629 lines
73 KiB
XML
629 lines
73 KiB
XML
<document id="D5F8783A59B7E3AE1D0692DA105077E8" ID-CLB-Dataset="36839" ID-DOI="10.11646/zootaxa.3814.1.1" ID-GBIF-Dataset="493d9746-7061-41d2-8bff-0713bd07f56f" ID-ISSN="1175-5326" ID-Zenodo-Dep="286211" ID-ZooBank="A024EBCC-AD60-4104-AD4A-4E4843CE6983" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460432840378" checkinUser="plazi" docAuthor="Branch, William R., Bayliss, Julian & Tolley, Krystal A." docDate="2014" docId="03F5304CFFCAFFECFF21FF7D4345EE7D" docLanguage="en" docName="zt03814p036.pdf" docOrigin="Zootaxa 3814 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhampholeon tilburyi Branch, Bayliss & Tolley, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="FFCC4834FFD2FFF7FFB6FFEA4265E875" masterDocTitle="Pygmy chameleons of the Rhampholeon platyceps compex (Squamata: Chamaeleonidae): Description of four new species from isolated ‘ sky islands’ of northern Mozambique" masterLastPageNumber="36" masterPageNumber="1" pageNumber="25" updateTime="1698331660971" updateUser="ExternalLinkService">
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<mods:mods id="BC76CD317EC04122B25BB524113331E4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="55BAA89E207433629948BD82444B7400">
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<mods:title id="6FCA9C692B2BFA96DB4806C70C83D769">Pygmy chameleons of the Rhampholeon platyceps compex (Squamata: Chamaeleonidae): Description of four new species from isolated ‘ sky islands’ of northern Mozambique</mods:title>
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<mods:name id="08BB0E72044C17C8565FC1DB8DDEECDA" type="personal">
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<mods:namePart id="4A3F92515C8C7DE48F30FB7AEC6E6B43">Branch, William R.</mods:namePart>
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<mods:namePart id="6D33F52A54395CF24F1E1F08A1C081FB">Bayliss, Julian</mods:namePart>
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<mods:namePart id="59B2765A2E1B3F069F6C161D24D6F955">Tolley, Krystal A.</mods:namePart>
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<mods:typeOfResource id="F8F2C7AF86918AA2B109BD8BEB1BEEC6">text</mods:typeOfResource>
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<mods:title id="D98EB65E16E94C649B80FF7B4DC50C0B">Zootaxa</mods:title>
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<mods:part id="E93267B72FEEDAF5A67D9A91BF33D818">
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<mods:date id="AB307503857740720942F08D1DBC30D2">2014</mods:date>
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<mods:detail id="854674746B39D1F2A745EB086364C853" type="volume">
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<mods:number id="D5B7651FBB17A3F597C2076607E73F92">3814</mods:number>
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<mods:identifier id="7B715474AA8D0F55EE72C34CCDAFE614" type="DOI">10.11646/zootaxa.3814.1.1</mods:identifier>
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<treatment id="03F5304CFFCAFFECFF21FF7D4345EE7D" ID-DOI="http://doi.org/10.5281/zenodo.6123217" ID-GBIF-Taxon="127668186" ID-Zenodo-Dep="6123217" LSID="urn:lsid:plazi:treatment:03F5304CFFCAFFECFF21FF7D4345EE7D" httpUri="http://treatment.plazi.org/id/03F5304CFFCAFFECFF21FF7D4345EE7D" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
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<subSubSection id="C346D2D1FFCAFFEFFF21FF7D410AE96D" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BE3815AFFCAFFEFFF21FF7D4063E8C7" blockId="24.[151,538,151,211]" box="[151,518,151,178]" pageId="24" pageNumber="25">
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<heading id="D0AB3636FFCAFFEFFF21FF7D4063E8C7" bold="true" box="[151,518,151,178]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
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<emphasis id="B9285D48FFCAFFEFFF21FF7D4063E8C7" bold="true" box="[151,518,151,178]" pageId="24" pageNumber="25">
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<taxonomicName id="4C5CFAD9FFCAFFEFFF21FF7D43C4E8C4" box="[151,417,151,177]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tilburyi" status="sp. nov.">
|
||
<emphasis id="B9285D48FFCAFFEFFF21FF7D43C4E8C4" bold="true" box="[151,417,151,177]" italics="true" pageId="24" pageNumber="25">Rhampholeon tilburyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21BE033FFCAFFEFFE1EFF724063E8C7" box="[424,518,152,178]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCAFFEFFF21FF51407FE8A6" blockId="24.[151,538,151,211]" box="[151,538,186,211]" pageId="24" pageNumber="25">Mount Namuli Pygmy Chameleon</paragraph>
|
||
<paragraph id="8BE3815AFFCAFFEFFF21FEE8410AE96D" blockId="24.[151,1115,258,403]" box="[151,879,258,280]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF21FEE84374E96D" bold="true" box="[151,273,258,280]" pageId="24" pageNumber="25">Synonomy:</emphasis>
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFEE5FEE84140E96D" authority="Branch & Ryan 2001" authorityName="Branch & Ryan" authorityYear="2001" box="[339,805,258,280]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="B9285D48FFCAFFEFFEE5FEE84023E96D" box="[339,582,258,280]" italics="true" pageId="24" pageNumber="25">Rhampholeon platyceps</emphasis>
|
||
<bibRefCitation id="EFCDFCABFFCAFFEFFDFAFEE84140E96D" author="Branch" box="[588,805,258,280]" pageId="24" pageNumber="25" refString="Branch, W. R. & Ryan, P. G. (2001) Additions to the Mozambique Herpetofauna: Two new lizards from the Namuli Massif, Mozambique. Herpetological Review, 32 (4), 281 - 282." type="journal article" year="2001">Branch & Ryan 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, p 282.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D2D1FFCAFFEFFEE5FECB4087E9E6" pageId="24" pageNumber="25" type="reference_group">
|
||
<paragraph id="8BE3815AFFCAFFEFFEE5FECB4087E9E6" blockId="24.[151,1115,258,403]" pageId="24" pageNumber="25">
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFEE5FECB417AE942" authority="Timberlake et al. 2010" authorityName="Timberlake et al." authorityYear="2010" box="[339,799,289,311]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tilburyi">
|
||
<emphasis id="B9285D48FFCAFFEFFEE5FECB4054E942" box="[339,561,289,311]" italics="true" pageId="24" pageNumber="25">Rhampholeon tilburyi</emphasis>
|
||
<bibRefCitation id="EFCDFCABFFCAFFEFFD81FECB417AE942" author="Timberlake" box="[567,799,289,311]" pageId="24" pageNumber="25" refString="Bayliss, J., Monteiro, J., Fishpool, L., Congdon, C., Bampton, I., Bruessow, C., Matimele, H., Banze, A. & Timberlake, J. R. (2010) Biodiversity and Conservation of Mount Inago, Mozambique. Report produced under Darwin Initiative Award 15 / 0 36. Mulanje Mountain Conservation Trust, Malawi, 32 pp." type="book" year="2010">
|
||
Timberlake
|
||
<emphasis id="B9285D48FFCAFFEFFD07FEC840BBE942" box="[689,734,289,311]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, p57
|
||
<emphasis id="B9285D48FFCAFFEFFCE3FEC84385E923" italics="true" pageId="24" pageNumber="25">
|
||
nomen nudum
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFEE5FEAA4385E923" box="[339,480,320,342]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">Rhampholeon</taxonomicName>
|
||
</emphasis>
|
||
nov. sp. (
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFDF4FEAA40E8E923" box="[578,653,320,342]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tilburyi">
|
||
<emphasis id="B9285D48FFCAFFEFFDF4FEAA40E8E923" box="[578,653,320,342]" italics="true" pageId="24" pageNumber="25">tilburyi</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<bibRefCitation id="EFCDFCABFFCAFFEFFD2AFEAA41E1E923" author="Timberlake" box="[668,900,320,342]" pageId="24" pageNumber="25" refString="Bayliss, J., Monteiro, J., Fishpool, L., Congdon, C., Bampton, I., Bruessow, C., Matimele, H., Banze, A. & Timberlake, J. R. (2010) Biodiversity and Conservation of Mount Inago, Mozambique. Report produced under Darwin Initiative Award 15 / 0 36. Mulanje Mountain Conservation Trust, Malawi, 32 pp." type="book" year="2010">
|
||
Timberlake
|
||
<emphasis id="B9285D48FFCAFFEFFCA0FEAB4127E923" box="[790,834,320,342]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
, p61
|
||
<emphasis id="B9285D48FFCAFFEFFC0CFEAB4023E901" italics="true" pageId="24" pageNumber="25">
|
||
nomen nudum
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFEE5FEB44023E901" box="[339,582,350,372]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="platyceps">Rhampholeon platyceps</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFDE2FEB440FAE901" box="[596,671,350,372]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tilburyi">
|
||
<emphasis id="B9285D48FFCAFFEFFDE2FEB440FAE901" box="[596,671,350,372]" italics="true" pageId="24" pageNumber="25">tilburyi</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<bibRefCitation id="EFCDFCABFFCAFFEFFD1BFEB441F3E901" author="Timberlake" box="[685,918,350,373]" pageId="24" pageNumber="25" refString="Bayliss, J., Monteiro, J., Fishpool, L., Congdon, C., Bampton, I., Bruessow, C., Matimele, H., Banze, A. & Timberlake, J. R. (2010) Biodiversity and Conservation of Mount Inago, Mozambique. Report produced under Darwin Initiative Award 15 / 0 36. Mulanje Mountain Conservation Trust, Malawi, 32 pp." type="book" year="2010">
|
||
Timberlake
|
||
<emphasis id="B9285D48FFCAFFEFFC9EFE8A4131E901" box="[808,852,350,373]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
, p61
|
||
<emphasis id="B9285D48FFCAFFEFFC7AFE8A4385E9E6" italics="true" pageId="24" pageNumber="25">
|
||
nomen nudum
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFEE5FE974385E9E6" authority="Portik et al. 2013" authorityName="Portik et al." authorityYear="2013" box="[339,480,381,403]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">Rhampholeon</taxonomicName>
|
||
</emphasis>
|
||
<bibRefCitation id="EFCDFCABFFCAFFEFFE53FE9740FDE9E6" author="Portik" box="[485,664,381,403]" pageId="24" pageNumber="25" refString="Portik, D. M., Travers, S., Bauer, A. M. & Branch, W. R. (2013 b) A new species of Lygodactylus (Squamata: Gekkonidae) endemic to Mount Namuli, an isolated ' sky' island of northern Mozambique. Zootaxa, 3710 (5), 415 - 435. http: // dx. doi. org / 10.11646 / zootaxa. 3710.5.2" type="journal article" year="2013">
|
||
Portik
|
||
<emphasis id="B9285D48FFCAFFEFFD9CFE944033E9E6" box="[554,598,381,403]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
, p 416.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D2D1FFCAFFEFFF21FE554112EAF8" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8BE3815AFFCAFFEFFF21FE5547FDEA1D" blockId="24.[151,1437,447,1157]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF21FE554373E9AD" bold="true" box="[151,278,447,472]" pageId="24" pageNumber="25">Etymology</emphasis>
|
||
. The specific epithet is a patronym for our good friend Dr. Colin Tilbury for his outstanding contributions to knowledge of African chameleons, most recently exemplified in his
|
||
<emphasis id="B9285D48FFCAFFEFFB0DFE0F4707E989" box="[1211,1378,485,508]" italics="true" pageId="24" pageNumber="25">magnum opus</emphasis>
|
||
, the
|
||
<emphasis id="B9285D48FFCAFFEFFF21FDED43E8EA55" box="[151,397,519,544]" italics="true" pageId="24" pageNumber="25">Chameleons of Africa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCDFCABFFCAFFEFFE2AFDED4053EA55" author="Tilbury" box="[412,566,519,544]" pageId="24" pageNumber="25" refString="Tilbury, C. R. (2010) Chameleons of Africa. An Atlas including the Chameleons of Europe, the Middle East and Asia. Edition Chimaira, Frankfurt am Main, 831 pp." type="book" year="2010">Tilbury 2010</bibRefCitation>
|
||
). For the last 30 years his travels throughout Africa in search of chameleons, combined with his stunning photography, have been a source of inspiration. His studies have done much to enhance our understanding of African chameleons, and it is with great pleasure that we name this chameleon in his honour.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCAFFEFFF71FD9E4112EAF8" blockId="24.[151,1437,447,1157]" box="[199,887,628,653]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF71FD9E4376EAF8" bold="true" box="[199,275,628,653]" pageId="24" pageNumber="25">
|
||
<typeStatus id="54E73FF8FFCAFFEFFF71FD9E436AEAF8" box="[199,271,628,653]" pageId="24" pageNumber="25">Types</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="54E73FF8FFCAFFEFFEFBFD9F4318EAF8" box="[333,381,629,653]" pageId="24" pageNumber="25">type</typeStatus>
|
||
series comprises eight specimens, including:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D2D1FFCAFFEFFF71FD7D4702ECF1" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8BE3815AFFCAFFEFFF71FD7D40DBEA8D" blockId="24.[151,1437,447,1157]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF71FD7D434BEAC5" box="[199,302,663,688]" italics="true" pageId="24" pageNumber="25">
|
||
<typeStatus id="54E73FF8FFCAFFEFFF71FD7D434BEAC5" box="[199,302,663,688]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
. An adult female (
|
||
<collectionCode id="ED4D199FFFCAFFEFFDBBFD72402DEAC5" box="[525,584,664,688]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R14921;
|
||
<figureCitation id="13679DDFFFCAFFEFFD0BFD7D4167EAC5" box="[701,770,663,688]" captionStart="FIGURE 8" captionStartId="26.[151,250,1908,1930]" captionTargetBox="[169,1421,211,1859]" captionTargetId="figure@26.[151,1436,193,1887]" captionTargetPageId="26" captionText="FIGURE 8. Rhampholeon tiburyi sp. nov.: A—Allotype (PEM R 17132, K. A. Tolley); B—Paratype (PEM R 17134, K Tolley); C—Head of male paratype (PEM R 20372) showing snout flexure; D—Preserved female holotype (PEM R 14921); E—Manho Forest, Mt. Namuli (J. Bayliss)." httpUri="https://zenodo.org/record/286219/files/figure.png" pageId="24" pageNumber="25">Fig. 8</figureCitation>
|
||
D) collected by a local guide,
|
||
<date id="FFE2A79AFFCAFFEFFBDFFD72472EEAC5" box="[1129,1355,663,688]" pageId="24" pageNumber="25" value="1998-11-30">30 November 1998</date>
|
||
, in the Ukalini Forest that nestles under the south face of the main Namuli peak, Namuli Massif, Zambézia Province,
|
||
<collectingCountry id="F34BC1CAFFCAFFEFFF21FD35434EEA8D" box="[151,299,735,760]" name="Mozambique" pageId="24" pageNumber="25">Mozambique</collectingCountry>
|
||
(15˚22'S 37˚04'E, ca
|
||
<quantity id="4CA42CBFFFCAFFEFFDABFD354017EA82" box="[541,626,735,760]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="24" pageNumber="25" unit="m" value="1550.0">1550 m</quantity>
|
||
a.s.l.).
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCAFFEFFF71FCEE4143EBFD" blockId="24.[151,1437,447,1157]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF71FCEE4344EB68" box="[199,289,772,797]" italics="true" pageId="24" pageNumber="25">Allotype</emphasis>
|
||
. An adult male (
|
||
<collectionCode id="ED4D199FFFCAFFEFFE69FCEE407CEB69" box="[479,537,772,796]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R17132,
|
||
<figureCitation id="13679DDFFFCAFFEFFD26FCEE40B5EB69" box="[656,720,772,797]" captionStart="FIGURE 8" captionStartId="26.[151,250,1908,1930]" captionTargetBox="[169,1421,211,1859]" captionTargetId="figure@26.[151,1436,193,1887]" captionTargetPageId="26" captionText="FIGURE 8. Rhampholeon tiburyi sp. nov.: A—Allotype (PEM R 17132, K. A. Tolley); B—Paratype (PEM R 17134, K Tolley); C—Head of male paratype (PEM R 20372) showing snout flexure; D—Preserved female holotype (PEM R 14921); E—Manho Forest, Mt. Namuli (J. Bayliss)." httpUri="https://zenodo.org/record/286219/files/figure.png" pageId="24" pageNumber="25">Fig. 8</figureCitation>
|
||
A) with everted hemipenes, collected by K.A. Tolley and S. van Noort,
|
||
<date id="FFE2A79AFFCAFFEFFF5EFCCD43E1EB35" box="[232,388,807,832]" pageId="24" pageNumber="25" value="2006-05-26">26 May 2006</date>
|
||
, at night perched on a dead branch
|
||
<quantity id="4CA42CBFFFCAFFEFFC90FCCD4108EB4A" box="[806,877,807,832]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="cm" value="50.0">50 cm</quantity>
|
||
above ground in a patch of forest dominated by overgrown tea (
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFEFEFCA6406EEB11" box="[328,523,844,869]" class="Magnoliopsida" family="Theaceae" genus="Camellia" kingdom="Plantae" order="Ericales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="sinensis">
|
||
<emphasis id="B9285D48FFCAFFEFFEFEFCA6406EEB11" box="[328,523,844,869]" italics="true" pageId="24" pageNumber="25">Camellia sinensis</emphasis>
|
||
</taxonomicName>
|
||
) on the SDZ Cha Sarl Tea Estate at Gurué, Namuli Massif, Zambézia Province,
|
||
<collectingCountry id="F34BC1CAFFCAFFEFFF21FC854342EBFD" box="[151,295,879,904]" name="Mozambique" pageId="24" pageNumber="25">Mozambique</collectingCountry>
|
||
(15˚26'51.6”S, 37˚00’,32.6”E, ca
|
||
<quantity id="4CA42CBFFFCAFFEFFD2FFC9A40BBEBF2" box="[665,734,880,904]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="24" pageNumber="25" unit="m" value="839.0">839 m</quantity>
|
||
a.s.l.).
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCAFFEFFF71FC7F41B8EC15" blockId="24.[151,1437,447,1157]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF71FC7F4350EBD9" box="[199,309,917,940]" italics="true" pageId="24" pageNumber="25">
|
||
<typeStatus id="54E73FF8FFCAFFEFFF71FC7F4350EBD9" box="[199,309,917,940]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
. Six specimens, comprising two females (
|
||
<collectionCode id="ED4D199FFFCAFFEFFCBBFC7E4122EBD9" box="[781,839,916,940]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R17134,
|
||
<figureCitation id="13679DDFFFCAFFEFFC00FC7E419DEBD9" box="[950,1016,916,941]" captionStart="FIGURE 8" captionStartId="26.[151,250,1908,1930]" captionTargetBox="[169,1421,211,1859]" captionTargetId="figure@26.[151,1436,193,1887]" captionTargetPageId="26" captionText="FIGURE 8. Rhampholeon tiburyi sp. nov.: A—Allotype (PEM R 17132, K. A. Tolley); B—Paratype (PEM R 17134, K Tolley); C—Head of male paratype (PEM R 20372) showing snout flexure; D—Preserved female holotype (PEM R 14921); E—Manho Forest, Mt. Namuli (J. Bayliss)." httpUri="https://zenodo.org/record/286219/files/figure.png" pageId="24" pageNumber="25">Fig. 8</figureCitation>
|
||
B, same collecting details as allotype;
|
||
<collectionCode id="ED4D199FFFCAFFEFFF21FC5242B5EBBA" box="[151,208,952,975]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R17135, perched on a tea bush in an overgrown patch of tea, on SDZ Cha Sarl Tea Estate at Gurué, 15˚26'42.8”S, 37˚00’,19.2”E); three males (
|
||
<collectionCode id="ED4D199FFFCAFFEFFD21FC3640B4EB81" box="[663,721,988,1012]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R17131, SDZ Cha Sarl Tea Estate at Gurué, 15˚26'49”S, 37˚00’,29”E;
|
||
<collectionCode id="ED4D199FFFCAFFEFFE87FBEA430CEC62" box="[305,361,1024,1047]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R17133, same collecting details as allotype;
|
||
<collectionCode id="ED4D199FFFCAFFEFFCC1FBEA41D5EC62" box="[887,944,1024,1047]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R20372, Muretha Plateau, Namuli Massif, Zambézia Province,
|
||
<collectingCountry id="F34BC1CAFFCAFFEFFECCFBCE406FEC48" box="[378,522,1060,1085]" name="Mozambique" pageId="24" pageNumber="25">Mozambique</collectingCountry>
|
||
(15˚23'26”S, 37˚02’,03”E, ca
|
||
<quantity id="4CA42CBFFFCAFFEFFCEDFBCE41CAEC49" box="[859,943,1060,1084]" metricMagnitude="3" metricUnit="m" metricValue="1.8039999999999998" pageId="24" pageNumber="25" unit="m" value="1804.0">1804 m</quantity>
|
||
a.s.l.), collected by J. Bayliss
|
||
<date id="FFE2A79AFFCAFFEFFAB6FBCE47FDEC49" box="[1280,1432,1060,1085]" pageId="24" pageNumber="25" value="2007-05-27">27 May 2007</date>
|
||
; and a very small juvenile (
|
||
<collectionCode id="ED4D199FFFCAFFEFFE03FBA2438AEC15" box="[437,495,1096,1120]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="24" pageNumber="25">PEM</collectionCode>
|
||
R20373, same details as previous specimen).
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCAFFEFFF71FB864702ECF1" blockId="24.[151,1437,447,1157]" box="[199,1383,1132,1157]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF71FB864359ECF0" bold="true" box="[199,316,1132,1157]" pageId="24" pageNumber="25">Meristics.</emphasis>
|
||
Measurements for the
|
||
<typeStatus id="54E73FF8FFCAFFEFFD88FB87400BECF0" box="[574,622,1133,1157]" pageId="24" pageNumber="25">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFD6DFB8641AAECF0" box="[731,975,1132,1157]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tilburyi" status="sp. nov.">
|
||
<emphasis id="B9285D48FFCAFFEFFD6DFB8641AAECF0" box="[731,975,1132,1157]" italics="true" pageId="24" pageNumber="25">Rhampholeon tilburyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9285D48FFCAFFEFFC60FB864655ECF0" bold="true" box="[982,1072,1132,1157]" pageId="24" pageNumber="25">
|
||
<taxonomicNameLabel id="A21BE033FFCAFFEFFC60FB864655ECF0" box="[982,1072,1132,1157]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are summarized in
|
||
<tableCitation id="C6DEB4E1FFCAFFEFFABBFB864707ECF1" box="[1293,1378,1132,1157]" captionStart="TABLE 8" captionStartId="24.[151,235,1204,1227]" captionTargetBox="[159,1407,1287,1824]" captionTargetPageId="24" captionText="TABLE 8. Measurements (mm) for the type series of Rhampholeon tilburyi sp. nov. from Mount Namuli, Mozambique (PEM: Port Elizabeth Museum)." httpUri="http://table.plazi.org/id/DF23D1D2FFCAFFEFFF21FB5E4394EC9E" pageId="24" pageNumber="25" tableUuid="DF23D1D2FFCAFFEFFF21FB5E4394EC9E">Table 8</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF23D1D2FFCAFFEFFF21FB5E4394EC9E" ID-Table-UUID="DF23D1D2FFCAFFEFFF21FB5E4394EC9E" httpUri="http://table.plazi.org/id/DF23D1D2FFCAFFEFFF21FB5E4394EC9E" pageId="24" pageNumber="25" targetBox="[159,1407,1287,1824]" targetIsTable="true" targetPageId="24">
|
||
<paragraph id="8BE3815AFFCAFFEFFF21FB5E4394EC9E" blockId="24.[151,1435,1203,1259]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF21FB5E436DECBE" bold="true" box="[151,264,1204,1227]" pageId="24" pageNumber="25">TABLE 8.</emphasis>
|
||
Measurements (mm) for the type series of
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFD69FB5941A6ECBE" box="[735,963,1203,1227]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="tilburyi" status="sp. nov.">
|
||
<emphasis id="B9285D48FFCAFFEFFD69FB5941A6ECBE" box="[735,963,1203,1227]" italics="true" pageId="24" pageNumber="25">Rhampholeon tilburyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9285D48FFCAFFEFFC7BFB5E4646ECBE" bold="true" box="[973,1059,1204,1227]" pageId="24" pageNumber="25">
|
||
<taxonomicNameLabel id="A21BE033FFCAFFEFFC7BFB5E4646ECBE" box="[973,1059,1204,1227]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Mount Namuli, Mozambique (PEM: Port Elizabeth Museum).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE3815AFFCAFFEFFE33FAED4722EF55" pageId="24" pageNumber="25">
|
||
<table id="F95C73FAFFCA0008FF29FAED471AEF55" box="[159,1407,1287,1824]" gridcols="9" gridrows="12" pageId="24" pageNumber="25">
|
||
<tr id="356C8318FFCA0008FF29FAED471AED2F" box="[159,1407,1287,1370]" gridrow="0" pageId="24" pageNumber="25" rowspan-0="1">
|
||
<th id="76BDEA64FFCA0008FE33FAED4386ED2F" box="[389,483,1287,1370]" gridcol="1" gridrow="0" pageId="24" pageNumber="25">PEM R14921 Holotype</th>
|
||
<th id="76BDEA64FFCA0008FDB8FAED4003ED2F" box="[526,614,1287,1370]" gridcol="2" gridrow="0" pageId="24" pageNumber="25">PEM R17132 Allotype</th>
|
||
<th id="76BDEA64FFCA0008FD21FAED408AED2F" box="[663,751,1287,1370]" gridcol="3" gridrow="0" pageId="24" pageNumber="25">PEM R17131 Paratype</th>
|
||
<th id="76BDEA64FFCA0008FC96FAED411DED2F" box="[800,888,1287,1370]" gridcol="4" gridrow="0" pageId="24" pageNumber="25">PEM R17133 Paratype</th>
|
||
<th id="76BDEA64FFCA0008FC1FFAED4664ED2F" box="[937,1025,1287,1370]" gridcol="5" gridrow="0" pageId="24" pageNumber="25">PEM R17134 Paratype</th>
|
||
<th id="76BDEA64FFCA0008FB9DFAED46E1ED2F" box="[1067,1156,1287,1370]" gridcol="6" gridrow="0" pageId="24" pageNumber="25">PEM R17135 Paratype</th>
|
||
<th id="76BDEA64FFCA0008FB1FFAED4767ED2F" box="[1193,1282,1287,1370]" gridcol="7" gridrow="0" pageId="24" pageNumber="25">PEM R20372 Paratype</th>
|
||
<th id="76BDEA64FFCA0008FA91FAED471AED2F" box="[1319,1407,1287,1370]" gridcol="8" gridrow="0" pageId="24" pageNumber="25">PEM R20373 Paratype</th>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29FA87471AEDF6" box="[159,1407,1389,1411]" gridrow="1" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29FA874336EDF6" box="[159,339,1389,1411]" gridcol="0" gridrow="1" pageId="24" pageNumber="25">Sex</th>
|
||
<td id="76BDEA64FFCA0008FE33FA874386EDF6" box="[389,483,1389,1411]" gridcol="1" gridrow="1" pageId="24" pageNumber="25">F</td>
|
||
<td id="76BDEA64FFCA0008FDB8FA874003EDF6" box="[526,614,1389,1411]" gridcol="2" gridrow="1" pageId="24" pageNumber="25">M</td>
|
||
<td id="76BDEA64FFCA0008FD21FA87408AEDF6" box="[663,751,1389,1411]" gridcol="3" gridrow="1" pageId="24" pageNumber="25">M</td>
|
||
<td id="76BDEA64FFCA0008FC96FA87411DEDF6" box="[800,888,1389,1411]" gridcol="4" gridrow="1" pageId="24" pageNumber="25">M</td>
|
||
<td id="76BDEA64FFCA0008FC1FFA874664EDF6" box="[937,1025,1389,1411]" gridcol="5" gridrow="1" pageId="24" pageNumber="25">F</td>
|
||
<td id="76BDEA64FFCA0008FB9DFA8746E1EDF6" box="[1067,1156,1389,1411]" gridcol="6" gridrow="1" pageId="24" pageNumber="25">F</td>
|
||
<td id="76BDEA64FFCA0008FB1FFA874767EDF6" box="[1193,1282,1389,1411]" gridcol="7" gridrow="1" pageId="24" pageNumber="25">M</td>
|
||
<td id="76BDEA64FFCA0008FA91FA87471AEDF6" box="[1319,1407,1389,1411]" gridcol="8" gridrow="1" pageId="24" pageNumber="25">J</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29FA7C471AEDD9" box="[159,1407,1430,1452]" gridrow="2" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29FA7C4336EDD9" box="[159,339,1430,1452]" gridcol="0" gridrow="2" pageId="24" pageNumber="25">Snout-vent length</th>
|
||
<td id="76BDEA64FFCA0008FE33FA7C4386EDD9" box="[389,483,1430,1452]" gridcol="1" gridrow="2" pageId="24" pageNumber="25">55.3</td>
|
||
<td id="76BDEA64FFCA0008FDB8FA7C4003EDD9" box="[526,614,1430,1452]" gridcol="2" gridrow="2" pageId="24" pageNumber="25">39.6</td>
|
||
<td id="76BDEA64FFCA0008FD21FA7C408AEDD9" box="[663,751,1430,1452]" gridcol="3" gridrow="2" pageId="24" pageNumber="25">33.2</td>
|
||
<td id="76BDEA64FFCA0008FC96FA7C411DEDD9" box="[800,888,1430,1452]" gridcol="4" gridrow="2" pageId="24" pageNumber="25">34.6</td>
|
||
<td id="76BDEA64FFCA0008FC1FFA7C4664EDD9" box="[937,1025,1430,1452]" gridcol="5" gridrow="2" pageId="24" pageNumber="25">48.8</td>
|
||
<td id="76BDEA64FFCA0008FB9DFA7C46E1EDD9" box="[1067,1156,1430,1452]" gridcol="6" gridrow="2" pageId="24" pageNumber="25">50</td>
|
||
<td id="76BDEA64FFCA0008FB1FFA7C4767EDD9" box="[1193,1282,1430,1452]" gridcol="7" gridrow="2" pageId="24" pageNumber="25">48.8</td>
|
||
<td id="76BDEA64FFCA0008FA91FA7C471AEDD9" box="[1319,1407,1430,1452]" gridcol="8" gridrow="2" pageId="24" pageNumber="25">21.4</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29FA2A471AEDA3" box="[159,1407,1472,1494]" gridrow="3" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29FA2A4336EDA3" box="[159,339,1472,1494]" gridcol="0" gridrow="3" pageId="24" pageNumber="25">Tail length</th>
|
||
<td id="76BDEA64FFCA0008FE33FA2A4386EDA3" box="[389,483,1472,1494]" gridcol="1" gridrow="3" pageId="24" pageNumber="25">15.4</td>
|
||
<td id="76BDEA64FFCA0008FDB8FA2A4003EDA3" box="[526,614,1472,1494]" gridcol="2" gridrow="3" pageId="24" pageNumber="25">12.5</td>
|
||
<td id="76BDEA64FFCA0008FD21FA2A408AEDA3" box="[663,751,1472,1494]" gridcol="3" gridrow="3" pageId="24" pageNumber="25">10.5</td>
|
||
<td id="76BDEA64FFCA0008FC96FA2A411DEDA3" box="[800,888,1472,1494]" gridcol="4" gridrow="3" pageId="24" pageNumber="25">11.8</td>
|
||
<td id="76BDEA64FFCA0008FC1FFA2A4664EDA3" box="[937,1025,1472,1494]" gridcol="5" gridrow="3" pageId="24" pageNumber="25">10.9</td>
|
||
<td id="76BDEA64FFCA0008FB9DFA2A46E1EDA3" box="[1067,1156,1472,1494]" gridcol="6" gridrow="3" pageId="24" pageNumber="25">13.8</td>
|
||
<td id="76BDEA64FFCA0008FB1FFA2A4767EDA3" box="[1193,1282,1472,1494]" gridcol="7" gridrow="3" pageId="24" pageNumber="25">15.5</td>
|
||
<td id="76BDEA64FFCA0008FA91FA2A471AEDA3" box="[1319,1407,1472,1494]" gridcol="8" gridrow="3" pageId="24" pageNumber="25">7.4</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29FA03471AED8A" box="[159,1407,1513,1535]" gridrow="4" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29FA034336ED8A" box="[159,339,1513,1535]" gridcol="0" gridrow="4" pageId="24" pageNumber="25">Total length</th>
|
||
<td id="76BDEA64FFCA0008FE33FA034386ED8A" box="[389,483,1513,1535]" gridcol="1" gridrow="4" pageId="24" pageNumber="25">70.7</td>
|
||
<td id="76BDEA64FFCA0008FDB8FA034003ED8A" box="[526,614,1513,1535]" gridcol="2" gridrow="4" pageId="24" pageNumber="25">52.1</td>
|
||
<td id="76BDEA64FFCA0008FD21FA03408AED8A" box="[663,751,1513,1535]" gridcol="3" gridrow="4" pageId="24" pageNumber="25">43.7</td>
|
||
<td id="76BDEA64FFCA0008FC96FA03411DED8A" box="[800,888,1513,1535]" gridcol="4" gridrow="4" pageId="24" pageNumber="25">46.4</td>
|
||
<td id="76BDEA64FFCA0008FC1FFA034664ED8A" box="[937,1025,1513,1535]" gridcol="5" gridrow="4" pageId="24" pageNumber="25">59.7</td>
|
||
<td id="76BDEA64FFCA0008FB9DFA0346E1ED8A" box="[1067,1156,1513,1535]" gridcol="6" gridrow="4" pageId="24" pageNumber="25">63.8</td>
|
||
<td id="76BDEA64FFCA0008FB1FFA034767ED8A" box="[1193,1282,1513,1535]" gridcol="7" gridrow="4" pageId="24" pageNumber="25">64.4</td>
|
||
<td id="76BDEA64FFCA0008FA91FA03471AED8A" box="[1319,1407,1513,1535]" gridcol="8" gridrow="4" pageId="24" pageNumber="25">28.8</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29F9F8471AEE5D" box="[159,1407,1554,1576]" gridrow="5" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29F9F84336EE5D" box="[159,339,1554,1576]" gridcol="0" gridrow="5" pageId="24" pageNumber="25">Head length</th>
|
||
<td id="76BDEA64FFCA0008FE33F9F84386EE5D" box="[389,483,1554,1576]" gridcol="1" gridrow="5" pageId="24" pageNumber="25">21.2</td>
|
||
<td id="76BDEA64FFCA0008FDB8F9F84003EE5D" box="[526,614,1554,1576]" gridcol="2" gridrow="5" pageId="24" pageNumber="25">12.5</td>
|
||
<td id="76BDEA64FFCA0008FD21F9F8408AEE5D" box="[663,751,1554,1576]" gridcol="3" gridrow="5" pageId="24" pageNumber="25">11.0</td>
|
||
<td id="76BDEA64FFCA0008FC96F9F8411DEE5D" box="[800,888,1554,1576]" gridcol="4" gridrow="5" pageId="24" pageNumber="25">11.8</td>
|
||
<td id="76BDEA64FFCA0008FC1FF9F84664EE5D" box="[937,1025,1554,1576]" gridcol="5" gridrow="5" pageId="24" pageNumber="25">15.5</td>
|
||
<td id="76BDEA64FFCA0008FB9DF9F846E1EE5D" box="[1067,1156,1554,1576]" gridcol="6" gridrow="5" pageId="24" pageNumber="25">20.8</td>
|
||
<td id="76BDEA64FFCA0008FB1FF9F84767EE5D" box="[1193,1282,1554,1576]" gridcol="7" gridrow="5" pageId="24" pageNumber="25">17.3</td>
|
||
<td id="76BDEA64FFCA0008FA91F9F8471AEE5D" box="[1319,1407,1554,1576]" gridcol="8" gridrow="5" pageId="24" pageNumber="25">8.6</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29F9D6471AEE27" box="[159,1407,1596,1618]" gridrow="6" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29F9D64336EE27" box="[159,339,1596,1618]" gridcol="0" gridrow="6" pageId="24" pageNumber="25">Head width</th>
|
||
<td id="76BDEA64FFCA0008FE33F9D64386EE27" box="[389,483,1596,1618]" gridcol="1" gridrow="6" pageId="24" pageNumber="25">10.0</td>
|
||
<td id="76BDEA64FFCA0008FDB8F9D64003EE27" box="[526,614,1596,1618]" gridcol="2" gridrow="6" pageId="24" pageNumber="25">7.3</td>
|
||
<td id="76BDEA64FFCA0008FD21F9D6408AEE27" box="[663,751,1596,1618]" gridcol="3" gridrow="6" pageId="24" pageNumber="25">5.5</td>
|
||
<td id="76BDEA64FFCA0008FC96F9D6411DEE27" box="[800,888,1596,1618]" gridcol="4" gridrow="6" pageId="24" pageNumber="25">5.3</td>
|
||
<td id="76BDEA64FFCA0008FC1FF9D64664EE27" box="[937,1025,1596,1618]" gridcol="5" gridrow="6" pageId="24" pageNumber="25">9.1</td>
|
||
<td id="76BDEA64FFCA0008FB9DF9D646E1EE27" box="[1067,1156,1596,1618]" gridcol="6" gridrow="6" pageId="24" pageNumber="25">6.4</td>
|
||
<td id="76BDEA64FFCA0008FB1FF9D64767EE27" box="[1193,1282,1596,1618]" gridcol="7" gridrow="6" pageId="24" pageNumber="25">9.6</td>
|
||
<td id="76BDEA64FFCA0008FA91F9D6471AEE27" box="[1319,1407,1596,1618]" gridcol="8" gridrow="6" pageId="24" pageNumber="25">4.3</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29F98F471AEE0E" box="[159,1407,1637,1659]" gridrow="7" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29F98F4336EE0E" box="[159,339,1637,1659]" gridcol="0" gridrow="7" pageId="24" pageNumber="25">Orbit diameter</th>
|
||
<td id="76BDEA64FFCA0008FE33F98F4386EE0E" box="[389,483,1637,1659]" gridcol="1" gridrow="7" pageId="24" pageNumber="25">5.1</td>
|
||
<td id="76BDEA64FFCA0008FDB8F98F4003EE0E" box="[526,614,1637,1659]" gridcol="2" gridrow="7" pageId="24" pageNumber="25">3.2</td>
|
||
<td id="76BDEA64FFCA0008FD21F98F408AEE0E" box="[663,751,1637,1659]" gridcol="3" gridrow="7" pageId="24" pageNumber="25">2.8</td>
|
||
<td id="76BDEA64FFCA0008FC96F98F411DEE0E" box="[800,888,1637,1659]" gridcol="4" gridrow="7" pageId="24" pageNumber="25">3.0</td>
|
||
<td id="76BDEA64FFCA0008FC1FF98F4664EE0E" box="[937,1025,1637,1659]" gridcol="5" gridrow="7" pageId="24" pageNumber="25">3.9</td>
|
||
<td id="76BDEA64FFCA0008FB9DF98F46E1EE0E" box="[1067,1156,1637,1659]" gridcol="6" gridrow="7" pageId="24" pageNumber="25">4.6</td>
|
||
<td id="76BDEA64FFCA0008FB1FF98F4767EE0E" box="[1193,1282,1637,1659]" gridcol="7" gridrow="7" pageId="24" pageNumber="25">4.5</td>
|
||
<td id="76BDEA64FFCA0008FA91F98F471AEE0E" box="[1319,1407,1637,1659]" gridcol="8" gridrow="7" pageId="24" pageNumber="25">2.4</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29F964471AEED1" box="[159,1407,1678,1700]" gridrow="8" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29F9644336EED1" box="[159,339,1678,1700]" gridcol="0" gridrow="8" pageId="24" pageNumber="25">Snout length</th>
|
||
<td id="76BDEA64FFCA0008FE33F9644386EED1" box="[389,483,1678,1700]" gridcol="1" gridrow="8" pageId="24" pageNumber="25">6.4</td>
|
||
<td id="76BDEA64FFCA0008FDB8F9644003EED1" box="[526,614,1678,1700]" gridcol="2" gridrow="8" pageId="24" pageNumber="25">3.7</td>
|
||
<td id="76BDEA64FFCA0008FD21F964408AEED1" box="[663,751,1678,1700]" gridcol="3" gridrow="8" pageId="24" pageNumber="25">3.8</td>
|
||
<td id="76BDEA64FFCA0008FC96F964411DEED1" box="[800,888,1678,1700]" gridcol="4" gridrow="8" pageId="24" pageNumber="25">3.9</td>
|
||
<td id="76BDEA64FFCA0008FC1FF9644664EED1" box="[937,1025,1678,1700]" gridcol="5" gridrow="8" pageId="24" pageNumber="25">5.3</td>
|
||
<td id="76BDEA64FFCA0008FB9DF96446E1EED1" box="[1067,1156,1678,1700]" gridcol="6" gridrow="8" pageId="24" pageNumber="25">5.5</td>
|
||
<td id="76BDEA64FFCA0008FB1FF9644767EED1" box="[1193,1282,1678,1700]" gridcol="7" gridrow="8" pageId="24" pageNumber="25">6.0</td>
|
||
<td id="76BDEA64FFCA0008FA91F964471AEED1" box="[1319,1407,1678,1700]" gridcol="8" gridrow="8" pageId="24" pageNumber="25">3.1</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29F952471AEEBB" box="[159,1407,1720,1742]" gridrow="9" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29F9524336EEBB" box="[159,339,1720,1742]" gridcol="0" gridrow="9" pageId="24" pageNumber="25">Inter-orbital</th>
|
||
<td id="76BDEA64FFCA0008FE33F9524386EEBB" box="[389,483,1720,1742]" gridcol="1" gridrow="9" pageId="24" pageNumber="25">5.8</td>
|
||
<td id="76BDEA64FFCA0008FDB8F9524003EEBB" box="[526,614,1720,1742]" gridcol="2" gridrow="9" pageId="24" pageNumber="25">4.7</td>
|
||
<td id="76BDEA64FFCA0008FD21F952408AEEBB" box="[663,751,1720,1742]" gridcol="3" gridrow="9" pageId="24" pageNumber="25">4.5</td>
|
||
<td id="76BDEA64FFCA0008FC96F952411DEEBB" box="[800,888,1720,1742]" gridcol="4" gridrow="9" pageId="24" pageNumber="25">5.0</td>
|
||
<td id="76BDEA64FFCA0008FC1FF9524664EEBB" box="[937,1025,1720,1742]" gridcol="5" gridrow="9" pageId="24" pageNumber="25">5.5</td>
|
||
<td id="76BDEA64FFCA0008FB9DF95246E1EEBB" box="[1067,1156,1720,1742]" gridcol="6" gridrow="9" pageId="24" pageNumber="25">5.8</td>
|
||
<td id="76BDEA64FFCA0008FB1FF9524767EEBB" box="[1193,1282,1720,1742]" gridcol="7" gridrow="9" pageId="24" pageNumber="25">5.6</td>
|
||
<td id="76BDEA64FFCA0008FA91F952471AEEBB" box="[1319,1407,1720,1742]" gridcol="8" gridrow="9" pageId="24" pageNumber="25">3.5</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29F90B471AEE82" box="[159,1407,1761,1783]" gridrow="10" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29F90B4336EE82" box="[159,339,1761,1783]" gridcol="0" gridrow="10" pageId="24" pageNumber="25">Rostral process</th>
|
||
<td id="76BDEA64FFCA0008FE33F90B4386EE82" box="[389,483,1761,1783]" gridcol="1" gridrow="10" pageId="24" pageNumber="25">1.8</td>
|
||
<td id="76BDEA64FFCA0008FDB8F90B4003EE82" box="[526,614,1761,1783]" gridcol="2" gridrow="10" pageId="24" pageNumber="25">2.0</td>
|
||
<td id="76BDEA64FFCA0008FD21F90B408AEE82" box="[663,751,1761,1783]" gridcol="3" gridrow="10" pageId="24" pageNumber="25">1.8</td>
|
||
<td id="76BDEA64FFCA0008FC96F90B411DEE82" box="[800,888,1761,1783]" gridcol="4" gridrow="10" pageId="24" pageNumber="25">1.9</td>
|
||
<td id="76BDEA64FFCA0008FC1FF90B4664EE82" box="[937,1025,1761,1783]" gridcol="5" gridrow="10" pageId="24" pageNumber="25">1.6</td>
|
||
<td id="76BDEA64FFCA0008FB9DF90B46E1EE82" box="[1067,1156,1761,1783]" gridcol="6" gridrow="10" pageId="24" pageNumber="25">1.7</td>
|
||
<td id="76BDEA64FFCA0008FB1FF90B4767EE82" box="[1193,1282,1761,1783]" gridcol="7" gridrow="10" pageId="24" pageNumber="25">D</td>
|
||
<td id="76BDEA64FFCA0008FA91F90B471AEE82" box="[1319,1407,1761,1783]" gridcol="8" gridrow="10" pageId="24" pageNumber="25">0.8</td>
|
||
</tr>
|
||
<tr id="356C8318FFCA0008FF29F8E0471AEF55" box="[159,1407,1802,1824]" gridrow="11" pageId="24" pageNumber="25">
|
||
<th id="76BDEA64FFCA0008FF29F8E04336EF55" box="[159,339,1802,1824]" gridcol="0" gridrow="11" pageId="24" pageNumber="25">Casque to Eye</th>
|
||
<td id="76BDEA64FFCA0008FE33F8E04386EF55" box="[389,483,1802,1824]" gridcol="1" gridrow="11" pageId="24" pageNumber="25">10.6</td>
|
||
<td id="76BDEA64FFCA0008FDB8F8E04003EF55" box="[526,614,1802,1824]" gridcol="2" gridrow="11" pageId="24" pageNumber="25">8.1</td>
|
||
<td id="76BDEA64FFCA0008FD21F8E0408AEF55" box="[663,751,1802,1824]" gridcol="3" gridrow="11" pageId="24" pageNumber="25">5.5</td>
|
||
<td id="76BDEA64FFCA0008FC96F8E0411DEF55" box="[800,888,1802,1824]" gridcol="4" gridrow="11" pageId="24" pageNumber="25">7.3</td>
|
||
<td id="76BDEA64FFCA0008FC1FF8E04664EF55" box="[937,1025,1802,1824]" gridcol="5" gridrow="11" pageId="24" pageNumber="25">10.5</td>
|
||
<td id="76BDEA64FFCA0008FB9DF8E046E1EF55" box="[1067,1156,1802,1824]" gridcol="6" gridrow="11" pageId="24" pageNumber="25">10.7</td>
|
||
<td id="76BDEA64FFCA0008FB1FF8E04767EF55" box="[1193,1282,1802,1824]" gridcol="7" gridrow="11" pageId="24" pageNumber="25">10.7</td>
|
||
<td id="76BDEA64FFCA0008FA91F8E0471AEF55" box="[1319,1407,1802,1824]" gridcol="8" gridrow="11" pageId="24" pageNumber="25">6.5</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C346D2D1FFCAFFEEFF71F88F40F6E980" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph id="8BE3815AFFCAFFEEFF71F88F40F6E980" blockId="24.[151,1436,1893,2026]" lastBlockId="25.[151,1437,151,1979]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<emphasis id="B9285D48FFCAFFEFFF71F88F435FEF0B" bold="true" box="[199,314,1893,1918]" pageId="24" pageNumber="25">Diagnosis</emphasis>
|
||
. The Mt. Namuli pygmy chameleon is referable to the
|
||
<taxonomicName id="4C5CFAD9FFCAFFEFFC6BF88F461CEF0B" box="[989,1145,1893,1918]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9285D48FFCAFFEFFC6BF88F461CEF0B" box="[989,1145,1893,1918]" italics="true" pageId="24" pageNumber="25">Rhampholeon</emphasis>
|
||
</taxonomicName>
|
||
(subgenus
|
||
<emphasis id="B9285D48FFCAFFEFFAB2F88F47F9EF0B" box="[1284,1436,1893,1918]" italics="true" pageId="24" pageNumber="25">Rhinodigitum</emphasis>
|
||
Matthee
|
||
<emphasis id="B9285D48FFCAFFEFFF4FF860434FEFD7" box="[249,298,1929,1954]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
|
||
. 2004) by possessing a short hemipenis that is almost bag-like, acalyculate and adorned with a pair of simple apical “horns” with a variable number of thorn-like papillae arranged on the outer aspect of the horn; having an unpigmented parietal peritoneum, claws that are strongly bicuspid, smooth plantar surfaces, a rostral process, and short tail (<27% of total length in adult males). It can be distinguished from most other species in
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFF21FF564380E8A1" authority="Rhinodigitum" authorityName="Rhinodigitum" box="[151,485,188,213]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9285D48FFCBFFEEFF21FF564356E8A0" box="[151,307,188,213]" italics="true" pageId="25" pageNumber="26">Rhampholeon</emphasis>
|
||
(
|
||
<emphasis id="B9285D48FFCBFFEEFEF5FF5643BEE8A0" box="[323,475,188,213]" italics="true" pageId="25" pageNumber="26">Rhinodigitum</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
by having deep inguinal (absent or indistinct in
|
||
<emphasis id="B9285D48FFCBFFEEFBBFFF56434DE88D" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFBBFFF5646D7E8A0" box="[1033,1202,188,213]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="boulengeri">Rh. boulengeri</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFB08FF56470FE8A0" box="[1214,1386,188,213]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="nchisiensis">Rh. nchisiensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFAC0FF56434DE88D" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="uluguruensis">Rh. uluguruensis</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFEDEFF354380E88D" box="[360,485,223,248]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="moyeri">
|
||
<emphasis id="B9285D48FFCBFFEEFEDEFF354380E88D" box="[360,485,223,248]" italics="true" pageId="25" pageNumber="26">Rh. moyeri</emphasis>
|
||
</taxonomicName>
|
||
) and axillary pits (also absent in
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFCD3FF35466AE88D" box="[869,1039,223,248]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="nchisiensis">
|
||
<emphasis id="B9285D48FFCBFFEEFCD3FF35466AE88D" box="[869,1039,223,248]" italics="true" pageId="25" pageNumber="26">Rh. nchisiensis</emphasis>
|
||
</taxonomicName>
|
||
). It differs from
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFB64FF35470EE88D" box="[1234,1387,223,248]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="B9285D48FFCBFFEEFB64FF35470EE88D" box="[1234,1387,223,248]" italics="true" pageId="25" pageNumber="26">Rh. platyceps</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFF21FEEE4353E969" box="[151,310,260,285]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="maspictus" status="sp. nov.">
|
||
<emphasis id="B9285D48FFCBFFEEFF21FEEE4353E969" box="[151,310,260,285]" italics="true" pageId="25" pageNumber="26">Rh. maspictus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9285D48FFCBFFEEFEF6FEEE43F9E968" bold="true" box="[320,412,260,285]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A21BE033FFCBFFEEFEF6FEEE43F9E968" box="[320,412,260,285]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its smaller size (<
|
||
<quantity id="4CA42CBFFFCBFFEEFD20FEEE40BAE968" box="[662,735,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="25" pageNumber="26" unit="mm" value="65.0">65mm</quantity>
|
||
total length) and weak crenulations of dorsal crest. It differs from all other members of the
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFE5AFECD40E1E935" box="[492,644,295,320]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="B9285D48FFCBFFEEFE5AFECD40E1E935" box="[492,644,295,320]" italics="true" pageId="25" pageNumber="26">Rh. platyceps</emphasis>
|
||
</taxonomicName>
|
||
complex, including the populations from Mt Mabu, Mt Chiperone and Mt Inago, by retaining in adult males a prominent flexure (>32˚) of the snout in front of the orbit (flat or <10˚ in adult male
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFEA3FE8543C8E9FD" box="[277,429,367,392]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="B9285D48FFCBFFEEFEA3FE8543C8E9FD" box="[277,429,367,392]" italics="true" pageId="25" pageNumber="26">Rh. platyceps</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFE53FE8540E6E9FD" box="[485,643,367,392]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="maspictus" status="sp. nov.">
|
||
<emphasis id="B9285D48FFCBFFEEFE53FE8540E6E9FD" box="[485,643,367,392]" italics="true" pageId="25" pageNumber="26">Rh. maspictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21BE033FFCBFFEEFD3DFE854080E9FD" box="[651,741,367,392]" pageId="25" pageNumber="26" rank="species">
|
||
<emphasis id="B9285D48FFCBFFEEFD3DFE854080E9FD" bold="true" box="[651,741,367,392]" pageId="25" pageNumber="26">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; <18% in the only known adult males of
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFB0BFE854717E9FD" box="[1213,1394,367,392]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="nebulauctor" status="sp. nov.">
|
||
<emphasis id="B9285D48FFCBFFEEFB0BFE854717E9FD" box="[1213,1394,367,392]" italics="true" pageId="25" pageNumber="26">Rh. nebulauctor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9285D48FFCBFFEEFACCFE8542ADE9D8" bold="true" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A21BE033FFCBFFEEFACCFE8542ADE9D8" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFEB4FE7E43B3E9D8" box="[258,470,404,429]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="bruessoworum" status="sp. nov.">
|
||
<emphasis id="B9285D48FFCBFFEEFEB4FE7E43B3E9D8" box="[258,470,404,429]" italics="true" pageId="25" pageNumber="26">Rh. bruessoworum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9285D48FFCBFFEEFE69FE7E405FE9D8" bold="true" box="[479,570,404,429]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A21BE033FFCBFFEEFE69FE7E405FE9D8" box="[479,570,404,429]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), and a narrower head (HW/HL% 49.4%; 53.2–59.8% in all other species). Finally, the species is also genetically well differentiated from all other
|
||
<taxonomicName id="4C5CFAD9FFCBFFEEFC0EFE5D4631E9A5" box="[952,1108,439,464]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9285D48FFCBFFEEFC0EFE5D4631E9A5" box="[952,1108,439,464]" italics="true" pageId="25" pageNumber="26">Rhampholeon</emphasis>
|
||
</taxonomicName>
|
||
, and all specimens examined form a monophyletic clade from Mt. Namuli.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D2D1FFCBFFECFF71FE154050EB60" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="8BE3815AFFCBFFEEFF71FE154342EA48" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
|
||
<emphasis id="B9285D48FFCBFFEEFF71FE154380EA6D" bold="true" box="[199,485,511,536]" pageId="25" pageNumber="26">
|
||
Description of
|
||
<typeStatus id="54E73FF8FFCBFFEEFECEFE154380EA6D" box="[376,485,511,536]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
. Adult female, viscera exposed by a single ventral incision and two lateral incisions on left flank.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71FDAD4306EC66" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
|
||
Head: Flattened, with only a small upward flexure of the snout at the front of the orbit (angling upwards at approximately 28˚ from a line between tip of snout and top of casque); casque weak, flat with small, conical tubercular edges; temporal crest moderate, forming a straight ridge of 7 raised, rounded tubercles, 3rd and last largest; lateral crests weak, composed of small conical tubercles that demarcate the edge of the casque; temporal crest moderate with a single row of interrupted large tubercles; parietal crest almost absent, composed of a few enlarged tubercles in the mid-line; supraorbital ridge well marked but without obvious clusters of tubercles (vaguely present at the posterior edge), and with a very small ‘soft horn’ of enlarged tubercles at the junction with the shallow interorbital ridge; interorbital ridge composed of 12 small granular tubercles that demarcate the posterior edge of a prominent frontal depression; two enlarged tubercles on inferior orbital rim; rostral ridges very well marked, and fusing at tip of the snout which bears a rostral process (
|
||
<quantity id="4CA42CBFFFCBFFEEFC43FC644634EBD2" box="[1013,1105,910,935]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, in fixation flattened against rostrum), about 7 small granules long and 5-6 granules wide at base; nares opening posterio-ventrally; no gular or mental appendages; scales on throat homogenous, more stellate than those on crown of head and subequal in size to those on the belly.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71FBF446A8ED46" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">Body: Dorsal crest weak, composed of a double row of enlarged, rounded tubercles along the backbone; the crest undulates between 13 equally spaced clusters of enlarged, spinose tubercles that are largest on the middle of the body, flatter and smooth over the forelimb and sacral regions, and only vaguely visible on the tail; deep axillary and inguinal pits are present; flank scalation heterogenous, composed of small, stellate granules with scattered, enlarged spinose tubercles, the largest occurring over the shoulder; chest, belly and lower surface of tail smooth; limb scalation more irregular, with numerous larger, spinose tubercles on forearms; soles of feet sub-spinose; claws strongly bicuspid; accessory planter spines at base of claws very small, almost absent; tail gently tapering, dorsoventrally flattened, 21.7% of total length, and covered with homogenous conical tubercles.</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71FAD44284EDEA" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">Colour in preservative: Body mottled brown with reddish brown reticulation on flanks; head darker above, blotched with dark brown; belly and inner surfaces of limbs paler with darker specking; tail dark brown above and below.</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71FA404604ED92" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
|
||
<emphasis id="B9285D48FFCBFFEEFF71FA404381EDB6" bold="true" box="[199,484,1450,1475]" pageId="25" pageNumber="26">Description of Allotype</emphasis>
|
||
(as for
|
||
<typeStatus id="54E73FF8FFCBFFEEFDFEFA4040C8EDB6" box="[584,685,1450,1475]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
, unless noted). Adult male, with everted hemipenes and viscera exposed by a single, ragged ventral incision and liver tissue excised for DNA analysis.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71FA18471AEE02" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
|
||
Head: A very prominent angular flexure of the snout at the front of the orbit (approx 40˚ from line between tip of snout and casque); supraorbital ridge well marked with clusters of tubercles at posterior and anterior level of orbit, the latter forming a ‘soft horn’ two scales high; rostral ridge fusing anteriorly with an obvious rostral process (
|
||
<quantity id="4CA42CBFFFCBFFEEFF29F9B5429FEE02" box="[159,250,1631,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="26" unit="mm" value="1.3">1.3 mm</quantity>
|
||
), that is rounded in profile and composed of about 6 small granules long and 5-6 granules wide at base.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71F9684378EECA" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">Body: Claws strongly bicuspid; accessory planter spines at the base of claws weakly developed; tail 24.0% of total length.</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71F92046FFEF06" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">
|
||
Coloration in life (
|
||
<figureCitation id="13679DDFFFCBFFEEFE2DF92043BBEE96" box="[411,478,1738,1763]" captionStart="FIGURE 8" captionStartId="26.[151,250,1908,1930]" captionTargetBox="[169,1421,211,1859]" captionTargetId="figure@26.[151,1436,193,1887]" captionTargetPageId="26" captionText="FIGURE 8. Rhampholeon tiburyi sp. nov.: A—Allotype (PEM R 17132, K. A. Tolley); B—Paratype (PEM R 17134, K Tolley); C—Head of male paratype (PEM R 20372) showing snout flexure; D—Preserved female holotype (PEM R 14921); E—Manho Forest, Mt. Namuli (J. Bayliss)." httpUri="https://zenodo.org/record/286219/files/figure.png" pageId="25" pageNumber="26">Fig. 8</figureCitation>
|
||
A): Head, body and tail tan to brown with orange flush along midflank and greenish flush over back and outer surfaces of hind limbs; two prominent diagonal lines on flanks, formed from three darkish spots joined by grey infusion; belly, throat, inner surfaces of limbs and underside of tail light tan; outer skin of limbs darker brown; snout with wide orange band running diagonally through nostril; skin of orbit and temporal region darker brown with scattered, intense green-blue individual tubercles; iris orange-red.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFCBFFEEFF71F89443F0EFCE" blockId="25.[151,1437,151,1979]" pageId="25" pageNumber="26">Coloration after preservation: Body very dark, almost blackish; all normal coloration lost due to fixation in concentrated formalin.</paragraph>
|
||
<caption id="DF23D1D2FFC8FFEDFF21F89E43BDEFBD" httpUri="https://zenodo.org/record/286219/files/figure.png" pageId="26" pageNumber="27" targetBox="[169,1421,211,1859]" targetPageId="26">
|
||
<paragraph id="8BE3815AFFC8FFEDFF21F89E43BDEFBD" blockId="26.[151,1436,1908,1992]" pageId="26" pageNumber="27">
|
||
<emphasis id="B9285D48FFC8FFEDFF21F89E4377EFFF" bold="true" box="[151,274,1908,1930]" pageId="26" pageNumber="27">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C5CFAD9FFC8FFEDFEAEF89E438AEFFF" box="[280,495,1908,1930]" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="tiburyi" status="sp. nov.">
|
||
<emphasis id="B9285D48FFC8FFEDFEAEF89E438AEFFF" box="[280,495,1908,1930]" italics="true" pageId="26" pageNumber="27">Rhampholeon tiburyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9285D48FFC8FFEDFE42F89E4021EFFF" bold="true" box="[500,580,1908,1930]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A21BE033FFC8FFEDFE42F89E4021EFFF" box="[500,580,1908,1930]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A—Allotype (PEM R17132, K.A. Tolley); B—Paratype (PEM R17134, K Tolley); C—Head of male paratype (PEM R20372) showing snout flexure; D—Preserved female holotype (PEM R14921); E—Manho Forest, Mt. Namuli (J. Bayliss).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FF7D4681E935" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FF7D43D6E8C5" bold="true" box="[199,435,151,176]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54E73FF8FFC9FFECFF71FF7D4351E8C5" box="[199,308,151,176]" pageId="27" pageNumber="28" type="paratype">Paratype</typeStatus>
|
||
variation.
|
||
</emphasis>
|
||
The tail varies from 18.3–21.7% of total length in females, and from 24.0–25.8% in males; the rostral process varies from 7.9–10.4% of head length in females, and from 15.9–16.6% in males. In life the iris of a male
|
||
<typeStatus id="54E73FF8FFC9FFECFEEEFF0A43DDE88D" box="[344,440,224,248]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
(R17131) was reddish and the snout had an indistinct brown stripe rather than the orange band in the allotype. In addition, a few scattered green-blue tubercles extended onto the snout, as well as on to the orbit and temporal region. Prominent “leaf vein” dark lines on the flank were also well developed.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FEA64112E9D8" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FEA6429BE910" bold="true" box="[199,254,332,357]" pageId="27" pageNumber="28">Size.</emphasis>
|
||
Largest male—PEM R17132 (allotype) 39.6 + 12.5 =
|
||
<quantity id="4CA42CBFFFC9FFECFCE8FEA641A0E911" box="[862,965,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.21" pageId="27" pageNumber="28" unit="mm" value="52.1">52.1 mm</quantity>
|
||
; largest female—PEM R14921 (
|
||
<typeStatus id="54E73FF8FFC9FFECFA99FEA647F1E910" box="[1327,1428,332,357]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
) 55.3 + 15.4 =
|
||
<quantity id="4CA42CBFFFC9FFECFE83FE9A43F9E9F2" box="[309,412,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="7.07" pageId="27" pageNumber="28" unit="mm" value="70.7">70.7 mm</quantity>
|
||
. The smallest male (SVL
|
||
<quantity id="4CA42CBFFFC9FFECFD09FE9A4140E9FD" box="[703,805,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.32" pageId="27" pageNumber="28" unit="mm" value="33.2">33.2 mm</quantity>
|
||
) still has well-developed hemipenes, indicating that it is sexually mature (although testicular activity was not determined).
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FE5D467BEAF0" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FE5D43D9E9A5" bold="true" box="[199,444,439,464]" pageId="27" pageNumber="28">Sexual Dimorphism.</emphasis>
|
||
Whilst the
|
||
<typeStatus id="54E73FF8FFC9FFECFDF6FE524015E9A5" box="[576,624,440,464]" pageId="27" pageNumber="28">type</typeStatus>
|
||
series is limited in number, there are indications of sexual dimorphism. Females grow considerably larger than males, and the three females in the
|
||
<typeStatus id="54E73FF8FFC9FFECFC48FE37464BE980" box="[1022,1070,477,501]" pageId="27" pageNumber="28">type</typeStatus>
|
||
series have an average SVL of
|
||
<quantity id="4CA42CBFFFC9FFECFF21FE154369EA62" box="[151,268,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="5.167" pageId="27" pageNumber="28" unit="mm" value="51.67">51.67 mm</quantity>
|
||
, compared with only
|
||
<quantity id="4CA42CBFFFC9FFECFDB4FE154011EA62" box="[514,628,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.467" pageId="27" pageNumber="28" unit="mm" value="34.67">34.67 mm</quantity>
|
||
for the three males. In addition, males have a more angular head shape (slope 39-45˚,
|
||
<figureCitation id="13679DDFFFC9FFECFEF2FDCE43E6EA49" box="[324,387,548,573]" captionStart="FIGURE 9" captionStartId="28.[151,250,1953,1975]" captionTargetBox="[177,1407,214,1913]" captionTargetId="figure@28.[164,1423,193,1932]" captionTargetPageId="28" captionText="FIGURE 9. Rhampholeon bruessoworum sp. nov.: A—Holotype; B—Allotype; C—Habitat on Mt. Inago (J. Bayliss)." httpUri="https://zenodo.org/record/286220/files/figure.png" pageId="27" pageNumber="28">Fig 9</figureCitation>
|
||
B) than females (slope 22-28˚), and also proportionately longer tails and a longer rostral process (see above). However, the small number of specimens precludes statistical confirmation. More subtle difference in scalation details cannot be assessed until larger series are available.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FD654050EB60" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FD65432BEADD" bold="true" box="[199,334,655,680]" pageId="27" pageNumber="28">Hemipenis.</emphasis>
|
||
(based on everted hemipenes of all three males in
|
||
<typeStatus id="54E73FF8FFC9FFECFC21FD7A41A2EADD" box="[919,967,656,680]" pageId="27" pageNumber="28">type</typeStatus>
|
||
series). Hemipenis short with unadorned basal and apical sections, except for a pair of curved horns that emerge prominently from the expanded, flattened crown, and which bear well developed spines on their upper surfaces; the folds of the sulcus spermaticus flare to drain into the flattened apical region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346D2D1FFC9FFECFF71FCF54345EE7D" pageId="27" pageNumber="28" type="distribution">
|
||
<paragraph id="8BE3815AFFC9FFECFF71FCF546DBEBF5" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FCF54305EB4D" bold="true" box="[199,352,799,824]" pageId="27" pageNumber="28">Distribution.</emphasis>
|
||
Restricted to the
|
||
<typeStatus id="54E73FF8FFC9FFECFD9CFCCA403FEB4D" box="[554,602,800,824]" pageId="27" pageNumber="28">type</typeStatus>
|
||
locality; Mt. Namuli, Zambézia Province, northern
|
||
<collectingCountry id="F34BC1CAFFC9FFECFB1FFCF54725EB4D" box="[1193,1344,799,824]" name="Mozambique" pageId="27" pageNumber="28">Mozambique</collectingCountry>
|
||
, in both the evergreen forests on Manho and Ukalini forests on the Muretha Plateau (
|
||
<quantity id="4CA42CBFFFC9FFECFC4EFCAE462AEB29" box="[1016,1103,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.8039999999999998" pageId="27" pageNumber="28" unit="m" value="1804.0">1804 m</quantity>
|
||
a.s.l.) at the base of the main peaks, respectively, and also in fallow areas of tea plantations at lower elevations (
|
||
<quantity id="4CA42CBFFFC9FFECFB84FC82461FEB0A" box="[1074,1146,872,896]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="27" pageNumber="28" unit="m" value="838.0">838 m</quantity>
|
||
a.s.l).
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FC66434AEC08" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FC66434FEBD0" bold="true" box="[199,298,908,933]" pageId="27" pageNumber="28">Habitat.</emphasis>
|
||
The Namuli Massif shows habitat zonation and is surrounded by
|
||
<taxonomicName id="4C5CFAD9FFC9FFECFBBBFC6646C4EBD0" box="[1037,1185,908,933]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9285D48FFC9FFECFBBBFC6646C4EBD0" box="[1037,1185,908,933]" italics="true" pageId="27" pageNumber="28">Brachystegia</emphasis>
|
||
</taxonomicName>
|
||
woodland at the base. Grassland and scrub with forested river valleys cover the slopes, with grassy plateaus and patches of dense moist evergreen forest near the summit (
|
||
<figureCitation id="13679DDFFFC9FFECFDAFFC3E4039EB99" box="[537,604,980,1005]" captionStart="FIGURE 8" captionStartId="26.[151,250,1908,1930]" captionTargetBox="[169,1421,211,1859]" captionTargetId="figure@26.[151,1436,193,1887]" captionTargetPageId="26" captionText="FIGURE 8. Rhampholeon tiburyi sp. nov.: A—Allotype (PEM R 17132, K. A. Tolley); B—Paratype (PEM R 17134, K Tolley); C—Head of male paratype (PEM R 20372) showing snout flexure; D—Preserved female holotype (PEM R 14921); E—Manho Forest, Mt. Namuli (J. Bayliss)." httpUri="https://zenodo.org/record/286219/files/figure.png" pageId="27" pageNumber="28">Fig. 8</figureCitation>
|
||
E). The largest surviving blocks include the Manho Forest (c.1,000–1,100 ha) and the Ukalini Forest (c.100 ha), the latter lodged against the base of the Namuli dome. Dominant emergent trees include
|
||
<emphasis id="B9285D48FFC9FFECFE83FBF7415BEC40" box="[309,830,1052,1077]" italics="true" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C5CFAD9FFC9FFECFE83FBF740C8EC41" authority="Faurea" authorityName="Faurea" box="[309,685,1052,1077]" class="Magnoliopsida" family="Lauraceae" genus="Cryptocarya" kingdom="Plantae" order="Laurales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="liebertiana">Cryptocarya liebertiana, Faurea</taxonomicName>
|
||
wentzeliana
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C5CFAD9FFC9FFECFCCBFBF6472FEC41" authority="Timberlake et al. 2009" authorityName="Timberlake et al." authorityYear="2009" box="[893,1354,1052,1077]" class="Gastropoda" family="Limapontiidae" genus="Olea" kingdom="Animalia" order="Sacoglossa" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="capensis">
|
||
<emphasis id="B9285D48FFC9FFECFCCBFBF6467AEC41" box="[893,1055,1052,1077]" italics="true" pageId="27" pageNumber="28">Olea capensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCDFCABFFC9FFECFB84FBF64727EC41" author="Timberlake" box="[1074,1346,1052,1077]" pageId="27" pageNumber="28" refString="Timberlake, J. R., F. Dowsett-Lemaire, F., Bayliss, J., Alves, T., Baena, S., Bento, C., Cook, K., Francisco, J., Harris, T., Smith, P. & de Sousa, C. (2009) Mt. Namuli, Mozambique: Biodiversity and Conservation. Report produced under the Darwin Initiative Award 15 / 036. Royal Botanic Gardens, Kew, London, 114 pp." type="book" year="2009">Timberlake et al. 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. These forests are under high anthropogenic threat as they are cleared for potato cultivation by surrounding rural communities.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FB6D438EED45" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
The allotype and most
|
||
<typeStatus id="54E73FF8FFC9FFECFE65FB63405BECD4" box="[467,574,1161,1185]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
|
||
were collected at night on perches from
|
||
<quantity id="4CA42CBFFFC9FFECFBA9FB6D46E9ECD5" box="[1055,1164,1159,1184]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="27" pageNumber="28" unit="cm" value="40.0" valueMax="50.0" valueMin="30.0">30-50 cm</quantity>
|
||
above ground in forest adjacent to a mountain stream in a fallow areas of an operational tea estate that have overgrown into a forest.
|
||
<bibRefCitation id="EFCDFCABFFC9FFECFAEEFB464341EC9D" pageId="27" pageNumber="28" refString="Portik, D. M., Travers, S., Bauer, A. M. & Branch, W. R. (2013 b) A new species of Lygodactylus (Squamata: Gekkonidae) endemic to Mount Namuli, an isolated ' sky' island of northern Mozambique. Zootaxa, 3710 (5), 415 - 435. http: // dx. doi. org / 10.11646 / zootaxa. 3710.5.2" type="journal article">
|
||
Portik
|
||
<emphasis id="B9285D48FFC9FFECFF21FB3B42ACEC9D" box="[151,201,1231,1256]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
noted that adults and juveniles were found sheltering on branches of small trees or on the fronds of epiphytic ferns within Ukalini Forest. This contrasts with perches heights of
|
||
<quantity id="4CA42CBFFFC9FFECFBEAFB1E46A7ED79" box="[1116,1218,1268,1293]" metricMagnitude="0" metricUnit="m" metricValue="3.15" metricValueMax="5.0" metricValueMin="1.3" pageId="27" pageNumber="28" unit="m" value="3.15" valueMax="5.0" valueMin="1.3">1.3–5 m</quantity>
|
||
recorded for
|
||
<taxonomicName id="4C5CFAD9FFC9FFECFAC0FB1E4382ED45" authority="Tilbury 1992" authorityName="Tilbury" authorityYear="1992" class="Reptilia" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="chapmanorum">
|
||
<emphasis id="B9285D48FFC9FFECFAC0FB1E435CED45" italics="true" pageId="27" pageNumber="28">Rh. chapmanorum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCDFCABFFC9FFECFEF0FAFD43BAED45" author="Tilbury" box="[326,479,1303,1328]" pageId="27" pageNumber="28" refString="Tilbury, C. R. (1992) A new dwarf forest chameleon (Sauria: Rhampholeon) from Malawi, Central Africa. Tropical Zoology, 5, 1 - 9. http: // dx. doi. org / 10.1080 / 03946975.1992.10539176" type="journal article" year="1992">Tilbury 1992</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FAD64700ED0D" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FAD64317ED20" bold="true" box="[199,370,1340,1365]" pageId="27" pageNumber="28">Reproduction.</emphasis>
|
||
The female
|
||
<typeStatus id="54E73FF8FFC9FFECFDBEFAD6400FED20" box="[520,618,1340,1365]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
was gravid with seven almost spherical eggs (largest diameter
|
||
<quantity id="4CA42CBFFFC9FFECFAFBFAD642A0ED0D" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="27" pageNumber="28" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.1-4.5 mm</quantity>
|
||
, four in the left oviduct and three in the right oviduct) that lack obvious signs of embryonic development.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FA6E46AFEDE8" blockId="27.[151,1437,151,1545]" box="[199,1226,1412,1437]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FA6E4365EDE8" bold="true" box="[199,256,1412,1437]" pageId="27" pageNumber="28">Diet.</emphasis>
|
||
The stomach of the
|
||
<typeStatus id="54E73FF8FFC9FFECFE53FA6E4022EDE8" box="[485,583,1412,1437]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
contained insect fragments, including coleopteran elytra.
|
||
</paragraph>
|
||
<paragraph id="8BE3815AFFC9FFECFF71FA4D4345EE7D" blockId="27.[151,1437,151,1545]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9285D48FFC9FFECFF71FA4D4321EDB5" bold="true" box="[199,324,1447,1472]" pageId="27" pageNumber="28">Predation.</emphasis>
|
||
A small chameleon, possibly referable to this species, was seen being carried by a Crowned Hornbill (
|
||
<taxonomicName id="4C5CFAD9FFC9FFECFEB0FA264066ED90" box="[262,515,1484,1509]" class="Aves" family="Bucerotidae" genus="Tockus" kingdom="Animalia" order="Bucerotiformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="alboterminatus">
|
||
<emphasis id="B9285D48FFC9FFECFEB0FA264066ED90" box="[262,515,1484,1509]" italics="true" pageId="27" pageNumber="28">Tockus alboterminatus</emphasis>
|
||
</taxonomicName>
|
||
) that was flying over the canopy of Ukalini Forest,
|
||
<date id="FFE2A79AFFC9FFECFBF1FA26477BED91" box="[1095,1310,1484,1509]" pageId="27" pageNumber="28" value="1998-11-29">29 November 1998</date>
|
||
(
|
||
<bibRefCitation id="EFCDFCABFFC9FFECFA9BFA264371EE7D" author="Branch" pageId="27" pageNumber="28" refString="Branch, W. R. & Ryan, P. G. (2001) Additions to the Mozambique Herpetofauna: Two new lizards from the Namuli Massif, Mozambique. Herpetological Review, 32 (4), 281 - 282." type="journal article" year="2001">Branch & Ryan 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |