as mammalian dung or carcasses (
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<bibRefCitation id="6676C72AFC8ECB165733AD919BF50278" pageId="68" pageNumber="69">Lackner 2010</bibRefCitation>
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). Their large and well-developed sensory patches on the ventral side of their antennal clubs probably enable them to locate the carrion from a long distance (but these are lacking in
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<taxonomicName id="5265E0163D33952F15CFEA9B860DB409" lsidName="S. grandiclava" pageId="68" pageNumber="69" rank="species" species="grandiclava">S. grandiclava</taxonomicName>
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). The natural history of several rare species (
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<taxonomicName id="A3CFA9973B93D6E36943BCFF51C01F27" lsidName="S. grandiclava" pageId="68" pageNumber="69" rank="species" species="grandiclava">S. grandiclava</taxonomicName>
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,
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<taxonomicName id="89FFFDD728B62FB9F456C70607355BD2" lsidName="S. amethystinus" pageId="68" pageNumber="69" rank="species" species="amethystinus">S. amethystinus</taxonomicName>
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and
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<taxonomicName id="316F7F7401C262E89C6B565D96C3ACF3" lsidName="S. tyrrhenus" pageId="68" pageNumber="69" rank="species" species="tyrrhenus">S. tyrrhenus</taxonomicName>
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) is completely unknown.
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<taxonomicName id="46968E9F659781930A47DE2C3E59ECFB" lsidName="S. rarus" pageId="68" pageNumber="69" rank="species" species="rarus">S. rarus</taxonomicName>
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sp. n. was collected in the nest of the arboreal Tree termite (
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<taxonomicName id="88B463EE9C0646C119EA9BE969324E40" class="Insecta" family="Termitidae" genus="Nasutitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nasutitermes walkeri" order="Blattodea" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="walkeri">Nasutitermes walkeri</taxonomicName>
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(Hill, 1942)) and is presumed to be a specialised termitophile.
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</paragraph>
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</subSubSection>
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<subSubSection id="EA5FF887C31632FC4CBAD213F209AB85" pageId="68" pageNumber="69" type="distribution">
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<paragraph id="DD5F73F8A2A8EE2712B6D2E48234680D" pageId="68" pageNumber="69">Distribution.</paragraph>
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<paragraph id="E337D0BD70CA08EC2993F707A89146D8" pageId="68" pageNumber="69">
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There are eighteen species of Australopacific
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<taxonomicName id="EAD2DEBB7D4ABA8D48E41BCA82D71B99" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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: nine native and two introduced (
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<taxonomicName id="073C442D06BDCDCBE963A567B83FD4C6" lsidName="S. chalcites" pageId="68" pageNumber="69" rank="species" species="chalcites">S. chalcites</taxonomicName>
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and
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<taxonomicName id="092D238137A24FC70E85A25DC2A5C901" lsidName="S. cupreus" pageId="68" pageNumber="69" rank="species" species="cupreus">S. cupreus</taxonomicName>
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) are known from Australia; three species of
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<taxonomicName id="33AA9166AA352E615919E570A6AEADAB" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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are present in New Zealand/Chatham Islands; New Caledonia has one endemic and one non-endemic species. New Guinea has a single endemic species, with two other, non-endemic congeners. We describe one endemic species from the Kiribati atoll (Figs 753, 755, 758-765).
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</paragraph>
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</subSubSection>
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<subSubSection id="F8A2A4A40AD601E656790518D01F6801" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="remarks">
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<paragraph id="A4D4BA9F2938207E97072D51909AED8A" pageId="68" pageNumber="69">Remarks.</paragraph>
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<paragraph id="DE8BF00C50B6B771BB7C5FE63AD212D6" pageId="68" pageNumber="69">
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Australopacific species of the genus
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<taxonomicName id="E054676FC6B86703E5BA326284495A19" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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differ from the similar-looking species
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<taxonomicName id="CB6EB1C0D9C3D3F24EC7535FD8932EB8" class="Insecta" family="Histeridae" genus="Notosaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notosaprinus irinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="irinus">Notosaprinus irinus</taxonomicName>
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(Marseul, 1862) by the presence of lateral prosternal striae (usually both carinal and lateral prosternal striae are present). See discussion of
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<taxonomicName id="2AC46946561AC34C4F6340CD8613DFD0" class="Insecta" family="Histeridae" genus="Notosaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notosaprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Notosaprinus</taxonomicName>
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for more details. The genus
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<taxonomicName id="F125A98282A56B25461E0C385CB6376F" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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is the most species-rich genus of the subfamily (see e.g.
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<bibRefCitation id="D082E152E2F1C05E9FEAE946D582A63F" pageId="68" pageNumber="69">Lackner 2010</bibRefCitation>
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or
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<bibRefCitation id="4F2A5998ED12793091A294B188FEB0B5" pageId="68" pageNumber="69">Mazur 2011</bibRefCitation>
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|
for details) with the bulk of its representatives occurring in the Palaearctic and Afrotropical Regions. Although moderately diverse also in the Oriental (Indo-Malayan) and Nearctic Regions,
|
|
<taxonomicName id="4820F42C9EC40D4B1BDE9CCEB5ADB12C" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
of the Australopacific Region with 16 native species are rather species-poor when compared to their counterparts from the other regions. Only the Neotropical Region, with only 7 known species has fewer than Australopacific Region.
|
|
</paragraph>
|
|
<paragraph id="00A609C8D9B67E3957F8A69D7B439C92" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
|
|
Based on the structure of the aedeagus there are two main groups of Australopacific
|
|
<taxonomicName id="A3FEC998448E1E6EED3A3989D7F90DF5" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
: in each group there are eight species with widely separated parameres and with
|
|
<pageBreakToken id="FF43689AA35FA4F9C693B2D064B2B519" pageId="69" pageNumber="70" start="start">fused</pageBreakToken>
|
|
parameres. Regarding the group of species with fused parameres, the following species are morphologically very similar:
|
|
<taxonomicName id="A32F0D854C69DA17BDB918B086A7C7F9" lsidName="S. cyaneus" pageId="69" pageNumber="70" rank="subSpecies" species="cyaneus" subSpecies="cyaneus">S. cyaneus cyaneus</taxonomicName>
|
|
,
|
|
<taxonomicName id="FA6E59A4D23B72EE11B614E024BBCF2E" lsidName="S. artensis" pageId="69" pageNumber="70" rank="species" species="artensis">S. artensis</taxonomicName>
|
|
,
|
|
<taxonomicName id="0730E1F9F058255356E74BFCD5B31ECA" lsidName="S. nitiduloides" pageId="69" pageNumber="70" rank="species" species="nitiduloides">S. nitiduloides</taxonomicName>
|
|
and
|
|
<taxonomicName id="767E2C7AD5C48DD574916C0D696DFBBE" lsidName="S. pacificus" pageId="69" pageNumber="70" rank="species" species="pacificus">S. pacificus</taxonomicName>
|
|
(compare Figs 335, 420, 493 and 510). Based on their overall similarities, including the structure of male genitalia we believe that they likely share a recent common ancestor that probably reached the region from the north, since a subspecies of
|
|
<taxonomicName id="7EC131A0FA7FB51C68616EEDED8F3638" lsidName="S. cyaneus" pageId="69" pageNumber="70" rank="species" species="cyaneus">S. cyaneus</taxonomicName>
|
|
,
|
|
<taxonomicName id="CF9C480B11E3628C6CDB19F3DD18D50F" lsidName="S. cyaneus" pageId="69" pageNumber="70" rank="subSpecies" species="cyaneus" subSpecies="auricollis">S. cyaneus auricollis</taxonomicName>
|
|
, is distributed also in the Indo-Malayan Region up to Southern Japan. These species can possibly be regarded among the so-called northern invaders, sensu Mitchell and Bouchard (2008). Another species with fused parameres,
|
|
<taxonomicName id="1BFED022BD36DEC7E2D5057FBC211C17" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus splendens" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="splendens">Saprinus splendens</taxonomicName>
|
|
, is widely distributed in the tropical regions of Africa/Arabian peninsula and the Oriental (Indo-Malayan) Region as far east and north as Japan, and its occurrence in the Australopacific Region (Australia and Marianna Islands) is not surprising. The three remaining species with fused parameres (
|
|
<taxonomicName id="515BBF5CF20A197844E7529F82978062" lsidName="S. rarus" pageId="69" pageNumber="70" rank="species" species="rarus">S. rarus</taxonomicName>
|
|
,
|
|
<taxonomicName id="35620DC497A3BC2F2981CAA3632F3BB1" lsidName="S. viridanus" pageId="69" pageNumber="70" rank="species" species="viridanus">S. viridanus</taxonomicName>
|
|
, and
|
|
<taxonomicName id="E9A963F844271F415E3850400CC908AE" lsidName="S. viridipennis" pageId="69" pageNumber="70" rank="species" species="viridipennis">S. viridipennis</taxonomicName>
|
|
) are Australian endemics.
|
|
<taxonomicName id="363FE262E964C4F321DCD77315101204" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus rarus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="rarus">Saprinus rarus</taxonomicName>
|
|
has a uniquely triangulate dilated and thickened antennal scape (Fig. 529), strongly dilated tibiae (Fig. 536) and elongate body shape (Fig. 528), features that are probably reflecting its termitophilous lifestyle, and may protect it against termites (thickened antennal club and dilated tibiae protect larger space of body venter when in repose).
|
|
<taxonomicName id="36EE9DA2A9C81C52094B8A632862A011" lsidName="S. viridipennis" pageId="69" pageNumber="70" rank="species" species="viridipennis">S. viridipennis</taxonomicName>
|
|
lacks the carinal prosternal striae entirely (Fig. 609; an autapomorphic feature within
|
|
<taxonomicName id="E53F4F4C1255DA9F158BCB34BB3A0DBD" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
), while the antennal club of
|
|
<taxonomicName id="17F6DF796D1A5B286F2BB02703B46E84" lsidName="S. viridanus" pageId="69" pageNumber="70" rank="species" species="viridanus">S. viridanus</taxonomicName>
|
|
is unusually large and circular, resembling the form present in the New Guinean endemic
|
|
<taxonomicName id="392595900A79830CCA923618AF135F2D" lsidName="S. grandiclava" pageId="69" pageNumber="70" rank="species" species="grandiclava">S. grandiclava</taxonomicName>
|
|
(compare Figs 460 and 586).
|
|
</paragraph>
|
|
<paragraph id="965FEDD9A99A97D52663C84D061CDDD4" pageId="69" pageNumber="70">
|
|
The remaining eight species with (widely) separate parameres (Australian
|
|
<taxonomicName id="82DE7C40D7096CC77716F87BFBAF8142" lsidName="S. amethystinus" pageId="69" pageNumber="70" rank="species" species="amethystinus">S. amethystinus</taxonomicName>
|
|
,
|
|
<taxonomicName id="3ED747A0970AC19CFC053251AE45328E" lsidName="S. australis" pageId="69" pageNumber="70" rank="species" species="australis">S. australis</taxonomicName>
|
|
,
|
|
<taxonomicName id="6BFE8C32D464E9EA690BCB7B16432754" lsidName="S. tyrrhenus" pageId="69" pageNumber="70" rank="species" species="tyrrhenus">S. tyrrhenus</taxonomicName>
|
|
,
|
|
<taxonomicName id="FD53116C3F84644648F367884580E935" lsidName="S. laetus" pageId="69" pageNumber="70" rank="species" species="laetus">S. laetus</taxonomicName>
|
|
; New Guinean
|
|
<taxonomicName id="4B0FD017F18A9D81ACC69381F64ECA30" lsidName="S. grandiclava" pageId="69" pageNumber="70" rank="species" species="grandiclava">S. grandiclava</taxonomicName>
|
|
and New Zealand
|
|
<taxonomicName id="8ECA32309A66AB3358091AD595073E8F" lsidName="S. detritus" pageId="69" pageNumber="70" rank="species" species="detritus">S. detritus</taxonomicName>
|
|
,
|
|
<taxonomicName id="7F5F023186EAC7F4C131142A9E1A090B" lsidName="S. pseudodetritus" pageId="69" pageNumber="70" rank="species" species="pseudodetritus">S. pseudodetritus</taxonomicName>
|
|
and
|
|
<taxonomicName id="3798ACC259EC64327169A9108BA16C03" lsidName="S. chathamensis" pageId="69" pageNumber="70" rank="species" species="chathamensis">S. chathamensis</taxonomicName>
|
|
) are unique among the 157 described species of
|
|
<taxonomicName id="D29CB7C79364DEAF8E98DB99CD74D2E3" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
(sensu
|
|
<bibRefCitation id="2E60D03C00080BA7CC0E19E2565309C7" pageId="69" pageNumber="70">Mazur 2011</bibRefCitation>
|
|
): no other species of
|
|
<taxonomicName id="691F91D9409537A19AA2CCD386CAA6AA" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
have this character. In the published phylogenetic analysis of the senior author (
|
|
<bibRefCitation id="BB7B3077238A8AE2DB452D062883F891" author="Lackner, T" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="216" pageNumber="217" pagination="521 - 555" title="Phylogeny of the Saprininae subfamily reveals interesting ecological shifts in the history of the subfamily (Coleoptera: Histeridae)." url="http://dx.doi.org/10.1111/zoj.12182" volume="172" year="2014 d">Lackner 2014d</bibRefCitation>
|
|
) only the type species of the genus,
|
|
<taxonomicName id="5A5A1D1E70325178B2F9472B0C524093" lsidName="S. (S.) semistriatus" pageId="69" pageNumber="70" rank="species" species="semistriatus" subGenus="S.">S. (S.) semistriatus</taxonomicName>
|
|
, a taxon with fused parameres, was analysed. As noted above (see remarks section of
|
|
<taxonomicName id="8F66FA1D7E6E3632A861A172682C8863" class="Insecta" family="Histeridae" genus="Notosaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notosaprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Notosaprinus</taxonomicName>
|
|
),
|
|
<taxonomicName id="8A97CA70C5CAF8C51F45C12E167637EE" lsidName="S. (S.) semistriatus" pageId="69" pageNumber="70" rank="species" species="semistriatus" subGenus="S.">S. (S.) semistriatus</taxonomicName>
|
|
was recovered sister to
|
|
<taxonomicName id="456EDB5D68F8014A97DB7BFFEBEAAB02" class="Insecta" family="Histeridae" genus="Notosaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notosaprinus irinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="irinus">Notosaprinus irinus</taxonomicName>
|
|
; this clade itself was recovered sister to a larger clade containing North American
|
|
<taxonomicName id="F9488EA1BF491C000CA7E869421DB03B" class="Insecta" family="Histeridae" genus="Xerosaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xerosaprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Xerosaprinus</taxonomicName>
|
|
, African
|
|
<taxonomicName id="6C632868EFCD4030615B99EE0838E9B1" class="Insecta" family="Histeridae" genus="Pilisaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilisaprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Pilisaprinus</taxonomicName>
|
|
and
|
|
<taxonomicName id="BD03BA5694D4D9861C582343DBACD6FD" class="Insecta" family="Histeridae" genus="Paraphilothis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraphilothis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Paraphilothis</taxonomicName>
|
|
, Middle-Asian
|
|
<taxonomicName id="7ADCC79C1DEBE825228029AF6AE75965" class="Insecta" family="Histeridae" genus="Styphrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Styphrus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Styphrus</taxonomicName>
|
|
, Palaearctic
|
|
<taxonomicName id="168A99A1E055398119E098241DD3D1BA" class="Insecta" family="Histeridae" genus="Phaonius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phaonius" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Phaonius</taxonomicName>
|
|
(itself subgenus of
|
|
<taxonomicName id="B7F30232B18FDBABEA4BB5B0BE9CEBE9" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
). None of these relatives of
|
|
<taxonomicName id="EDA51EB8EACBA1A837025CEEAB6E9B8D" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
possess separated parameres. On the other hand, two members of the genus
|
|
<taxonomicName id="B53F9C65557A38C05153D47B113BC984" class="Insecta" family="Histeridae" genus="Microsaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microsaprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Microsaprinus</taxonomicName>
|
|
Kryzhanovskij, 1976 that was recovered in the aforementioned analysis closer to the root, possess separated parameres of the aedeagus (albeit not widely; see Secq and Secq 1995 for figures). Looking at the character value nodes of
|
|
<taxonomicName id="5CCDFD6D80C84ED2243BC866AF78327B" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
near and even more distant relatives the basal aedeagal condition does not appear therefore to be the separated-parameres character state, with the single exception of the two
|
|
<taxonomicName id="BD2C5F2E002149A39909E4A1A4E4F686" class="Insecta" family="Histeridae" genus="Microsaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microsaprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Microsaprinus</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4D6E416E12B40F35F5009077C54859AE" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70" type="key">
|
|
<paragraph id="DE83E45F35618EE8DBFB301E87D8B508" pageId="69" pageNumber="70">
|
|
Key to the Australopacific species of
|
|
<taxonomicName id="8F37CE8133F15F14AFF94AF7F8383586" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
|
|
Erichson, 1834
|
|
</paragraph>
|
|
<paragraph id="54252485150F3CAA4404286E20D59BBB" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
|
|
<table id="B6C09680590E1CC9A085164D1F49A8F5" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
|
|
<tr id="CAE40CD92B8780DA43478E204927C3AA" pageId="69" pageNumber="70">
|
|
<td id="CD0424DAA0AFF6E797705F40F6F58CDD" colspan="1" pageId="69" pageNumber="70" rowspan="1">466</td>
|
|
</tr>
|
|
<tr id="00F9F0C9EA83AEB716E4090897B4BCE4" pageId="70" pageNumber="71">
|
|
<td id="02746F033E0BD89B40BB0257F853B8CB" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<pageBreakToken id="0CCF777B873D450BF63AA0EEA48B715A" pageId="70" pageNumber="71" start="start">458475</pageBreakToken>
|
|
</td>
|
|
<td id="37CD5B3053009DC4052C3DC2259A4D39" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="AFC887A086DB5F2C361859D89F1D17CF" lsidName="S. (S.) laetus" pageId="70" pageNumber="71" rank="species" species="laetus" subGenus="S.">S. (S.) laetus</taxonomicName>
|
|
Erichson, 1834
|
|
</td>
|
|
</tr>
|
|
<tr id="56C80FD352179B925C2903FC88F80964" pageId="70" pageNumber="71">
|
|
<td id="E1CC06C2BFE7D2D45F829C12B9E59B34" colspan="1" pageId="70" pageNumber="71" rowspan="1">511526</td>
|
|
</tr>
|
|
<tr id="745646285BBDEF554CDF2F4A44A759E7" pageId="70" pageNumber="71">
|
|
<td id="7E21977439D20EE2EF462C2A0358453A" colspan="1" pageId="70" pageNumber="71" rowspan="1">511515</td>
|
|
<td id="96D744E0E75E2E8402681AA706651C4F" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="D9BC76147A05CB45D0DDFA9B3DA8FC72" lsidName="S. (S.) pseudodetritus" pageId="70" pageNumber="71" rank="species" species="pseudodetritus" subGenus="S.">S. (S.) pseudodetritus</taxonomicName>
|
|
sp. n.
|
|
</td>
|
|
</tr>
|
|
<tr id="61365E5AD508734349DFADE5BD2D992A" pageId="70" pageNumber="71">
|
|
<td id="9AA9C714FB4EAC2A98CE7F38E833C0B4" colspan="1" pageId="70" pageNumber="71" rowspan="1">372376</td>
|
|
</tr>
|
|
<tr id="60A5411F1A2276C50D8A7DB499868885" pageId="70" pageNumber="71">
|
|
<td id="F164EA5791F251E536B1A79F6662251D" colspan="1" pageId="70" pageNumber="71" rowspan="1">422432-438</td>
|
|
<td id="02384F702E192810EBCD15A5458726ED" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="6FB49AB2CD0F40A6CC9CCBF8A59991FF" lsidName="S. (S.) detritus" pageId="70" pageNumber="71" rank="species" species="detritus" subGenus="S.">S. (S.) detritus</taxonomicName>
|
|
(Fabricius, 1775)
|
|
</td>
|
|
</tr>
|
|
<tr id="059BC9A64852F8239BFF8A50A1DED74D" pageId="70" pageNumber="71">
|
|
<td id="63FDF9C19DEE4E3776D5EAAF697D6A66" colspan="1" pageId="70" pageNumber="71" rowspan="1">372382-388</td>
|
|
<td id="A10AC6DD1851AB6740D396EAB33D9867" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="6861A29F951181DB7F3CD8E165C31F87" lsidName="S. (S.) chathamensis" pageId="70" pageNumber="71" rank="species" species="chathamensis" subGenus="S.">S. (S.) chathamensis</taxonomicName>
|
|
sp. n.
|
|
</td>
|
|
</tr>
|
|
<tr id="25DC5E596418C4DD0FC80616F141CFE9" pageId="70" pageNumber="71">
|
|
<td id="3D20660AA591E0D9BB0D676E2D37FCC0" colspan="1" pageId="70" pageNumber="71" rowspan="1">603</td>
|
|
</tr>
|
|
<tr id="DFF364AED09BE802D5A721B19E7CD4AE" pageId="70" pageNumber="71">
|
|
<td id="6B85240F8E9D802753548DF6CBFE1BDC" colspan="1" pageId="70" pageNumber="71" rowspan="1">609612-618</td>
|
|
<td id="A60882A473EB45344FD347B3BD5100EF" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="CD24CA97AA38269B7469EE1D1B4AE8EB" lsidName="S. (S.) viridipennis" pageId="70" pageNumber="71" rank="species" species="viridipennis" subGenus="S.">S. (S.) viridipennis</taxonomicName>
|
|
Lewis, 1901
|
|
</td>
|
|
</tr>
|
|
<tr id="5351C531FE364CEC853912B3F6B605A2" pageId="70" pageNumber="71">
|
|
<td id="8776EF0126F1B8F6F0238F9315A48FB0" colspan="1" pageId="70" pageNumber="71" rowspan="1">589</td>
|
|
</tr>
|
|
<tr id="82B80E4E9359C044ED5A09738296DCB2" pageId="70" pageNumber="71">
|
|
<td id="1224FE4B7B0752ABDFE095C5DF0D565B" colspan="1" pageId="70" pageNumber="71" rowspan="1">PEL587601</td>
|
|
<td id="07D00E66A990685A305E2B576FFE67A5" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="9EDA946AD2C3A8A6B4B25F5ADCBFDAA1" lsidName="S. (S.) viridanus" pageId="70" pageNumber="71" rank="species" species="viridanus" subGenus="S.">S. (S.) viridanus</taxonomicName>
|
|
Lewis, 1899
|
|
</td>
|
|
</tr>
|
|
<tr id="F0979794871B9FE73011277165FD8A44" pageId="70" pageNumber="71">
|
|
<td id="0CAB1CBF576F4EE362B42556A3D7D79F" colspan="1" pageId="70" pageNumber="71" rowspan="1">PEL303318456</td>
|
|
</tr>
|
|
<tr id="48D0E8214E2A46D4F752EF06D9B66383" pageId="70" pageNumber="71">
|
|
<td id="58D2A750364B0BCC47C3E448F2B4E282" colspan="1" pageId="70" pageNumber="71" rowspan="1">301PEL306311-319</td>
|
|
<td id="573B53EE13C774F6E758A206B499E648" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="2FC799F0E283C5C9DD6D73F0FE0B164F" lsidName="S. (S.) amethystinus" pageId="70" pageNumber="71" rank="species" species="amethystinus" subGenus="S.">S. (S.) amethystinus</taxonomicName>
|
|
Lewis, 1900
|
|
</td>
|
|
</tr>
|
|
<tr id="AB86394CA62A4A2B63D264DAFF7720E2" pageId="70" pageNumber="71">
|
|
<td id="B34A9E778C7DC2492BC03FA63FF06CA6" colspan="1" pageId="70" pageNumber="71" rowspan="1">565PEL571578-584</td>
|
|
<td id="DE9E532E738EB11BC7719D3A8D1ACEE0" colspan="1" pageId="70" pageNumber="71" rowspan="1">
|
|
<taxonomicName id="15B956CCBDFE39C97A705C9AA6389D7D" lsidName="S. (S.) tyrrhenus" pageId="70" pageNumber="71" rank="species" species="tyrrhenus" subGenus="S.">S. (S.) tyrrhenus</taxonomicName>
|
|
Blackburn, 1903
|
|
</td>
|
|
</tr>
|
|
<tr id="FFACC7C26492ED242999F0EABFCA6A16" pageId="71" pageNumber="72">
|
|
<td id="059868A1ED3B534A85C77A7C70122C3A" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<pageBreakToken id="0FF7D362D88EB1A56862FC1D3040FE83" pageId="71" pageNumber="72" start="start">441442</pageBreakToken>
|
|
</td>
|
|
<td id="9DC99C9E28E34D74C2E5FBBB75C5139F" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<taxonomicName id="1B3E334DF60C39FB43DCDCDB1C5C0A61" lsidName="S. (S.) grandiclava" pageId="71" pageNumber="72" rank="species" species="grandiclava" subGenus="S.">S. (S.) grandiclava</taxonomicName>
|
|
Kanaar, 1989
|
|
</td>
|
|
</tr>
|
|
<tr id="09792DEB2D29AB2BB0F7B012E0C07598" pageId="71" pageNumber="72">
|
|
<td id="4D54922F44B6D670CA4D0FFCE32414EB" colspan="1" pageId="71" pageNumber="72" rowspan="1">549</td>
|
|
</tr>
|
|
<tr id="FA5B54BDE04C958A7614E9235D14365F" pageId="71" pageNumber="72">
|
|
<td id="0911A302CA377308DC428E9E382B5127" colspan="1" pageId="71" pageNumber="72" rowspan="1">528536537529</td>
|
|
<td id="E7DBCD7A6DFC9E6F992232D5D7A8599B" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<taxonomicName id="02DF7A592AB237CF613D5190F5F51851" lsidName="S. (S.) rarus" pageId="71" pageNumber="72" rank="species" species="rarus" subGenus="S.">S. (S.) rarus</taxonomicName>
|
|
sp. n.
|
|
</td>
|
|
</tr>
|
|
<tr id="C950482C771452D027C08E606113E5BF" pageId="71" pageNumber="72">
|
|
<td id="7323D425650DCA9D8359E3B714B90A05" colspan="1" pageId="71" pageNumber="72" rowspan="1">389397398390</td>
|
|
</tr>
|
|
<tr id="8953E2817FDB990CCC4D95F9BCA9295F" pageId="71" pageNumber="72">
|
|
<td id="C2C31F216D3CBE0AD76CF8D7F6A62254" colspan="1" pageId="71" pageNumber="72" rowspan="1">PEL390</td>
|
|
</tr>
|
|
<tr id="14EE996E6C5DF821E52CF43E090D73CA" pageId="71" pageNumber="72">
|
|
<td id="0B413C1F6BFE036C019EB400B8AB0FFD" colspan="1" pageId="71" pageNumber="72" rowspan="1">389399-407</td>
|
|
<td id="EFDE0F0C2430BED07E3ADE0B72AC1A38" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<taxonomicName id="E6C0CC0064BD507BC69EF81A1BB6C856" lsidName="S. (S.) cupreus" pageId="71" pageNumber="72" rank="species" species="cupreus" subGenus="S.">S. (S.) cupreus</taxonomicName>
|
|
Erichson, 1834
|
|
</td>
|
|
</tr>
|
|
<tr id="424EFFD7D89D65B6EB871B74FDC0AE3B" pageId="71" pageNumber="72">
|
|
<td id="C462F5F1389CC1A3C7C7A977E780C62A" colspan="1" pageId="71" pageNumber="72" rowspan="1">353363-371</td>
|
|
<td id="48CC900FBCDD6C70DA762489BA11D4F9" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<taxonomicName id="FB59CC16663141292EF30A3966DC3A90" lsidName="S. (S.) chalcites" pageId="71" pageNumber="72" rank="species" species="chalcites" subGenus="S.">S. (S.) chalcites</taxonomicName>
|
|
(Illiger, 1807)
|
|
</td>
|
|
</tr>
|
|
<tr id="6AF97D3A7F4FEC4EB4B628C5ADCEE91C" pageId="71" pageNumber="72">
|
|
<td id="D3F9ECF03A562061D9E7A181DF484180" colspan="1" pageId="71" pageNumber="72" rowspan="1">PEL338</td>
|
|
</tr>
|
|
<tr id="BD2A38017AE05E14F603036B1FEA6C03" pageId="71" pageNumber="72">
|
|
<td id="5D62230E3769794E4051E5C4189547F5" colspan="1" pageId="71" pageNumber="72" rowspan="1">337351</td>
|
|
<td id="A35CAEF4EB02C069C3D5E3209CEE2982" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<taxonomicName id="FF1CA5D935F55CED6A43BF5CD4B33F6F" lsidName="S. (S.) australis" pageId="71" pageNumber="72" rank="species" species="australis" subGenus="S.">S. (S.) australis</taxonomicName>
|
|
(Boisduval, 1835)
|
|
</td>
|
|
</tr>
|
|
<tr id="4461D188CA007EFB46A1741B037262DB" pageId="71" pageNumber="72">
|
|
<td id="1B5C6E80ECFF26362D7C8F8DF7CCAD6A" colspan="1" pageId="71" pageNumber="72" rowspan="1">408</td>
|
|
</tr>
|
|
<tr id="7AADAAEA1FD4DFBA746DF09CE3238B88" pageId="71" pageNumber="72">
|
|
<td id="BD429F79D1556C5233E39C45215C8546" colspan="1" pageId="71" pageNumber="72" rowspan="1">547561</td>
|
|
<td id="588B3B57B9D0CB6CBAC6970C9E48AAD2" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<taxonomicName id="C8642162C27B4A390874BB90DA3C9119" lsidName="S. (S.) splendens" pageId="71" pageNumber="72" rank="species" species="splendens" subGenus="S.">S. (S.) splendens</taxonomicName>
|
|
(Paykull, 1811)
|
|
</td>
|
|
</tr>
|
|
<tr id="F32ED7161F3449CE846F9E95738B05FC" pageId="71" pageNumber="72">
|
|
<td id="1B784532F1B5F56A81AD5A41688CDECD" colspan="1" pageId="71" pageNumber="72" rowspan="1">408420510</td>
|
|
</tr>
|
|
<tr id="F88D663D103B1DADF0C51E7001833196" pageId="71" pageNumber="72">
|
|
<td id="4F8C6C4C759F300D5496A6AE6558E893" colspan="1" pageId="71" pageNumber="72" rowspan="1">408415-421</td>
|
|
<td id="5D582F70AC006258F460C405E966FB63" colspan="1" pageId="71" pageNumber="72" rowspan="1">
|
|
<taxonomicName id="D6063412FBEED93ED5BDDD3212648B90" lsidName="S. (S.) cyaneus" pageId="71" pageNumber="72" rank="subSpecies" species="cyaneus" subGenus="S." subSpecies="cyaneus">S. (S.) cyaneus cyaneus</taxonomicName>
|
|
(Fabricius, 1775)
|
|
</td>
|
|
</tr>
|
|
<tr id="66E3749C7F2D9E446EC7CFAD53A69FB3" pageId="72" pageNumber="73">
|
|
<td id="9198E13B757605CC3DE6180ABBD1E0B1" colspan="1" pageId="72" pageNumber="73" rowspan="1">
|
|
<pageBreakToken id="3505F93E916FBE4090D79C112A70F455" pageId="72" pageNumber="73" start="start">320</pageBreakToken>
|
|
</td>
|
|
</tr>
|
|
<tr id="C3027C9478971FC3DCE408C98151FC44" pageId="72" pageNumber="73">
|
|
<td id="3A03BF70E75331F578BC744E2FD0F53C" colspan="1" pageId="72" pageNumber="73" rowspan="1">320330-336</td>
|
|
<td id="876C51CE89835E1A6E637C39A21C2C43" colspan="1" pageId="72" pageNumber="73" rowspan="1">
|
|
<taxonomicName id="694DCEF864BD4591D4F5088652A49C9A" lsidName="S. (S.) artensis" pageId="72" pageNumber="73" rank="species" species="artensis" subGenus="S.">S. (S.) artensis</taxonomicName>
|
|
Marseul, 1862
|
|
</td>
|
|
</tr>
|
|
<tr id="02FD9C5FFDFA423F64B4C79741F6F354" pageId="72" pageNumber="73">
|
|
<td id="F2FB27E37A84F067E714944F4C290FAF" colspan="1" pageId="72" pageNumber="73" rowspan="1">477</td>
|
|
</tr>
|
|
<tr id="195CA32254E19A2C7AF2E7B377C84D23" pageId="72" pageNumber="73">
|
|
<td id="77DD2EBAA0C546374E31A15F30A5D35F" colspan="1" pageId="72" pageNumber="73" rowspan="1">487-493</td>
|
|
<td id="D0BECB65107B385C7E38DD32C13D5B6F" colspan="1" pageId="72" pageNumber="73" rowspan="1">
|
|
<taxonomicName id="4FFC26A49ADFF37125C1B3DAD4714195" lsidName="S. (S.) nitiduloides" pageId="72" pageNumber="73" rank="species" species="nitiduloides" subGenus="S.">S. (S.) nitiduloides</taxonomicName>
|
|
Fairmaire, 1883
|
|
</td>
|
|
</tr>
|
|
<tr id="AFBE25BB7E2BA7ED6FEE3EE1C05C9FBB" pageId="72" pageNumber="73">
|
|
<td id="B1BB6CAF375AB10D09117DD294C64597" colspan="1" pageId="72" pageNumber="73" rowspan="1">504-510</td>
|
|
<td id="714761263CD6D938D3633A940B2F67D9" colspan="1" pageId="72" pageNumber="73" rowspan="1">
|
|
<taxonomicName id="5418642DBD1F4BDEB658B849FA2CC300" lsidName="S. (S.) pacificus" pageId="72" pageNumber="73" rank="species" species="pacificus" subGenus="S.">S. (S.) pacificus</taxonomicName>
|
|
sp. n.
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |