474 lines
61 KiB
XML
474 lines
61 KiB
XML
<document id="220576BDA0DCAE9EA8428D092FF7A994" ID-DOI="10.11646/zootaxa.4032.3.4" ID-GBIF-Dataset="dfc837d7-f020-4fa6-b4a4-e0fcbbb353ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="238343" ID-ZooBank="EC2FE508-40E1-4464-868D-E2ED0BC56960" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461293766316" checkinUser="plazi" docAuthor="Zeiri, Asma, Nel, Andre & Garrouste, Romain" docDate="2015" docId="0B5287CD4801FFF9FF429984FAA7A085" docLanguage="en" docName="zt04032p296.pdf" docOrigin="Zootaxa 4032 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Urolibellula eocenica Zeiri, Nel & Garrouste, 2015, sp. n." docType="treatment" docVersion="7" lastPageNumber="295" masterDocId="F76BFFB54800FFFCFFD59F5FFFE8A10D" masterDocTitle="A new libelluloid family from the Eocene Green River Formation (Colorado, USA) (Odonata, Anisoptera)" masterLastPageNumber="296" masterPageNumber="290" pageNumber="291" updateTime="1698616086538" updateUser="plazi">
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<mods:titleInfo id="989F1562566A6F7ED9CC7707EE490E1D">
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<mods:title id="F5EB8AF3F27EB553CF11FF3208017E69">A new libelluloid family from the Eocene Green River Formation (Colorado, USA) (Odonata, Anisoptera)</mods:title>
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<mods:name id="CC882579B694DC8DD53B137CA9ED43DC" type="personal">
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<mods:namePart id="D54D73BF45152F871308B04BB16B6EF3">Zeiri, Asma</mods:namePart>
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<mods:namePart id="82AE6CFE6873345EDDF2B570F44DBADF">Nel, Andre</mods:namePart>
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<mods:name id="96F1753EF4B2571ECC4076E59D674370" type="personal">
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<mods:roleTerm id="0B27D46F3F3398230C84CB3D89A5CE86">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="546AC66C41D5F0328BD10E41612EBDB9">Garrouste, Romain</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DC06314C05305ED4BDE0D0D3E0CD7C31">text</mods:typeOfResource>
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<mods:title id="135ABBBBF5D779EC8B991C984160FEEC">Zootaxa</mods:title>
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<mods:part id="0964FC4DD96324AE080146F98BB7916E">
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<mods:date id="A21A2CBE205D3065E7EADFB552C171D9">2015</mods:date>
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<mods:number id="723C88B706E868F9A33B98D1E30EF4D9">4032</mods:number>
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<mods:classification id="179004750B0919D9F06088A7D8898C3A">journal article</mods:classification>
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<mods:identifier id="011AF88A9D71B5252EC2CE5F2E3A32BB" type="DOI">10.11646/zootaxa.4032.3.4</mods:identifier>
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<mods:identifier id="402407CFAD9D6C67100067721D612619" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0B5287CD4801FFF9FF429984FAA7A085" ID-DOI="http://doi.org/10.5281/zenodo.6111617" ID-GBIF-Taxon="119642245" ID-Zenodo-Dep="6111617" LSID="urn:lsid:plazi:treatment:0B5287CD4801FFF9FF429984FAA7A085" httpUri="http://treatment.plazi.org/id/0B5287CD4801FFF9FF429984FAA7A085" lastPageId="5" lastPageNumber="295" pageId="1" pageNumber="291">
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<subSubSection id="CBE165504801FFFDFF429984FEF9A61A" pageId="1" pageNumber="291" type="nomenclature">
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<paragraph id="834436DB4801FFFDFF429984FE0CA7FB" blockId="1.[151,484,1755,1815]" box="[151,484,1755,1782]" pageId="1" pageNumber="291">
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<heading id="D80C81B74801FFFDFF429984FE0CA7FB" bold="true" box="[151,484,1755,1782]" fontSize="11" level="1" pageId="1" pageNumber="291" reason="1">
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<emphasis id="B18FEAC94801FFFDFF429984FE0CA7FB" bold="true" box="[151,484,1755,1782]" pageId="1" pageNumber="291">
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<taxonomicName id="44FB4D584801FFFDFF429984FE73A7F8" box="[151,411,1755,1781]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="291" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="B18FEAC94801FFFDFF429984FE73A7F8" bold="true" box="[151,411,1755,1781]" italics="true" pageId="1" pageNumber="291">Urolibellula eocenica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AABC57B24801FFFDFE779983FE0CA7FB" box="[418,484,1756,1782]" pageId="1" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="834436DB4801FFFDFF4299A1FEF9A61A" blockId="1.[151,484,1755,1815]" box="[151,273,1790,1815]" pageId="1" pageNumber="291">
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(
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<figureCitation id="1BC02A5E4801FFFDFF4A99A1FEE1A61A" box="[159,265,1790,1815]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1021,1043]" captionStartId-1="2.[151,250,1454,1476]" captionStartId-2="2.[151,250,2009,2031]" captionTargetBox-0="[264,1322,179,999]" captionTargetBox-1="[282,1305,1101,1426]" captionTargetBox-2="[293,1287,1521,1979]" captionTargetId-0="figure@2.[264,1323,179,1000]" captionTargetId-1="figure@2.[273,1315,1093,1433]" captionTargetId-2="figure@2.[289,1297,1508,1987]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. Urolibellula eocenica sp. n., holotype specimen, photograph of general habitus (scale bar represents 3 mm)." captionText-1="FIGURE 2. Urolibellula eocenica sp. n., holotype specimen, drawing of forewing (scale bar represents 4 mm)." captionText-2="FIGURE 3. Urolibellula eocenica sp. n., holotype specimen, drawing of hindwing (scale bar represents 4 mm)." httpUri-0="https://zenodo.org/record/238344/files/figure.png" httpUri-1="https://zenodo.org/record/238345/files/figure.png" httpUri-2="https://zenodo.org/record/238346/files/figure.png" pageId="1" pageNumber="291">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="CBE165504801FFFDFF429819FE8EA6C6" pageId="1" pageNumber="291" type="materials_examined">
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<paragraph id="834436DB4801FFFDFF429819FE58A68E" blockId="1.[151,1436,1862,2031]" pageId="1" pageNumber="291">
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<emphasis id="B18FEAC94801FFFDFF429819FEEEA652" bold="true" box="[151,262,1862,1887]" pageId="1" pageNumber="291">Material.</emphasis>
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<typeStatus id="5C4088794801FFFDFED89819FE9DA652" box="[269,373,1862,1887]" pageId="1" pageNumber="291" type="holotype">Holotype</typeStatus>
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specimen without number, labelled “Dragonfly, Anisoptera gen.
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<taxonomicName id="44FB4D584801FFFDFB859819FB32A652" box="[1104,1242,1862,1887]" class="Insecta" family="Corduliidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="291" phylum="Arthropoda" rank="family">Corduliidae</taxonomicName>
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”,
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<collectionCode id="E5EAAE1E4801FFFDFB3B9818FAD7A652" LSID="urn:lsid:biocol.org:col:34871" box="[1262,1343,1863,1887]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="291">USNM</collectionCode>
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(United State National Museum).
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</paragraph>
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<paragraph id="834436DB4801FFFDFF1298D1FE8EA6C6" blockId="1.[151,1436,1862,2031]" pageId="1" pageNumber="291">
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<emphasis id="B18FEAC94801FFFDFF1298D1FE78A6AA" bold="true" box="[199,400,1934,1959]" pageId="1" pageNumber="291">Age and outcrop</emphasis>
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. Eocene, Green River Formation, Parachute Creek Member, 43–47 Myrs, Wighton, Garfiel County, Colorado.
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</paragraph>
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</subSubSection>
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<subSubSection id="CBE165504801FFFDFF129889FDE0A6E2" box="[199,520,2006,2031]" pageId="1" pageNumber="291" type="diagnosis">
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<paragraph id="834436DB4801FFFDFF129889FDE0A6E2" blockId="1.[151,1436,1862,2031]" box="[199,520,2006,2031]" pageId="1" pageNumber="291">
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<emphasis id="B18FEAC94801FFFDFF129889FED2A6E2" bold="true" box="[199,314,2006,2031]" pageId="1" pageNumber="291">Diagnosis</emphasis>
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.
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<collectingCountry id="FBEC764B4801FFFDFE9D9888FE8FA6E2" box="[328,359,2007,2031]" name="American Samoa" pageId="1" pageNumber="291">As</collectingCountry>
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for the genus.
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</paragraph>
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</subSubSection>
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||
<caption id="D78466534802FFFEFF429CA2FA88A51E" box="[151,1376,1021,1043]" httpUri="https://zenodo.org/record/238344/files/figure.png" pageId="2" pageNumber="292" targetBox="[264,1322,179,999]" targetPageId="2">
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<paragraph id="834436DB4802FFFEFF429CA2FA88A51E" blockId="2.[151,1376,1021,1043]" box="[151,1376,1021,1043]" pageId="2" pageNumber="292">
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<emphasis id="B18FEAC94802FFFEFF429CA2FEFBA51F" bold="true" box="[151,275,1021,1043]" pageId="2" pageNumber="292">FIGURE 1.</emphasis>
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<taxonomicName id="44FB4D584802FFFEFECC9CA2FE1EA51E" box="[281,502,1021,1043]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="B18FEAC94802FFFEFECC9CA2FE1EA51E" box="[281,502,1021,1043]" italics="true" pageId="2" pageNumber="292">Urolibellula eocenica</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B18FEAC94802FFFEFE299CA1FDDCA51E" bold="true" box="[508,564,1022,1043]" pageId="2" pageNumber="292">
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||
<taxonomicNameLabel id="AABC57B24802FFFEFE299CA1FDDCA51E" box="[508,564,1022,1043]" pageId="2" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
, holotype specimen, photograph of general habitus (scale bar represents 3 mm).
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||
</paragraph>
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||
</caption>
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||
<caption id="D78466534802FFFEFF429AF1FAE9A4C9" box="[151,1281,1454,1476]" httpUri="https://zenodo.org/record/238345/files/figure.png" pageId="2" pageNumber="292" targetBox="[282,1305,1101,1426]" targetPageId="2">
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||
<paragraph id="834436DB4802FFFEFF429AF1FAE9A4C9" blockId="2.[151,1281,1454,1476]" box="[151,1281,1454,1476]" pageId="2" pageNumber="292">
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||
<emphasis id="B18FEAC94802FFFEFF429AF1FEFAA4CE" bold="true" box="[151,274,1454,1476]" pageId="2" pageNumber="292">FIGURE 2.</emphasis>
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||
<taxonomicName id="44FB4D584802FFFEFECC9AF1FE1EA4C9" box="[281,502,1454,1476]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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||
<emphasis id="B18FEAC94802FFFEFECC9AF1FE1EA4C9" box="[281,502,1454,1476]" italics="true" pageId="2" pageNumber="292">Urolibellula eocenica</emphasis>
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</taxonomicName>
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||
<emphasis id="B18FEAC94802FFFEFE299AF0FDDCA4C9" bold="true" box="[508,564,1455,1476]" pageId="2" pageNumber="292">
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||
<taxonomicNameLabel id="AABC57B24802FFFEFE299AF0FDDCA4C9" box="[508,564,1455,1476]" pageId="2" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, holotype specimen, drawing of forewing (scale bar represents 4 mm).
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</paragraph>
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</caption>
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||
<caption id="D78466534802FFFEFF429886FAEEA6E2" box="[151,1286,2009,2031]" httpUri="https://zenodo.org/record/238346/files/figure.png" pageId="2" pageNumber="292" targetBox="[293,1287,1521,1979]" targetPageId="2">
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<paragraph id="834436DB4802FFFEFF429886FAEEA6E2" blockId="2.[151,1286,2009,2031]" box="[151,1286,2009,2031]" pageId="2" pageNumber="292">
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<emphasis id="B18FEAC94802FFFEFF429886FEFAA6E3" bold="true" box="[151,274,2009,2031]" pageId="2" pageNumber="292">FIGURE 3.</emphasis>
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<taxonomicName id="44FB4D584802FFFEFECC9886FE1EA6E2" box="[281,502,2009,2031]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="B18FEAC94802FFFEFECC9886FE1EA6E2" box="[281,502,2009,2031]" italics="true" pageId="2" pageNumber="292">Urolibellula eocenica</emphasis>
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</taxonomicName>
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||
<emphasis id="B18FEAC94802FFFEFE299886FDDCA6E3" bold="true" box="[508,564,2009,2030]" pageId="2" pageNumber="292">
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||
<taxonomicNameLabel id="AABC57B24802FFFEFE299886FDDCA6E3" box="[508,564,2009,2030]" pageId="2" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, holotype specimen, drawing of hindwing (scale bar represents 4 mm).
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</paragraph>
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||
</caption>
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||
<subSubSection id="CBE165504803FFF8FF129FC8FA69A0F8" lastPageId="4" lastPageNumber="294" pageId="3" pageNumber="293" type="description">
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<paragraph id="834436DB4803FFFFFF129FC8FE56A1D8" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="293">
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<emphasis id="B18FEAC94803FFFFFF129FC8FEB1A1BD" bold="true" box="[199,345,151,176]" pageId="3" pageNumber="293">Description.</emphasis>
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Two forewings, one hind wing, and three legs attached to fragments of the thorax. Wings hyaline, pterostigmata dark brown.
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</paragraph>
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||
<paragraph id="834436DB4803FFFFFF129F80FCFBA5E5" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="293">
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Forewing length 38.0 mm; width at nodus 8.0 mm; distance from base to nodus 20.0 mm; nodus situated distinctly closer to apex than to base; distance from nodus to pterostigma 11.0 mm; distance from base to arculus 6.0 mm; primary antenodal crossveins Axl and Ax2 aligned but not stronger than other 10 secondary antenodals (all complete except most distal one, but not well aligned with antenodals of second row between ScP and RA); Axl 2.0 mm basal of arculus and Ax2 5.0 mm distal of Axl, a long “cordulegastrid gap” and a gap directly distal of arculus; seven postnodal crossveins between nodus and pterostigma not aligned with six postsubnodal crossveins; a “libellulid gap” directly distal of subnodus concerning one postnodal; pterostigma 4.0 mm long and 1.0 mm wide; pterostigma with a basal brace not aligned with its basal side, distally displaced and weakly oblique; pterostigma covering two cells; arculus nearly totally straight and aligned with basal side of discoidal triangle; origins of
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<collectingCountry id="FBEC764B4803FFFFFAAF9D5FFA74A31A" box="[1402,1436,512,535]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
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and MA (sectors of arculus) separated at their extreme bases in arculus; free portion of
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<collectingCountry id="FBEC764B4803FFFFFB7B9D7BFB39A331" box="[1198,1233,548,572]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
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+MA longer than posterior crossvein in arculus; hypertriangle
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<quantity id="44039B3E4803FFFFFD499D17FD1EA352" box="[668,758,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="293" unit="mm" value="3.8">3.8 mm</quantity>
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long and 1.0 mm wide, free; its costal side (MA) weakly curved; discoidal triangle not transverse but broad, divided into three cells by crossveins touching in one point; length of basal side of discoidal triangle
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||
<quantity id="44039B3E4803FFFFFD8F9DCFFD75A3AA" box="[602,669,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="293" unit="mm" value="3.0">3 mm</quantity>
|
||
, length of costal side
|
||
<quantity id="44039B3E4803FFFFFC5B9DCFFC0DA3AA" box="[910,997,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="293" unit="mm" value="2.8">2.8 mm</quantity>
|
||
, length of distal side MAb
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||
<quantity id="44039B3E4803FFFFFAC69DCFFABEA3AA" box="[1299,1366,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="293" unit="mm" value="4.0">4 mm</quantity>
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||
; MAb straight; pseudo-anal vein PsA distinct; subdiscoidal triangle three-celled; basal space free; submedian space only crossed by CuP; anal area max. 2.0 mm wide with two rows of cells; cubito-anal area max.
|
||
<quantity id="44039B3E4803FFFFFB479D88FB00A3E2" box="[1170,1256,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="293" unit="mm" value="4.5">4.5 mm</quantity>
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||
wide with up to five rows of cells; MP weakly curved and ending just distal of nodus; area between CuA and MP not widened; basal part of postdiscoidal area with 2–3 rows of cells; postdiscoidal area not widened distally (width near discoidal triangle
|
||
<quantity id="44039B3E4803FFFFFF2D9C1BFEBDA251" box="[248,341,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="293" unit="mm" value="2.5">2.5 mm</quantity>
|
||
; width at hind margin
|
||
<quantity id="44039B3E4803FFFFFDB29C1BFD2EA251" box="[615,710,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="293" unit="mm" value="2.3">2.3 mm</quantity>
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||
); Mspl nearly straight, ending at MAa, with one row of cells between it and MAa; MAa weakly curved;
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||
<collectingCountry id="FBEC764B4803FFFFFDAB9C37FD4AA28D" box="[638,674,872,896]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
3/4 and MA weakly undulate and closely parallel with only one row of cells between them up to hind margin; first branching of
|
||
<collectingCountry id="FBEC764B4803FFFFFCE09CD2FCBFA2A9" box="[821,855,909,932]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
<quantity id="44039B3E4803FFFFFC8A9CD3FC5DA2A9" box="[863,949,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="293" unit="mm" value="6.1">6.1 mm</quantity>
|
||
basal of subnodus; IR2 originating on RPl/ 2;
|
||
<collectingCountry id="FBEC764B4803FFFFFF619CEFFF3FA2C5" box="[180,215,944,968]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2 aligned with subnodus; only one lestine oblique vein “
|
||
<collectingCountry id="FBEC764B4803FFFFFC9F9CEFFC88A2C5" box="[842,864,944,968]" name="Iceland" pageId="3" pageNumber="293">O</collectingCountry>
|
||
” between
|
||
<collectingCountry id="FBEC764B4803FFFFFC029CEFFC12A2C5" box="[983,1018,944,968]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2 and IR2,
|
||
<quantity id="44039B3E4803FFFFFBA09CF0FB23A2CA" box="[1141,1227,943,968]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="293" unit="mm" value="0.5">0.5 mm</quantity>
|
||
and one cell distal of subnodus; presence of a post-oblique-vein gap; second crossvein between
|
||
<collectingCountry id="FBEC764B4803FFFFFBED9C8BFBB7A2E1" box="[1080,1119,980,1004]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="FBEC764B4803FFFFFB799C8BFB27A2E1" box="[1196,1231,980,1004]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2 developed as a “libellulid oblique vein;” only one bridge crossvein between
|
||
<collectingCountry id="FBEC764B4803FFFFFC869CA7FC9EA51D" box="[851,886,1016,1040]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2 and IR2 basal of subnodus; area between
|
||
<collectingCountry id="FBEC764B4803FFFFFAB99CA7FA67A51D" box="[1388,1423,1016,1040]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2 and IR2 distally widened, but only one row of cells between these veins, except near hind margin (two rows of cells); a well-defined Rspl with one row of cells between it and IR2;
|
||
<collectingCountry id="FBEC764B4803FFFFFC7B9B1FFC39A555" box="[942,977,1088,1112]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2 weakly undulate; RPl and
|
||
<collectingCountry id="FBEC764B4803FFFFFAC19B1FFADFA555" box="[1300,1335,1088,1112]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2 basally parallel with only one row of cells between them, but about 1.0 mm basal of pterostigma, area between these veins widening progressively; a well-defined pseudo-IRl originating on RPl beneath pterostigma; three rows of cells between pseudo-IRl and RPl and between pseudo-IRl and
|
||
<collectingCountry id="FBEC764B4803FFFFFCAA9BF3FC4DA5C9" box="[895,933,1196,1220]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2; no apical supplement (strong concave supplementary sector developed between
|
||
<collectingCountry id="FBEC764B4803FFFFFDBE9B8FFD7AA5E5" box="[619,658,1232,1256]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="FBEC764B4803FFFFFD079B8FFD10A5E5" box="[722,760,1232,1256]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
2).
|
||
</paragraph>
|
||
<paragraph id="834436DB4803FFF8FF129BABFA69A0F8" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="294" pageId="3" pageNumber="293">
|
||
Hindwing: length 37.0 mm; width at nodus 15.0 mm; distance from base to nodus 15.0 mm; nodus situated distinctly closer to base than to apex; distance from nodus to pterostigma
|
||
<quantity id="44039B3E4803FFFFFC089A48FBACA43D" box="[989,1092,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="293" unit="mm" value="15.5">15.5 mm</quantity>
|
||
; distance from base to arculus
|
||
<quantity id="44039B3E4803FFFFFF429A63FF1DA459" box="[151,245,1340,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="293" unit="mm" value="5.5">5.5 mm</quantity>
|
||
; primary antenodal crossveins Axl and Ax2 aligned but not stronger than other five antenodals (all complete); Axl
|
||
<quantity id="44039B3E4803FFFFFE9D9A3FFE62A475" box="[328,394,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="293" unit="mm" value="3.0">3 mm</quantity>
|
||
basal of arculus and Ax2 is
|
||
<quantity id="44039B3E4803FFFFFD189A00FCCBA475" box="[717,803,1375,1400]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="293" unit="mm" value="5.5">5.5 mm</quantity>
|
||
distal of Axl; only four antesubnodal crossveins, with a long “cordulegastrid gap” and a gap directly distal of arculus; 10 postnodal crossveins between nodus and pterostigma not aligned with eight postsubnodal crossveins; a “libellulid gap” directly distal of subnodus concerning two postnodals; pterostigma
|
||
<quantity id="44039B3E4803FFFFFD889A93FD5BA4E9" box="[605,691,1484,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="293" unit="mm" value="3.5">3.5 mm</quantity>
|
||
long and 1.0 mm wide; pterostigma with a basal brace not aligned with its basal side, distally displaced and weakly oblique; pterostigma covering two cells; arculus nearly totally straight and aligned with basal side of discoidal triangle; origins of
|
||
<collectingCountry id="FBEC764B4803FFFFFC4A994AFC29A721" box="[927,961,1557,1580]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
and MA (sectors of arculus) separated at their extreme bases in arculus; free portion of
|
||
<collectingCountry id="FBEC764B4803FFFFFD4F9967FD55A75D" box="[666,701,1592,1616]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
+MA longer than posterior crossvein in arculus; hypertriangle 3.0 mm long and
|
||
<quantity id="44039B3E4803FFFFFEE59903FE7BA779" box="[304,403,1628,1653]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="293" unit="mm" value="0.75">0.75 mm</quantity>
|
||
wide, free; its costal side (MA) distinctly curved; discoidal triangle not transverse but broad, and free; length of basal side of discoidal triangle
|
||
<quantity id="44039B3E4803FFFFFD1E99DFFCCBA795" box="[715,803,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="293" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, length of costal side 3.0 mm, length of distal side MAb 3.0 mm; MAb straight; pseudo-anal vein PsA distinct; subdiscoidal triangle unicellular; basal space free; submedian space only crossed by CuP; anal area max.
|
||
<quantity id="44039B3E4803FFFFFD299998FC88A7ED" box="[764,864,1735,1760]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="293" unit="mm" value="12.5">12.5 mm</quantity>
|
||
wide with about ten rows of cells; cubito-anal area max. 8.0 mm wide with up to six rows of cells; CuA strongly sigmoidally curved and with a strong CuAb; anal loop long and narrow, posteriorly closed and with a concave well-defined midrib with a distinct toe; MP strongly curved and ending just basal of nodus; area between CuA and MP not widened; basal part of postdiscoidal area with only two rows of cells; postdiscoidal area strongly widened distally (width near discoidal triangle
|
||
<quantity id="44039B3E4803FFFFFAE89808FA70A67D" box="[1341,1432,1879,1904]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="293" unit="mm" value="3.5">3.5 mm</quantity>
|
||
; width at hind margin 7.0 mm); a Mspl nearly straight ending on MAa, with one row of cells between it and MAa; MAa weakly curved but with a strong bend near posterior wing margin;
|
||
<collectingCountry id="FBEC764B4803FFFFFC0798FFFC1EA6B5" box="[978,1014,1952,1976]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
3/4 and MA not undulate and closely parallel with only one row of cells between them up to hind margin; first branching of
|
||
<collectingCountry id="FBEC764B4803FFFFFB79989AFB26A6D1" box="[1196,1230,1989,2012]" name="Poland" pageId="3" pageNumber="293">RP</collectingCountry>
|
||
5.0 mm basal of subnodus; IR2 originating on RPl/2;
|
||
<collectingCountry id="FBEC764B4804FFF8FDEF9FC7FDB5A1BD" box="[570,605,152,176]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 aligned with subnodus; only one lestine oblique vein “
|
||
<collectingCountry id="FBEC764B4804FFF8FB089FC7FB1BA1BD" box="[1245,1267,152,176]" name="Iceland" pageId="4" pageNumber="294">O</collectingCountry>
|
||
” between
|
||
<collectingCountry id="FBEC764B4804FFF8FAB99FC7FA67A1BD" box="[1388,1423,152,176]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 and IR2,
|
||
<quantity id="44039B3E4804FFF8FED49FE3FEB0A1D9" box="[257,344,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="294" unit="mm" value="0.5">0.5 mm</quantity>
|
||
and one cell distal of subnodus; presence of a post-oblique-vein gap; second crossvein between
|
||
<collectingCountry id="FBEC764B4804FFF8FF429FBFFF56A1F5" box="[151,190,224,248]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="FBEC764B4804FFF8FED49FBFFECCA1F5" box="[257,292,224,248]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 developed as a “libellulid oblique vein;” only one bridge crossvein between
|
||
<collectingCountry id="FBEC764B4804FFF8FB709FBFFB20A1F5" box="[1189,1224,224,248]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 and IR2 basal of subnodus; area between
|
||
<collectingCountry id="FBEC764B4804FFF8FE7E9E5BFE26A011" box="[427,462,260,284]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 and IR2 distally widened, but only one row of cells between these veins, except near hind margin (two rows of cells); a well-defined Rspl with one row of cells between it and IR2;
|
||
<collectingCountry id="FBEC764B4804FFF8FAC59E77FADBA04D" box="[1296,1331,296,320]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 weakly undulate; RPl and
|
||
<collectingCountry id="FBEC764B4804FFF8FEA19E13FE7FA069" box="[372,407,332,356]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 basally parallel with only one row of cells between them, but about
|
||
<quantity id="44039B3E4804FFF8FB0E9E13FADDA069" box="[1243,1333,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="294" unit="mm" value="1.5">1.5 mm</quantity>
|
||
basal of pterostigma, area between these veins widening progressively; a well-defined pseudo-IRl originating on RPl beneath pterostigma; three rows of cells between pseudo-IRl and RPl and between pseudo-IRl and
|
||
<collectingCountry id="FBEC764B4804FFF8FB239ECBFAF4A0A1" box="[1270,1308,404,428]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2; no apical supplement (strong concave supplementary sector developed between
|
||
<collectingCountry id="FBEC764B4804FFF8FC6D9EE7FC37A0DD" box="[952,991,440,464]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="FBEC764B4804FFF8FBF79EE7FBA0A0DD" box="[1058,1096,440,464]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2); basal hind margin not well preserved so that the presence of an anal angle or an anal triangle cannot be ascertained; no membranule visible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CBE165504804FFF9FF129EA0FAA7A085" lastPageId="5" lastPageNumber="295" pageId="4" pageNumber="294" type="discussion">
|
||
<paragraph id="834436DB4804FFF9FF129EA0FE2BA1D8" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,392]" lastPageId="5" lastPageNumber="295" pageId="4" pageNumber="294">
|
||
<emphasis id="B18FEAC94804FFF8FF129EA0FEADA315" bold="true" box="[199,325,511,536]" pageId="4" pageNumber="294">Discussion</emphasis>
|
||
. The presence of a post-oblique-vein gap is a specialized character present in the
|
||
<taxonomicName id="44FB4D584804FFF8FB0D9EA0FA85A315" box="[1240,1389,511,536]" class="Insecta" family="Corduliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Macromiidae</taxonomicName>
|
||
and most taxa of Haplohamulida (sensu
|
||
<bibRefCitation id="E76A4B2A4804FFF8FD9E9D7BFD0DA330" author="Bechly" box="[587,741,548,573]" pageId="4" pageNumber="294" refString="Bechly, G. (2005) A new fossil dragonfly (Anisoptera: Corduliidae) from the Paleocene Fur Formation (Mo clay) of Denmark. Stuttgarter Beitrage zur Naturkunde, (B), 358, 1 - 7." type="journal article" year="2005">Bechly 2005</bibRefCitation>
|
||
). The presence of a well-defined Mspl, a well-developed, elongate, boot-shaped anal loop with a distinct toe and a distinct Cuspl) is a synapomorphy of the clade Italoansida
|
||
<bibRefCitation id="E76A4B2A4804FFF8FF429D33FEC4A388" author="Bechly" box="[151,300,620,645]" pageId="4" pageNumber="294" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2,1 - 402." type="book" year="1996">Bechly, 1996</bibRefCitation>
|
||
(
|
||
<taxonomicName id="44FB4D584804FFF8FEE89D33FDE1A388" authority="Selys" authorityName="Selys" box="[317,521,620,645]" class="Insecta" family="Corduliidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Corduliidae Selys</taxonomicName>
|
||
in Selys & Hagen, 1850, + Anauriculida
|
||
<bibRefCitation id="E76A4B2A4804FFF8FC3C9D33FB6AA388" author="Bechly" box="[1001,1154,620,645]" pageId="4" pageNumber="294" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2,1 - 402." type="book" year="1996">Bechly, 1996</bibRefCitation>
|
||
). The following putative synapomorphies support a position in the Anauriculida (=
|
||
<taxonomicName id="44FB4D584804FFF8FCE99DD0FB73A3A5" authority="Bechly, 1996" authorityName="Bechly" authorityYear="1996" box="[828,1179,655,680]" class="Insecta" family="Hemicorduliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">
|
||
Hemicorduliidae
|
||
<bibRefCitation id="E76A4B2A4804FFF8FBD79DD0FB73A3A5" author="Bechly" box="[1026,1179,655,680]" pageId="4" pageNumber="294" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2,1 - 402." type="book" year="1996">Bechly, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
, + Libellulida
|
||
<bibRefCitation id="E76A4B2A4804FFF8FA9D9DD0FF38A3C0" author="Bechly" pageId="4" pageNumber="294" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2,1 - 402." type="book" year="1996">Bechly, 1996</bibRefCitation>
|
||
) for
|
||
<taxonomicName id="44FB4D584804FFF8FED29DEBFE78A3C1" box="[263,400,692,716]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FED29DEBFE78A3C1" box="[263,400,692,716]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AABC57B24804FFF8FE429DEBFE0FA3C1" box="[407,487,692,717]" pageId="4" pageNumber="294" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FE429DEBFE37A3C0" bold="true" box="[407,479,692,717]" pageId="4" pageNumber="294">gen. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
: two primary antenodal brackets Ax1 and Ax2 less distinct; besides anal crossing (= CuP) no cubito-anal crossveins present in hindwing (convergent to
|
||
<taxonomicName id="44FB4D584804FFF8FC619D87FB8CA3FD" box="[948,1124,728,752]" class="Insecta" family="Corduliidae" genus="Pseudocordulia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FC619D87FB8CA3FD" box="[948,1124,728,752]" italics="true" pageId="4" pageNumber="294">Pseudocordulia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="44FB4D584804FFF8FBA29D87FAF6A3FD" box="[1143,1310,728,752]" class="Insecta" family="Corduliidae" genus="Austrocordulia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FBA29D87FAF6A3FD" box="[1143,1310,728,752]" italics="true" pageId="4" pageNumber="294">Austrocordulia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="44FB4D584804FFF8FAE49D87FA7DA3FD" box="[1329,1429,728,752]" class="Insecta" family="Corduliidae" genus="Neophya" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FAE49D87FA7DA3FD" box="[1329,1429,728,752]" italics="true" pageId="4" pageNumber="294">Neophya</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="44FB4D584804FFF8FF429DA3FEBBA219" box="[151,339,764,788]" class="Insecta" family="Corduliidae" genus="Hesperocordulia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FF429DA3FEBBA219" box="[151,339,764,788]" italics="true" pageId="4" pageNumber="294">Hesperocordulia</emphasis>
|
||
</taxonomicName>
|
||
; maybe synapomorphic with some
|
||
<taxonomicName id="44FB4D584804FFF8FD3D9DA3FC98A218" box="[744,880,764,789]" class="Insecta" family="Corduliidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Corduliidae</taxonomicName>
|
||
; reversed in some derived libellulids) because of reduction of the basal side of the hindwing subdiscoidal triangle; characteristic curved shape of Rspl that is distally “rejoining” IR2; Mspl very distinct and curved, distally “rejoining” the MA.
|
||
<taxonomicName id="44FB4D584804FFF8FB939C1BFB38A251" box="[1094,1232,836,860]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FB939C1BFB38A251" box="[1094,1232,836,860]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
differs from the
|
||
<taxonomicName id="44FB4D584804FFF8FF429C38FEBBA28D" box="[151,339,871,896]" class="Insecta" family="Hemicorduliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Hemicorduliidae</taxonomicName>
|
||
(
|
||
<taxonomicName id="44FB4D584804FFF8FEB29C37FD7DA28D" authority="Selys, 1870" authorityName="Selys" authorityYear="1870" box="[359,661,871,896]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FEB29C37FDEAA28D" box="[359,514,872,896]" italics="true" pageId="4" pageNumber="294">Hemicordulia</emphasis>
|
||
Selys, 1870
|
||
</taxonomicName>
|
||
) in the most distal antenodal crossvein incomplete and oblique in forewing, presence of a one-celled subdiscoidal triangle in hindwing and discoidal triangle of forewing with three cells not separated by parallel crossveins.
|
||
<taxonomicName id="44FB4D584804FFF8FDA39CEFFD17A2C5" box="[630,767,944,968]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FDA39CEFFD17A2C5" box="[630,767,944,968]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
shares with the Libellulida (=
|
||
<taxonomicName id="44FB4D584804FFF8FBB49CF0FA68A2C5" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" box="[1121,1408,943,968]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae Leach, 1815</taxonomicName>
|
||
, +
|
||
<taxonomicName id="44FB4D584804FFF8FF429C8BFDE2A2E0" authority="Lieftinck, 1954" authorityName="Lieftinck" authorityYear="1954" box="[151,522,980,1005]" class="Insecta" family="Urothemistidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Urothemistidae Lieftinck, 1954</taxonomicName>
|
||
) the following synapomorphies (note that in the phylogeny of the higher Libelluloidea proposed by
|
||
<bibRefCitation id="E76A4B2A4804FFF8FE109CA7FD7BA51D" box="[453,659,1016,1040]" pageId="4" pageNumber="294" refString="Ware, J. L., May, M. & Kjer, K. (2007) Phylogeny of the higher Libelluloidea (Anisoptera: Odonata): An exploration of the most speciose superfamily of dragonflies. Molecular Phylogenetic & Evolution, 45, 289 - 310. http: // dx. doi. org / 10.1016 / j. ympev. 2007.05.027" type="journal article">
|
||
Ware
|
||
<emphasis id="B18FEAC94804FFF8FDD39CA6FDD7A51D" box="[518,575,1016,1040]" italics="true" pageId="4" pageNumber="294">et al.</emphasis>
|
||
(2007)
|
||
</bibRefCitation>
|
||
, the
|
||
<taxonomicName id="44FB4D584804FFF8FD1C9CA8FC9EA51D" box="[713,886,1015,1040]" class="Insecta" family="Urothemistidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Urothemistidae</taxonomicName>
|
||
are considered as a subfamily Urothemistinae of the
|
||
<taxonomicName id="44FB4D584804FFF8FF119B43FEA3A538" box="[196,331,1052,1077]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
but still falls as sister group of the remaining
|
||
<taxonomicName id="44FB4D584804FFF8FCA29B43FBE9A538" box="[887,1025,1052,1077]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
, in the same position as in
|
||
<bibRefCitation id="E76A4B2A4804FFF8FA989B43FF07A555" author="Bechly" pageId="4" pageNumber="294" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2,1 - 402." type="book" year="1996">Bechly (1996))</bibRefCitation>
|
||
: the two primary antenodal brackets Ax1 and Ax2 are very indistinct in forewings and indistinguishable from secondaries in hindwings (even if some secondary antenodals of forewing are not well aligned); anal loop reaching hind margin of hindwing or nearly so (well-developed “toe”) (
|
||
<figureCitation id="1BC02A5E4804FFF8FC319BD8FBBEA5AD" box="[996,1110,1159,1184]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="5.[151,250,1174,1196]" captionStartId-1="5.[151,250,2004,2026]" captionTargetBox-0="[264,1322,447,1152]" captionTargetBox-1="[264,1322,1276,1981]" captionTargetId-0="figure@5.[264,1323,447,1152]" captionTargetId-1="figure@5.[264,1323,1276,1982]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="FIGURE 4. Aethriamanta rezia Kirby, 1889, specimen from Guinée Nimba, Marais Yafélé, alt. 500 m, 16 April 1991 (J. Legrand leg.)" captionText-1="FIGURE 5. Urothemis assignata (Selys, 1872), Sénégal, Casamance, Djibelor, 5 June 1981 (J. Etienne leg.)" httpUri-0="https://zenodo.org/record/238347/files/figure.png" httpUri-1="https://zenodo.org/record/238348/files/figure.png" pageId="4" pageNumber="294">Figs. 4–5</figureCitation>
|
||
).
|
||
<bibRefCitation id="E76A4B2A4804FFF8FBB89BD8FAFAA5AD" author="Bechly" box="[1133,1298,1159,1184]" pageId="4" pageNumber="294" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2,1 - 402." type="book" year="1996">Bechly (1996)</bibRefCitation>
|
||
proposed a further character for this clade, i.e. the second crossvein between
|
||
<collectingCountry id="FBEC764B4804FFF8FCA59BF3FC7FA5C9" box="[880,919,1196,1220]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="FBEC764B4804FFF8FC009BF3FC10A5C9" box="[981,1016,1196,1220]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2 is developed as a “libellulid oblique vein.” This character is not very pronounced in
|
||
<taxonomicName id="44FB4D584804FFF8FD7A9B8FFCD0A5E5" box="[687,824,1232,1256]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FD7A9B8FFCD0A5E5" box="[687,824,1232,1256]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
, as this vein is only weakly oblique, but some modern
|
||
<taxonomicName id="44FB4D584804FFF8FF429BABFEF6A400" box="[151,286,1268,1293]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="44FB4D584804FFF8FEB59BABFD4EA401" authority="Kirby, 1890" authorityName="Kirby" authorityYear="1890" box="[352,678,1268,1293]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="species" species="foliata">
|
||
<emphasis id="B18FEAC94804FFF8FEB59BABFDFEA401" box="[352,534,1268,1292]" italics="true" pageId="4" pageNumber="294">Libellula foliata</emphasis>
|
||
Kirby, 1890
|
||
</taxonomicName>
|
||
) also have this vein not very oblique.
|
||
<taxonomicName id="44FB4D584804FFF8FB8C9BABFB0AA401" box="[1113,1250,1268,1292]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FB8C9BABFB0AA401" box="[1113,1250,1268,1292]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
differs from the
|
||
<taxonomicName id="44FB4D584804FFF8FF429A48FEACA43D" box="[151,324,1303,1328]" class="Insecta" family="Urothemistidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Urothemistidae</taxonomicName>
|
||
in the libellulid oblique vein not very oblique, most distal antenodal incomplete in forewing, presence of a subdiscoidal triangle in hindwing, and forewing discoidal triangle much broader (see
|
||
<bibRefCitation id="E76A4B2A4804FFF8FAF39A63FA65A459" author="Ris" box="[1318,1421,1340,1365]" pageId="4" pageNumber="294" refString="Ris, F. (1909 - 1913) Libellulinen monographisch bearbeitet. In: Collection Zoologique du Baron Edm. de Selys Longchamps. Catalogue systematique et descriptif, 9 - 16 bis, 1 - 1278." type="book chapter" year="1909">
|
||
Ris
|
||
<date id="F745101B4804FFF8FA839A63FA65A459" box="[1366,1421,1340,1364]" pageId="4" pageNumber="294" value="1909" valueMax="1913">1909</date>
|
||
</bibRefCitation>
|
||
– 1913).
|
||
<taxonomicName id="44FB4D584804FFF8FF329A3FFE99A475" box="[231,369,1376,1400]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FF329A3FFE99A475" box="[231,369,1376,1400]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
shares with the
|
||
<taxonomicName id="44FB4D584804FFF8FDF99A00FD5BA475" box="[556,691,1375,1400]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
the following apomorphies: most distal antenodal between costal margin and ScP developed as antenodal oblique vein, with suppression of corresponding antenodal between ScP and RA in forewing; all antenodal crossveins developed as antenodal brackets, therefore the two primaries Ax1 and Ax2 completely indistinguishable from the secondaries in hindwings. This character is also present in the forewings of many
|
||
<taxonomicName id="44FB4D584804FFF8FE459AB0FDFFA705" box="[400,535,1519,1544]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
but not in all (e.g.
|
||
<taxonomicName id="44FB4D584804FFF8FCFE9AAFFB99A705" authority="Kirby, 1889" authorityName="Kirby" authorityYear="1889" box="[811,1137,1519,1544]" class="Insecta" family="Libellulidae" genus="Hydrobasileus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FCFE9AAFFC25A705" box="[811,973,1520,1544]" italics="true" pageId="4" pageNumber="294">Hydrobasileus</emphasis>
|
||
Kirby, 1889
|
||
</taxonomicName>
|
||
, or the oldest libellulid
|
||
<emphasis id="B18FEAC94804FFF8FF42994BFEACA721" box="[151,324,1556,1580]" italics="true" pageId="4" pageNumber="294">Palaeolibellula</emphasis>
|
||
). Nevertheless,
|
||
<taxonomicName id="44FB4D584804FFF8FDD0994BFD67A721" box="[517,655,1556,1580]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FDD0994BFD67A721" box="[517,655,1556,1580]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
lacks two other putative apomorphies of the
|
||
<taxonomicName id="44FB4D584804FFF8FB62994BFAA9A720" box="[1207,1345,1556,1581]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
, i.e. its sectors of arculus are not stalked and it has no apical supplement (strong concave supplementary sector developed between
|
||
<collectingCountry id="FBEC764B4804FFF8FF2A9903FECEA779" box="[255,294,1628,1652]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="FBEC764B4804FFF8FEB89903FE7BA779" box="[365,403,1628,1652]" name="Poland" pageId="4" pageNumber="294">RP</collectingCountry>
|
||
2). Its forewing discoidal triangle is clearly broader than in any modern
|
||
<taxonomicName id="44FB4D584804FFF8FB379903FA81A778" box="[1250,1385,1628,1653]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
and divided into three smaller cells by crossveins not longitudinally disposed. Its subdiscoidal triangle and triangle are both very large with approximatively the same sizes. The modern
|
||
<taxonomicName id="44FB4D584804FFF8FC7F99FBFBD9A7B0" box="[938,1073,1700,1725]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
with triangle and subdiscoidal triangle of similar sizes have a narrow triangle and a subdiscoidal triangle quite smaller than in
|
||
<taxonomicName id="44FB4D584804FFF8FB0D9997FA89A7ED" box="[1240,1377,1736,1760]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FB0D9997FA89A7ED" box="[1240,1377,1736,1760]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
. The forewing discoidal triangle and subdiscoidal triangle of
|
||
<taxonomicName id="44FB4D584804FFF8FCE199B3FC56A609" box="[820,958,1772,1796]" class="Insecta" family="Libellulidae" genus="Urolibellula" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B18FEAC94804FFF8FCE199B3FC56A609" box="[820,958,1772,1796]" italics="true" pageId="4" pageNumber="294">Urolibellula</emphasis>
|
||
</taxonomicName>
|
||
reminds one of some “corduliids” (e.g.
|
||
<taxonomicName id="44FB4D584804FFF8FF42984FFE3BA625" authority="Charpentier, 1825" authorityName="Charpentier" authorityYear="1825" box="[151,467,1807,1832]" class="Insecta" family="Corduliidae" genus="Epitheca" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FF42984FFF13A625" box="[151,251,1808,1832]" italics="true" pageId="4" pageNumber="294">Epitheca</emphasis>
|
||
Charpentier, 1825
|
||
</taxonomicName>
|
||
) (see
|
||
<bibRefCitation id="E76A4B2A4804FFF8FDCB9850FD59A625" author="Martin" box="[542,689,1807,1832]" pageId="4" pageNumber="294" refString="Martin, R. (1914) Odonata. Fam. Libellulidae. Subfam. Cordulinae. Genera Insectorum, 155, 1 - 32." type="journal article" year="1914">Martin 1914</bibRefCitation>
|
||
), even of some Petalurida (
|
||
<taxonomicName id="44FB4D584804FFF8FC23984FFA99A625" authority="Deichmuller, 1886" authorityName="Deichmuller" authorityYear="1886" box="[1014,1393,1807,1832]" class="Insecta" family="Cordulegasteridae" genus="Protolindenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94804FFF8FC23984FFB79A625" box="[1014,1169,1808,1832]" italics="true" pageId="4" pageNumber="294">Protolindenia</emphasis>
|
||
Deichmüller, 1886
|
||
</taxonomicName>
|
||
) or Aeshnoptera (
|
||
<emphasis id="B18FEAC94804FFF8FEE3986BFDEEA641" box="[310,518,1844,1868]" italics="true" pageId="4" pageNumber="294">Paramesuropetala</emphasis>
|
||
<bibRefCitation id="E76A4B2A4804FFF8FDDB986BFD00A641" author="Bechly" box="[526,744,1844,1869]" pageId="4" pageNumber="294" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B18FEAC94804FFF8FDB3986AFD77A641" box="[614,671,1844,1868]" italics="true" pageId="4" pageNumber="294">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
) (
|
||
<bibRefCitation id="E76A4B2A4804FFF8FCD4986BFC58A641" author="Nel" box="[769,944,1844,1869]" pageId="4" pageNumber="294" refString="Nel, A., Bechly, G., Jarzembowski, E. A. & Martinez-Delclos, X. (1998) A revision of the fossil petalurid dragonflies (Insecta: Odonata: Anisoptera: Petalurida). Paleontologia Lombarda, Milano, (N. S.), 10, 1 - 68." type="journal article" year="1998">
|
||
Nel
|
||
<emphasis id="B18FEAC94804FFF8FCE6986AFC85A641" box="[819,877,1844,1868]" italics="true" pageId="4" pageNumber="294">et al.</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E76A4B2A4804FFF8FC6B986BFB7CA641" author="Bechly" box="[958,1172,1844,1869]" pageId="4" pageNumber="294" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B18FEAC94804FFF8FBC0986AFBA7A641" box="[1045,1103,1844,1868]" italics="true" pageId="4" pageNumber="294">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
). Thus this character is clearly subject to convergencies. In conclusion, the Urolibellulidae would fall in a “basal” position as probable sister group of the
|
||
<taxonomicName id="44FB4D584804FFF8FEAB9823FDEDA698" box="[382,517,1916,1941]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="E76A4B2A4804FFF8FDB09823FD13A698" author="Bechly" box="[613,763,1916,1941]" pageId="4" pageNumber="294" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2,1 - 402." type="book" year="1996">Bechly 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E76A4B2A4804FFF8FCD99823FCA0A699" author="Bechly" box="[780,840,1916,1940]" pageId="4" pageNumber="294" refString="Bechly, G. (2014) Phylogenetic Systematics of Odonata. Available from: http: // www. bernstein. naturkundemuseum-bw. de / odonata / intro. htm (accessed 14 October 2014)" type="journal article" year="2014">2014</bibRefCitation>
|
||
), the
|
||
<taxonomicName id="44FB4D584804FFF8FC5A9823FBD4A698" box="[911,1084,1916,1941]" class="Insecta" family="Urothemistidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Urothemistidae</taxonomicName>
|
||
being the sister group of the (Urolibellulidae +
|
||
<taxonomicName id="44FB4D584804FFF8FEB298FFFE1CA6B4" box="[359,500,1952,1977]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
). The presence of a representative of the crown group of the
|
||
<taxonomicName id="44FB4D584804FFF8FB4298FFFAF6A6B4" box="[1175,1310,1952,1977]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="294" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
in the Late Cretaceous shows that the Urolibellulidae were certainly also present in the Late Mesozoic. The stem lineage of the
|
||
<taxonomicName id="44FB4D584805FFF9FF429FC8FEF6A1BD" box="[151,286,151,176]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="295" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
|
||
was probably diversifying during the Late Cretaceous-Eocene even if the fossil record of this clade is very scarce for this period.
|
||
</paragraph>
|
||
<paragraph id="834436DB4805FFF9FF129F80FAA7A085" blockId="5.[151,1437,151,392]" pageId="5" pageNumber="295">
|
||
<emphasis id="B18FEAC94805FFF9FF129F80FED3A1F5" bold="true" box="[199,315,223,248]" pageId="5" pageNumber="295">Remarks.</emphasis>
|
||
<bibRefCitation id="E76A4B2A4805FFF9FE919F80FDEDA1F5" author="Cockerell" box="[324,517,223,248]" pageId="5" pageNumber="295" refString="Cockerell, T. D. A. (1921) Eocene insects from the Rocky Mountains. Proceedings of the United States National Museum, 57, 233 - 260. http: // dx. doi. org / 10.5479 / si. 00963801.57 - 2313.233" type="journal article" year="1921">Cockerell (1921)</bibRefCitation>
|
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described a fossil abdomen from the Eocene Green River Formation under the name
|
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<taxonomicName id="44FB4D584805FFF9FF099E5BFE0FA011" box="[220,487,260,284]" class="Insecta" family="Corduliidae" genus="Stenogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="295" phylum="Arthropoda" rank="species" species="scudderi">
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<emphasis id="B18FEAC94805FFF9FF099E5BFE0FA011" box="[220,487,260,284]" italics="true" pageId="5" pageNumber="295">Stenogomphus scudderi</emphasis>
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</taxonomicName>
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. The ‘libelluloid’ genus
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<taxonomicName id="44FB4D584805FFF9FCD19E5BFBB9A011" authority="Scudder, 1892" authorityName="Scudder" authorityYear="1892" box="[772,1105,260,285]" class="Insecta" family="Corduliidae" genus="Stenogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="295" phylum="Arthropoda" rank="genus">
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<emphasis id="B18FEAC94805FFF9FCD19E5BFC4FA011" box="[772,935,260,284]" italics="true" pageId="5" pageNumber="295">Stenogomphus</emphasis>
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<bibRefCitation id="E76A4B2A4805FFF9FC7A9E5BFBB9A011" author="Scudder" box="[943,1105,260,285]" pageId="5" pageNumber="295" refString="Scudder, S. H. (1892) Some insects of special interest from Florissant, Colorado and other points of the Tertiaries of Colorado and Utah. Bulletin of the United States Geological Survey, 93, 25 pp." type="book" year="1892">Scudder, 1892</bibRefCitation>
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</taxonomicName>
|
||
is based on a forewing from the Earliest Oligocene Florissant Formation (
|
||
<bibRefCitation id="E76A4B2A4805FFF9FD599E78FCC4A04D" author="Scudder" box="[652,812,295,320]" pageId="5" pageNumber="295" refString="Scudder, S. H. (1892) Some insects of special interest from Florissant, Colorado and other points of the Tertiaries of Colorado and Utah. Bulletin of the United States Geological Survey, 93, 25 pp." type="book" year="1892">Scudder 1892</bibRefCitation>
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;
|
||
<bibRefCitation id="E76A4B2A4805FFF9FCE39E78FC75A04D" author="Ris" box="[822,925,295,320]" pageId="5" pageNumber="295" refString="Ris, F. (1910) The identity of two Odonata fossils. Bulletin of the Wisconsin Natural History Society, Milwaukee, 8, 102 - 105." type="journal article" year="1910">Ris 1910</bibRefCitation>
|
||
;
|
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<bibRefCitation id="E76A4B2A4805FFF9FC7D9E78FB75A04D" author="Nel" box="[936,1181,295,320]" pageId="5" pageNumber="295" refString="Nel, A. & Paicheler, J. - C. (1994) Les Libelluloidea fossiles autres que Libellulidae. Un inventaire critique (Odonata, Corduliidae, Macromiidae, Synthemistidae, Chlorogomphidae et Mesophlebiidae). Nouvelle Revue d'Entomologie, (N. S.), 11, 321 - 334." type="journal article" year="1994">Nel & Paicheler 1994</bibRefCitation>
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||
). There is no argument supporting the inclusion of
|
||
<taxonomicName id="44FB4D584805FFF9FE039E13FDBDA069" box="[470,597,332,356]" class="Insecta" family="Corduliidae" genus="Stenogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="295" phylum="Arthropoda" rank="species" species="scudderi">
|
||
<emphasis id="B18FEAC94805FFF9FE039E13FDBDA069" box="[470,597,332,356]" italics="true" pageId="5" pageNumber="295">S. scudderi</emphasis>
|
||
</taxonomicName>
|
||
in the genus
|
||
<taxonomicName id="44FB4D584805FFF9FD239E13FC70A069" box="[758,920,332,356]" class="Insecta" family="Corduliidae" genus="Stenogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B18FEAC94805FFF9FD239E13FC70A069" box="[758,920,332,356]" italics="true" pageId="5" pageNumber="295">Stenogomphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="44FB4D584805FFF9FC7C9E13FB32A069" authority="Scudder, 1892" authorityName="Scudder" authorityYear="1892" box="[937,1242,332,357]" class="Insecta" family="Corduliidae" genus="Stenogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="295" phylum="Arthropoda" rank="species" species="carletoni">
|
||
<emphasis id="B18FEAC94805FFF9FC7C9E13FBC5A069" box="[937,1069,332,356]" italics="true" pageId="5" pageNumber="295">S. carletoni</emphasis>
|
||
<bibRefCitation id="E76A4B2A4805FFF9FBE29E13FB32A069" author="Scudder" box="[1079,1242,332,357]" pageId="5" pageNumber="295" refString="Scudder, S. H. (1892) Some insects of special interest from Florissant, Colorado and other points of the Tertiaries of Colorado and Utah. Bulletin of the United States Geological Survey, 93, 25 pp." type="book" year="1892">Scudder, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
differs from our fossil in the complete last antenodal crossvein, and the second crossvein between
|
||
<collectingCountry id="FBEC764B4805FFF9FBFF9E2FFBB9A085" box="[1066,1105,368,392]" name="Poland" pageId="5" pageNumber="295">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="FBEC764B4805FFF9FB449E2FFB5CA085" box="[1169,1204,368,392]" name="Poland" pageId="5" pageNumber="295">RP</collectingCountry>
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2 not oblique.
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||
</paragraph>
|
||
</subSubSection>
|
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</treatment>
|
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</document> |