204 lines
20 KiB
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204 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.263.4158" ID-GBIF-Dataset="5fe6c912-0997-4d09-800d-101a2a9c26e8" ID-PMC="PMC3591764" ID-Pensoft-Pub="1313-2970-263-59" ID-PubMed="23653517" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-263-59" ModsDocOrigin="ZooKeys 263" ModsDocTitle="Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species" checkinTime="1451247727841" checkinUser="pensoft" docAuthor="Taylor, Christopher K." docDate="2013" docId="5A73C065345DAE7530261EF901601F17" docLanguage="en" docName="ZooKeys 263: 59-73" docOrigin="ZooKeys 263" docSource="http://dx.doi.org/10.3897/zookeys.263.4158" docTitle="Pantopsalis listeri White 1849" docType="treatment" docVersion="3" lastPageNumber="66" masterDocId="776AFFECFFB1FFC40947FF959858FFB7" masterDocTitle="Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species" masterLastPageNumber="73" masterPageNumber="59" pageNumber="66" updateTime="1668155226950" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Taylor, Christopher K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>263</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>59</mods:start>
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<mods:end>73</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.263.4158</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.263.4158</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-263-59</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040053" LSID="urn:lsid:plazi:treatment:5A73C065345DAE7530261EF901601F17" httpUri="http://treatment.plazi.org/id/5A73C065345DAE7530261EF901601F17" lastPageId="8" lastPageNumber="66" pageId="7" pageNumber="66">
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<subSubSection pageId="7" pageNumber="66" type="nomenclature">
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<paragraph pageId="7" pageNumber="66">
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<taxonomicName LSID="http://species-id.net/wiki/Pantopsalis_listeri" authority="White, 1849" authorityName="White" authorityYear="1849" class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">
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<pageBreakToken pageId="7" pageNumber="66" start="start">Pantopsalis</pageBreakToken>
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listeri (White, 1849)
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</taxonomicName>
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Figure 3
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="66" type="reference_group">
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<paragraph pageId="7" pageNumber="66">
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<taxonomicName class="Arachnida" family="Phalangiidae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangium listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Phalangium listeri</taxonomicName>
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<bibRefCitation pageId="7" pageNumber="66">White 1849</bibRefCitation>
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: 6 (reprinted 1850: 52).
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</paragraph>
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<paragraph pageId="7" pageNumber="66">
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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(
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<bibRefCitation pageId="7" pageNumber="66">White 1849</bibRefCitation>
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):
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<bibRefCitation pageId="7" pageNumber="66">Simon 1879</bibRefCitation>
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: 73,
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
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: 61 (as nomen dubium; further citations provided by
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="7" pageNumber="66">
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis luna" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="luna">Megalopsalis luna</taxonomicName>
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<bibRefCitation author="Forster, RR" journalOrPublisher="Records of the Dominion Museum" pageId="12" pageNumber="71" pagination="183 - 192" title="The genus Megalopsalis Roewer in New Zealand with keys to the New Zealand genera of Opiliones." volume="1" year="1944">Forster 1944</bibRefCitation>
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: 190, pl. 66 Figs 1-3 syn. n.
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</paragraph>
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<paragraph pageId="7" pageNumber="66">
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis luna" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="luna">Pantopsalis luna</taxonomicName>
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(
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<bibRefCitation author="Forster, RR" journalOrPublisher="Records of the Dominion Museum" pageId="12" pageNumber="71" pagination="183 - 192" title="The genus Megalopsalis Roewer in New Zealand with keys to the New Zealand genera of Opiliones." volume="1" year="1944">Forster 1944</bibRefCitation>
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):
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
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: 70-71, Fig. 9.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="66" type="neotype">
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<paragraph pageId="7" pageNumber="66">Neotype.</paragraph>
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<paragraph pageId="7" pageNumber="66">1 ♂, 'Ile du Milieu, Filhol, 1875-75' (MNHP no. 134).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="66" type="other specimens examined">
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<paragraph pageId="7" pageNumber="66">Other specimens examined.</paragraph>
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<paragraph pageId="7" pageNumber="66">1 ♂, same data as neotype; 3 ♂, New Zealand, WD. Waiho Gorge, Sth Westland, 21 July 1927 (MONZ); photographs of live males provided by Simon Pollard (Canterbury Museum, Christchurch).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="66" type="description">
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<paragraph pageId="7" pageNumber="66">Description.</paragraph>
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<paragraph pageId="7" pageNumber="66">
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As described by
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<bibRefCitation pageId="7" pageNumber="66">Simon (1879)</bibRefCitation>
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, with the following additions: Dorsum of opisthosoma with pale silvery, narrow, transverse stripes on posterior margins of segments (Fig. 3b). Segment II of chelicera inflated in neotype, slender in remaining specimens. Glans of penis without dorsal keel (
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
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: Fig. 9).
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</paragraph>
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<caption pageId="7" pageNumber="66">
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<paragraph pageId="7" pageNumber="66">
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Figure 3.
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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. A Neotype, lateral view B live specimen, photographed by Simon Pollard.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66" type="comments">
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<paragraph pageId="7" pageNumber="66">Comments.</paragraph>
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<paragraph pageId="7" pageNumber="66">
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The original type specimen(s) of
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<taxonomicName class="Arachnida" family="Phalangiidae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangium listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Phalangium listeri</taxonomicName>
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are lost; they have not been located in the collection of the Museum of Natural History, London (J. Beccaloni, pers. com.) or of the
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<normalizedToken originalValue="Muséum">Museum</normalizedToken>
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national
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<normalizedToken originalValue="d’Histoire">d'Histoire</normalizedToken>
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naturelle, Paris (M. Judson, pers. com.) It was therefore treated as a nomen dubium by
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor (2004)</bibRefCitation>
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. A redescription of
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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was provided by
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<bibRefCitation pageId="7" pageNumber="66">Simon (1879)</bibRefCitation>
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, who made it the type species of his new genus
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="7" pageNumber="66">
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Since the publication of
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor (2004)</bibRefCitation>
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, I have had the opportunity to examine two of the specimens used by
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<bibRefCitation pageId="7" pageNumber="66">Simon (1879)</bibRefCitation>
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in his redescription of this species. Their state of preservation is not ideal (they appear to have been subject to desiccation at some point in the past) and the genitalia have become distorted. Nevertheless, I was able to confirm the absence of a dorsal keel on the glans, demonstrating that
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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could not be conspecific with
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis albipalpis" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="albipalpis">Pantopsalis albipalpis</taxonomicName>
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,
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis coronata" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="coronata">Pantopsalis coronata</taxonomicName>
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or
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis phocator" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="phocator">Pantopsalis phocator</taxonomicName>
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among other South Island
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
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species. External characters (described by
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<bibRefCitation pageId="7" pageNumber="66">Simon 1879</bibRefCitation>
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), such as the unarmed ocularium with denticles restricted to the anterior propeltidium on the dorsal prosomal plate, are also consistent with male specimens referred to
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis luna" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="luna">Pantopsalis luna</taxonomicName>
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(
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<bibRefCitation author="Forster, RR" journalOrPublisher="Records of the Dominion Museum" pageId="12" pageNumber="71" pagination="183 - 192" title="The genus Megalopsalis Roewer in New Zealand with keys to the New Zealand genera of Opiliones." volume="1" year="1944">Forster 1944</bibRefCitation>
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) by
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor (2004)</bibRefCitation>
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(pers. obs.), and it is my judgement that that species is a junior synonym of
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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sensu
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<bibRefCitation pageId="7" pageNumber="66">Simon (1879)</bibRefCitation>
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. The opisthosoma has collapsed in both MNHP specimens, so
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<normalizedToken originalValue="Simon’s">Simon's</normalizedToken>
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(1879) failure to note the transverse striping present in this species may be an artefact of preservation and does not oppose the synonymy.
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</paragraph>
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<paragraph lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
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<normalizedToken originalValue="White’s">White's</normalizedToken>
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(1849) original description of
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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does not provide a more detailed type locality than 'New
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<normalizedToken originalValue="Zealand’">Zealand'</normalizedToken>
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, but
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<bibRefCitation author="Judson, MLI" journalOrPublisher="Occasional Papers on Systematic Entomology" pageId="13" pageNumber="72" pagination="1 - 54" title="Catalogue of the pseudoscorpion types (Arachnida: Chelonethi) in the Natural History Museum, London." volume="11" year="1997">Judson (1997)</bibRefCitation>
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suggested the Bay of Islands, North Island, as the likely type locality for
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<taxonomicName class="Arachnida" family="Cheliferidae" genus="Chelifer" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chelifer pallipes" order="Pseudoscorpiones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="pallipes">Chelifer pallipes</taxonomicName>
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<bibRefCitation pageId="7" pageNumber="66">White 1849</bibRefCitation>
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(now
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<taxonomicName class="Arachnida" family="Cheliferidae" genus="Philomaoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philomaoria pallipes" order="Pseudoscorpiones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="pallipes">Philomaoria pallipes</taxonomicName>
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), described in the same paper. If this was also the case for
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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, then
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<normalizedToken originalValue="Simon’s">Simon's</normalizedToken>
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(1879) specimens would be unlikely to represent the same species as
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<normalizedToken originalValue="White’s">White's</normalizedToken>
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<pageBreakToken pageId="8" pageNumber="67" start="start">original</pageBreakToken>
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type(s).
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<bibRefCitation pageId="8" pageNumber="67">Simon (1879)</bibRefCitation>
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did not explicitly indicate how he identified his specimens as
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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; as no other New Zealand enantiobunine had yet been described, Simon was probably simply unaware that more than one phalangioid species with enlarged chelicerae existed there. Nevertheless, one of the MNHP specimens (Fig. 3a) is here designated as neotype of
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<taxonomicName class="Arachnida" family="Phalangiidae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangium listeri" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="listeri">Phalangium listeri</taxonomicName>
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.
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<normalizedToken originalValue="White’s">White's</normalizedToken>
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original description was exceedingly rudimentary, giving basic characters of the chelicerae only, and inadequate for determining which of the genera
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
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,
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
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or
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus">Mangatangi</taxonomicName>
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was being examined.
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<normalizedToken originalValue="Simon’s">Simon's</normalizedToken>
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more detailed redescription was at least implicitly used as the basis for identification of
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
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by all subsequent authors (
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<bibRefCitation author="Pocock, RI" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="13" pageNumber="72" pagination="392 - 413" title="On some new harvest-spiders of the order Opiliones from the southern continents." volume="1902" year="1903 a">Pocock 1903a</bibRefCitation>
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, b,
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<bibRefCitation author="Hogg, HR" journalOrPublisher="Transactions and Proceedings of the New Zealand Institute, new series" pageId="12" pageNumber="71" pagination="273 - 283" title="Some New Zealand and Tasmanian Arachnidae." volume="42" year="1910">Hogg 1910</bibRefCitation>
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, 1920,
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<bibRefCitation pageId="8" pageNumber="67">Roewer 1923</bibRefCitation>
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,
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<bibRefCitation author="Taylor, CK" journalOrPublisher="Tuhinga" pageId="13" pageNumber="72" pagination="53 - 76" title="New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae) - the genus Pantopsalis." volume="15" year="2004">Taylor 2004</bibRefCitation>
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||
). Nomenclatural stability is best served by fixing
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<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis listeri" order="Opiliones" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="listeri">Pantopsalis listeri</taxonomicName>
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's identity as the species examined by Simon.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |