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<document id="9F4AF0F287CDB93D0C688F90C6A9770F" ID-CLB-Dataset="202625" ID-DOI="10.11646/zootaxa.5214.4.1" ID-GBIF-Dataset="05dc98be-c3f9-4039-a21d-b285d6059a8c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7397617" ID-ZooBank="80EE5BF7-A183-4089-950E-25DBD65EFCFC" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1670225684006" checkinUser="plazi" docAuthor="Carpenter, James M. &amp; Brown, Graham R." docDate="2022" docId="036C87E0B124FF80D3CF57A8614EC38B" docLanguage="en" docName="zootaxa.5214.4.1.pdf" docOrigin="Zootaxa 5214 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ropalidia impetuosa (Smith)" docType="treatment" docVersion="11" lastPageNumber="463" masterDocId="FF55FF98B12FFF8CD358516F6562C456" masterDocTitle="Catalogue of the Australian social wasps (Hymenoptera: Vespidae)" masterLastPageNumber="495" masterPageNumber="451" pageNumber="462" updateTime="1699412786959" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EFF4E0B40AADD9171BF548034FEA1FFF">Catalogue of the Australian social wasps (Hymenoptera: Vespidae)</mods:title>
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<mods:namePart id="EEBF293C9F567DE82D2EA75A4100D691">Carpenter, James M.</mods:namePart>
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<mods:affiliation id="C1482D70C3C789B5ED73DB7A8F275FE2">Museum and Art Gallery of the Northern Territory, GPO Box 4646, Darwin, NT 0801, Australia. &amp; Visiting Scientist, CSIRO Land and Water, Private Mail Bag 44, Winnellie, NT 0822, Australia. &amp; University Fellow, Research Institute for Environment and Livelihoods, Charles Darwin University, Darwin, NT 0909, Australia. thynnini @ outlook. com; https: // orcid. org / 0000 - 0002 - 4101 - 3823</mods:affiliation>
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<paragraph id="8B7A36F6B124FF87D3CF57A86405C2B7" blockId="11.[151,1436,1735,2013]" box="[151,359,1735,1761]" pageId="11" pageNumber="462">
<taxonomicName id="4CC54D75B124FF87D3CF57A86405C2B7" ID-CoL="4TBXP" authority="(Smith)" authorityName="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[151,359,1735,1761]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B124FF87D3CF57A86468C2B6" box="[151,266,1735,1760]" italics="true" pageId="11" pageNumber="462">impetuosa</emphasis>
(Smith)
</taxonomicName>
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<paragraph id="8B7A36F6B124FF87D3AF57846620C3C3" blockId="11.[151,1436,1735,2013]" pageId="11" pageNumber="462">
<taxonomicName id="4CC54D75B124FF87D3AF57846735C353" authority="Smith, 1860" authorityName="Smith" authorityYear="1860" baseAuthorityName="Smith" box="[247,599,1771,1797]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B124FF87D3AF578464D9C352" box="[247,443,1771,1796]" italics="true" pageId="11" pageNumber="462">Icaria impetuosa</emphasis>
<bibRefCitation id="EF544B07B124FF87D29057846735C353" author="Smith" box="[456,599,1771,1797]" firstAuthor="Smith" pageId="11" pageNumber="462" pagination="93 - 143" refId="ref49541" refString="Smith, F. (1860 [&quot; 1861 &quot;]) Descriptions of hymenopterous insects collected by Mr. A. R. Wallace in the islands of Batchian, Kaisaa, Amboyna, Gilolo and at Dory in New Guinea. Journal of the Proceedings of the Linnean Society, Zoology, 5, 93 - 143. https: // doi. org / 10.1111 / j. 1096 - 3642.1860. tb 01022. x" type="journal article" year="1860">Smith, 1860</bibRefCitation>
</taxonomicName>
[1861]: 131, female - “Batchian; Amboyna” (
<typeStatus id="547E8854B124FF87D7D85784618BC353" box="[1152,1257,1771,1797]" pageId="11" pageNumber="462" type="lectotype">lectotype</typeStatus>
Oxford); 1863 [1864]a: 41 (distr.); 1863 [1864]b: 138 (distr.); 1871: 380 (cat.). -
<bibRefCitation id="EF544B07B124FF87D71656606033C37F" author="Dalla Torre" box="[1102,1361,1807,1833]" firstAuthor="Dalla Torre" page="118" pageId="11" pageNumber="462" refId="ref30910" refString="Dalla Torre, K. W. (1894) Catalogus Hymenopterorum. Vol. 9. Vespidae (Diploptera). G. Engelmann, Leipzig, 181 pp." type="book" year="1894">Dalla Torre, 1894: 118</bibRefCitation>
(cat.); 1904: 73 (cat.). -
<bibRefCitation id="EF544B07B124FF87D141565C6653C31B" author="Vecht, J. van der" box="[537,817,1843,1869]" firstAuthor="van der Vecht" page="104" pageId="11" pageNumber="462" pagination="103 - 190" refId="ref51729" refString="Vecht, J. van der (1941) The Indo-Australian species of the genus Ropalidia (= Icaria) (Hym., Vespidae) (first part). Treubia, 18, 103 - 190." type="journal article" year="1941">van der Vecht, 1941: 104</bibRefCitation>
(syn. of
<taxonomicName id="4CC54D75B124FF87D0CC565C61DDC31B" ID-CoL="4TBXH" authority="(de Saussure))" authorityName="" baseAuthorityName="de Saussure" baseAuthorityYear="1854" box="[916,1215,1843,1869]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="B9B1EAE4B124FF87D0CC565C6176C31A" box="[916,1044,1843,1868]" italics="true" pageId="11" pageNumber="462">R. gregaria</emphasis>
(de Saussure))
</taxonomicName>
. -
<bibRefCitation id="EF544B07B124FF87D781565C60FEC31B" author="Kojima" box="[1241,1436,1843,1869]" firstAuthor="Kojima" page="9" pageId="11" pageNumber="462" pagination="1 - 22" refId="ref40107" refString="Kojima, J. (2001 a) Taxonomic notes on types of the social wasp tribe Ropalidiini (Insecta: Hymenoptera; Vespidae, Polistinae) housed mainly in the Museum d'Histoire Naturelle, Geneve. Natural History Bulletin of Ibaraki University, 5, 1 - 22." type="journal article" year="2001" yearSuffix="a">Kojima, 2001a: 9</bibRefCitation>
(syns.:
<taxonomicName id="4CC54D75B124FF87D2F0563867DDC327" ID-CoL="3P8LM" authority="Cameron" authorityName="Cameron" box="[424,703,1879,1905]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="spilocephala">
<emphasis id="B9B1EAE4B124FF87D2F05638672CC327" box="[424,590,1879,1905]" italics="true" pageId="11" pageNumber="462">I. spilocephala</emphasis>
Cameron
</taxonomicName>
,
<taxonomicName id="4CC54D75B124FF87D192563866C7C327" ID-CoL="3P8KL" authority="du Buysson" authorityName="du Buysson" box="[714,933,1879,1905]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="cohni">
<emphasis id="B9B1EAE4B124FF87D1925638667AC327" box="[714,792,1879,1905]" italics="true" pageId="11" pageNumber="462">I cohni</emphasis>
du Buysson
</taxonomicName>
,
<taxonomicName id="4CC54D75B124FF87D0E856386070C327" ID-CoL="4TBXT" authority="Cheesman" authorityName="Cheesman" box="[944,1298,1879,1905]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="inquieta">
<emphasis id="B9B1EAE4B124FF87D0E8563861F1C326" box="[944,1171,1879,1904]" italics="true" pageId="11" pageNumber="462">R. gregaria inquieta</emphasis>
Cheesman
</taxonomicName>
,
<taxonomicName id="4CC54D75B124FF87D64456386700C3C3" ID-CoL="4TC3B" authority="Cheesman" authorityName="Cheesman" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="variety" species="gregaria" variety="tolerans">
<emphasis id="B9B1EAE4B124FF87D644563860FEC326" box="[1308,1436,1879,1904]" italics="true" pageId="11" pageNumber="462">R. gregaria</emphasis>
var.
<emphasis id="B9B1EAE4B124FF87D2DF56146481C3C3" box="[391,483,1915,1941]" italics="true" pageId="11" pageNumber="462">tolerans</emphasis>
Cheesman
</taxonomicName>
,
<taxonomicName id="4CC54D75B124FF87D1355614678FC3C2" ID-CoL="4TBXH" authorityName="" baseAuthorityName="de Saussure" baseAuthorityYear="1854" box="[621,749,1915,1940]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="B9B1EAE4B124FF87D1355614678FC3C2" box="[621,749,1915,1940]" italics="true" pageId="11" pageNumber="462">R. gregaria</emphasis>
</taxonomicName>
auctt.).
</paragraph>
<paragraph id="8B7A36F6B124FF80D3AF56F061A3C483" blockId="11.[151,1436,1735,2013]" lastBlockId="12.[247,1437,151,2013]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
<treatmentCitationGroup id="ABD511D8B124FF80D3AF56F061A3C483" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
<treatmentCitation id="0A6410E7B124FF87D3AF56F067BEC3EF" author="Cameron, P." box="[247,732,1951,1977]" page="230" pageId="11" pageNumber="462" year="1906">
<taxonomicName id="4CC54D75B124FF87D3AF56F067BEC3EF" ID-CoL="3P8LM" authority="Cameron, 1906 b: 230" authorityName="Cameron" authorityPageNumber="230" authorityYear="1906" box="[247,732,1951,1977]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="spilocephala">
<emphasis id="B9B1EAE4B124FF87D3AF56F064B7C3EF" box="[247,469,1951,1977]" italics="true" pageId="11" pageNumber="462">Icaria spilocephala</emphasis>
<bibRefCitation id="EF544B07B124FF87D28756F067BEC3EF" author="Cameron" box="[479,732,1951,1977]" firstAuthor="Cameron" page="230" pageId="11" pageNumber="462" pagination="221 - 233" refId="ref28309" refString="Cameron, P. (1906 b) Hymenoptera of the Dutch Expedition to New Guinea in 1904 and 1905. Part. 1: Thynnidae, Scolidae, Pompilidae, Sphegidae and Vespidae. Tijdschrift voor Entomologie, 49, 221 - 233." type="journal article" year="1906" yearSuffix="b">Cameron, 1906b: 230</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, female - [New
<collectingCountry id="F3D27666B124FF87D0C556F06697C3EF" box="[925,1013,1951,1977]" name="Guinea-Bissau" pageId="11" pageNumber="462">Guinea</collectingCountry>
:] “Etna Bay” (
<typeStatus id="547E8854B124FF87D7C756F0606AC3EF" box="[1183,1288,1951,1977]" pageId="11" pageNumber="462" type="lectotype">lectotype</typeStatus>
Naturalis). -
<bibRefCitation id="EF544B07B124FF87D20F56AC6719C38B" author="Vecht, J. van der" box="[343,635,1987,2013]" firstAuthor="van der Vecht" page="104" pageId="11" pageNumber="462" pagination="103 - 190" refId="ref51729" refString="Vecht, J. van der (1941) The Indo-Australian species of the genus Ropalidia (= Icaria) (Hym., Vespidae) (first part). Treubia, 18, 103 - 190." type="journal article" year="1941">
van der
<treatmentCitation id="0A6410E7B124FF87D2EF56AC6719C38B" author="Vecht, J. van der" box="[439,635,1987,2013]" page="104" pageId="11" pageNumber="462" year="1941">Vecht, 1941: 104</treatmentCitation>
</bibRefCitation>
(syn. of
<taxonomicName id="4CC54D75B124FF87D1BF56AC617EC38B" ID-CoL="4TBXH" authority="(de Saussure))" authorityName="" baseAuthorityName="de Saussure" baseAuthorityYear="1854" box="[743,1052,1987,2013]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="B9B1EAE4B124FF87D1BF56AC6608C38A" box="[743,874,1987,2012]" italics="true" pageId="11" pageNumber="462">R. gregaria</emphasis>
(de Saussure))
</taxonomicName>
. -
<treatmentCitation id="0A6410E7B124FF87D76456AC60FEC38B" author="Kojima, J. &amp; Carpenter, J. M." box="[1084,1436,1986,2013]" page="5" pageId="11" pageNumber="462" year="1997">
<bibRefCitation id="EF544B07B124FF87D76456AC60FEC38B" author="Kojima &amp; Carpenter" box="[1084,1436,1986,2013]" firstAuthor="Kojima" page="5" pageId="11" pageNumber="462" pagination="1 - 96" refId="ref40310" refString="Kojima, J. &amp; Carpenter, J. M. (1997 a) Catalog of species in the polistine tribe Ropalidiini (Hymenoptera: Vespidae). American Museum Novitates, 3199, 1 - 96." type="journal article" year="1997" yearSuffix="a">Kojima &amp; Carpenter, 1997a: 5</bibRefCitation>
</treatmentCitation>
(designation of
<typeStatus id="547E8854B123FF80D15251F86714C4E7" box="[522,630,151,177]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
). -
<treatmentCitation id="0A6410E7B123FF80D1C451F86130C4E7" author="Kojima, J. &amp; van Achterberg, C." box="[668,1106,151,177]" page="166" pageId="12" pageNumber="463" year="1997">
<bibRefCitation id="EF544B07B123FF80D1C451F86130C4E7" author="Kojima &amp; van Achterberg" box="[668,1106,151,177]" firstAuthor="Kojima" page="166" pageId="12" pageNumber="463" pagination="157 - 169" refId="ref40201" refString="Kojima, J. &amp; van Achterberg, C. (1997 a) List of the primary types of social wasps (Hymenoptera: Vespidae) deposited in the Nationaal Natuurhistorisch Museum, Leiden, and the Zoologisch Museum, Amsterdam. Zoologische mededeelingen, 71 (14), 157 - 169." type="journal article" year="1997" yearSuffix="a">Kojima &amp; van Achterberg, 1997a: 166</bibRefCitation>
</treatmentCitation>
(list). -
<treatmentCitation id="0A6410E7B123FF80D7E951F860FAC4E7" author="Kojima, J." box="[1201,1432,151,177]" page="367" pageId="12" pageNumber="463" year="1999">
<bibRefCitation id="EF544B07B123FF80D7E951F860FAC4E7" author="Kojima" box="[1201,1432,151,177]" firstAuthor="Kojima" page="367" pageId="12" pageNumber="463" pagination="367 - 377" refId="ref39867" refString="Kojima, J. (1999 b) Taxonomic notes on Australian Ropalidia Guerin-Meneville, 1831 (Hymenoptera: Vespidae, Polistinae). Entomological Science, 2, 367 - 377." type="journal article" year="1999" yearSuffix="b">Kojima, 1999b: 367</bibRefCitation>
</treatmentCitation>
, 371 (syn. of
<taxonomicName id="4CC54D75B123FF80D2BD51D46793C483" ID-CoL="4TBXH" authority="(de Saussure))" authorityName="" baseAuthorityName="de Saussure" baseAuthorityYear="1854" box="[485,753,187,213]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="B9B1EAE4B123FF80D2BD51D46724C482" box="[485,582,187,212]" italics="true" pageId="12" pageNumber="463">gregaria</emphasis>
(de Saussure))
</taxonomicName>
; 2001a: 9 (syn. of
<taxonomicName id="4CC54D75B123FF80D09951D461DFC483" authority="(Smith))" authorityName="Smith" authorityYear="1860" baseAuthorityName="Smith" box="[961,1213,187,213]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B123FF80D09951D46131C482" box="[961,1107,187,212]" italics="true" pageId="12" pageNumber="463">R. impetuosa</emphasis>
(Smith))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF51B06003C517" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="ABD511D8B123FF80D3AF51B06003C517" pageId="12" pageNumber="463">
<treatmentCitation id="0A6410E7B123FF80D3AF51B067FBC4AF" author="Buysson, R. du" box="[247,665,223,249]" page="306" pageId="12" pageNumber="463" year="1909">
<taxonomicName id="4CC54D75B123FF80D3AF51B067FBC4AF" ID-CoL="3P8KL" authority="du Buysson, 1909: 306" authorityName="du Buysson" authorityPageNumber="306" authorityYear="1909" box="[247,665,223,249]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cohni">
<emphasis id="B9B1EAE4B123FF80D3AF51B064EAC4AF" box="[247,392,223,249]" italics="true" pageId="12" pageNumber="463">Icaria Cohni</emphasis>
<bibRefCitation id="EF544B07B123FF80D2C851B067FBC4AF" author="Buysson, R. du" box="[400,665,223,249]" firstAuthor="du Buysson" page="306" pageId="12" pageNumber="463" pagination="305 - 306" refId="ref27723" refString="Buysson, R. du (1909) Deux Hymenopteres nouveaux d'Oceanie. Bulletin de la Societe entomologique de France, 1909, 305 - 306. https: // doi. org / 10.3406 / bsef. 1909.24575" type="journal article" year="1909">du Buysson, 1909: 306</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, male - “
<collectingRegion id="4901F814B123FF80D1A751B066F6C4AF" box="[767,916,223,249]" country="Papua New Guinea" name="Bougainville" pageId="12" pageNumber="463">Bougainville</collectingRegion>
” (
<typeStatus id="547E8854B123FF80D0F651B06175C4AF" box="[942,1047,223,249]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
Paris). -
<bibRefCitation id="EF544B07B123FF80D7D851B060FEC4AF" author="Vecht, J. van der" box="[1152,1436,223,249]" firstAuthor="van der Vecht" page="104" pageId="12" pageNumber="463" pagination="103 - 190" refId="ref51729" refString="Vecht, J. van der (1941) The Indo-Australian species of the genus Ropalidia (= Icaria) (Hym., Vespidae) (first part). Treubia, 18, 103 - 190." type="journal article" year="1941">
van der
<treatmentCitation id="0A6410E7B123FF80D78451B060FEC4AF" author="Vecht, J. van der" box="[1244,1436,223,249]" page="104" pageId="12" pageNumber="463" year="1941">Vecht, 1941: 104</treatmentCitation>
</bibRefCitation>
(syn. of
<taxonomicName id="4CC54D75B123FF80D2E8506C67B5C54B" ID-CoL="4TBXH" authority="(de Saussure))" authorityName="" baseAuthorityName="de Saussure" baseAuthorityYear="1854" box="[432,727,259,285]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="B9B1EAE4B123FF80D2E8506C674DC54A" box="[432,559,259,284]" italics="true" pageId="12" pageNumber="463">R. gregaria</emphasis>
(de Saussure))
</taxonomicName>
. -
<treatmentCitation id="0A6410E7B123FF80D1B6506C66DDC54B" author="Richards, O. W." box="[750,959,259,285]" page="86" pageId="12" pageNumber="463" year="1978">
<bibRefCitation id="EF544B07B123FF80D1B6506C66DDC54B" author="Richards" box="[750,959,259,285]" firstAuthor="Richards" page="86" pageId="12" pageNumber="463" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 86</bibRefCitation>
</treatmentCitation>
(
<taxonomicName id="4CC54D75B123FF80D095506C6169C54B" ID-CoL="3P8L2" authorityName="Buysson" authorityYear="1909" box="[973,1035,259,285]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="kohni">
<emphasis id="B9B1EAE4B123FF80D095506C6169C54B" box="[973,1035,259,285]" italics="true" pageId="12" pageNumber="463">kohni</emphasis>
</taxonomicName>
[!]; syn. of
<taxonomicName id="4CC54D75B123FF80D7D2506C64BCC517" ID-CoL="5KXX3" authority="(Cameron))" baseAuthorityName="Cameron" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="spilocephala">
<emphasis id="B9B1EAE4B123FF80D7D2506C60FEC54B" box="[1162,1436,259,285]" italics="true" pageId="12" pageNumber="463">R. gregaria spilocephala</emphasis>
(Cameron))
</taxonomicName>
. -
<treatmentCitation id="0A6410E7B123FF80D2A0504867D9C517" author="Kojima, J." box="[504,699,294,321]" page="5" pageId="12" pageNumber="463" year="2001">
<bibRefCitation id="EF544B07B123FF80D2A0504867D9C517" author="Kojima" box="[504,699,294,321]" firstAuthor="Kojima" page="5" pageId="12" pageNumber="463" pagination="1 - 22" refId="ref40107" refString="Kojima, J. (2001 a) Taxonomic notes on types of the social wasp tribe Ropalidiini (Insecta: Hymenoptera; Vespidae, Polistinae) housed mainly in the Museum d'Histoire Naturelle, Geneve. Natural History Bulletin of Ibaraki University, 5, 1 - 22." type="journal article" year="2001" yearSuffix="a">Kojima, 2001a: 5</bibRefCitation>
</treatmentCitation>
(designation of
<typeStatus id="547E8854B123FF80D02A504866BFC517" box="[882,989,295,321]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
), 9 (syn. of
<taxonomicName id="4CC54D75B123FF80D73A5048603FC517" authority="(Smith))" authorityName="Smith" authorityYear="1860" baseAuthorityName="Smith" box="[1122,1373,295,321]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B123FF80D73A50486196C516" box="[1122,1268,295,320]" italics="true" pageId="12" pageNumber="463">R. impetuosa</emphasis>
(Smith))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF50246656C533" blockId="12.[247,1437,151,2013]" box="[247,820,331,357]" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF50246419C532" ID-CoL="6N4C8" authorityName="de Saussure" authorityYear="1853" box="[247,379,331,356]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B9B1EAE4B123FF80D3AF50246419C532" box="[247,379,331,356]" italics="true" pageId="12" pageNumber="463">Icaria picta</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF544B07B123FF80D2D150246731C533" author="Buysson, R. du" box="[393,595,331,357]" firstAuthor="du Buysson" pageId="12" pageNumber="463" pagination="227 - 230" refId="ref27793" refString="Buysson, R. du (1913 [&quot; 1911 &quot;]) Vespidae et Eumenidae. In: Friese, H., Hymenoptera der Aru-und Kei-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 34, pp. 227 - 230." type="journal article" year="1913">du Buysson, 1913</bibRefCitation>
[1911]: 229 (distr.).
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF50006752C70B" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF500064ACC5DE" ID-CoL="4TBXH" authorityName="" baseAuthorityName="de Saussure" baseAuthorityYear="1854" box="[247,462,367,393]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="B9B1EAE4B123FF80D3AF500064ACC5DE" box="[247,462,367,393]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF544B07B123FF80D285500067D8C5DF" author="Tillyard" box="[477,698,366,393]" firstAuthor="Tillyard" page="297" pageId="12" pageNumber="463" refId="ref51485" refString="Tillyard, R. J. (1925) The Insects of Australia and New Zealand. Angus &amp; Robertson, Ltd., Sydney, 560 pp., 44 pls." type="book" year="1925">Tillyard, 1925: 297</bibRefCitation>
. -
<bibRefCitation id="EF544B07B123FF80D18E500066CCC5DF" author="Williams" box="[726,942,367,393]" firstAuthor="Williams" page="79" pageId="12" pageNumber="463" pagination="53 - 118" refId="ref53003" refString="Williams, F. X. (1928) The natural history of a Philippine nipa house with descriptions of new wasps. The Philippine Journal of Science, 35, 53 - 118." type="journal article" year="1928">Williams, 1928: 79</bibRefCitation>
(prey:
<taxonomicName id="4CC54D75B123FF80D759500060F3C5DF" ID-CoL="4F64F" authority="Kirkaldy" authorityName="Kirkaldy" authorityYear="1903" box="[1025,1425,367,393]" class="Insecta" family="Delphacidae" genus="Perkinsiella" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="saccharicida">
<emphasis id="B9B1EAE4B123FF80D75950006043C5DF" box="[1025,1313,367,393]" italics="true" pageId="12" pageNumber="463">Perkinsiella saccharicida</emphasis>
Kirkaldy
</taxonomicName>
). -
<bibRefCitation id="EF544B07B123FF80D23150FC67EAC5FB" author="Vecht, J. van der" box="[361,648,403,429]" firstAuthor="van der Vecht" page="104" pageId="12" pageNumber="463" pagination="103 - 190" refId="ref51729" refString="Vecht, J. van der (1941) The Indo-Australian species of the genus Ropalidia (= Icaria) (Hym., Vespidae) (first part). Treubia, 18, 103 - 190." type="journal article" year="1941">van der Vecht, 1941: 104</bibRefCitation>
(syns.:
<taxonomicName id="4CC54D75B123FF80D1BC50FC669DC5FB" ID-CoL="3P8KL" authority="du Buysson" authorityName="du Buysson" box="[740,1023,403,429]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cohni">
<emphasis id="B9B1EAE4B123FF80D1BC50FC660DC5FB" box="[740,879,403,429]" italics="true" pageId="12" pageNumber="463">Icaria cohni</emphasis>
du Buysson
</taxonomicName>
,
<taxonomicName id="4CC54D75B123FF80D75550FC6185C5FB" authority="Smith" authorityName="Smith" authorityYear="1860" baseAuthorityName="Smith" box="[1037,1255,403,429]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B123FF80D75550FC61FBC5FA" box="[1037,1177,403,428]" italics="true" pageId="12" pageNumber="463">I. impetuosa</emphasis>
Smith
</taxonomicName>
,
<taxonomicName id="4CC54D75B123FF80D7AC50FC64A0C587" ID-CoL="3P8LM" authority="Cameron" authorityName="Cameron" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="spilocephala">
<emphasis id="B9B1EAE4B123FF80D7AC50FC60FEC5FB" box="[1268,1436,403,429]" italics="true" pageId="12" pageNumber="463">I. spilocephala</emphasis>
Cameron
</taxonomicName>
), 111 (key), 149 (distr. [including subspecies
<taxonomicName id="4CC54D75B123FF80D09750D8613CC587" ID-CoL="3P8LM" box="[975,1118,439,465]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="spilocephala">
<emphasis id="B9B1EAE4B123FF80D09750D8613CC587" box="[975,1118,439,465]" italics="true" pageId="12" pageNumber="463">spilocephala</emphasis>
</taxonomicName>
]). -
<bibRefCitation id="EF544B07B123FF80D7CC50D86013C587" author="Cheesman" box="[1172,1393,438,465]" firstAuthor="Cheesman" page="3" pageId="12" pageNumber="463" pagination="1 - 26" refId="ref29995" refString="Cheesman, L. E. (1952) Ropalidia of Papuasia. Annals and Magazine of Natural History, Series 12, 5, 1 - 26, https: // doi. org / 10.1080 / 00222935208654259" type="journal article" year="1952">Cheesman, 1952: 3</bibRefCitation>
, 20 (male). -
<bibRefCitation id="EF544B07B123FF80D29B50B467B4C5A3" author="Vecht, J. van der" box="[451,726,475,501]" firstAuthor="van der Vecht" page="27" pageId="12" pageNumber="463" pagination="1 - 72" refId="ref51768" refString="Vecht, J. van der (1962) The Indo-Australian species of the genus Ropalidia (Icaria) (Hymenoptera, Vespidae) (second part). Zoologische Verhandelingen, 57, 1 - 72." type="journal article" year="1962">van der Vecht, 1962: 27</bibRefCitation>
(distr.; nest). -
<bibRefCitation id="EF544B07B123FF80D0D150B4610AC5A3" author="Baltazar" box="[905,1128,475,501]" firstAuthor="Baltazar" page="291" pageId="12" pageNumber="463" pagination="1 - 488" refId="ref25375" refString="Baltazar, C. (1966) A catalogue of Philippine Hymenoptera. Pacific Insects Monograph, 8, 1 - 488." type="journal article" year="1966">Baltazar, 1966: 291</bibRefCitation>
(in subgenus
<taxonomicName id="4CC54D75B123FF80D65350B460F6C5A3" authority="White, 1841" box="[1291,1428,475,501]" class="Insecta" family="Vespidae" genus="Anthreneida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="B9B1EAE4B123FF80D65350B460F6C5A3" box="[1291,1428,475,501]" italics="true" pageId="12" pageNumber="463">Anthreneida</emphasis>
</taxonomicName>
; cat.). -
<bibRefCitation id="EF544B07B123FF80D2F050906704C64F" author="Iwata" box="[424,614,510,537]" firstAuthor="Iwata" page="295" pageId="12" pageNumber="463" refId="ref38023" refString="Iwata, K. (1976) Evolution of Instinct. Comparative Ethology of Hymenoptera. Amerind Publishing Company, New Delhi, 535 pp." type="book" year="1976">Iwata, 1976: 295</bibRefCitation>
(in subgenus
<taxonomicName id="4CC54D75B123FF80D05C509066EFC64F" authority="White, 1841" box="[772,909,511,537]" class="Insecta" family="Vespidae" genus="Anthreneida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="B9B1EAE4B123FF80D05C509066EFC64F" box="[772,909,511,537]" italics="true" pageId="12" pageNumber="463">Anthreneida</emphasis>
</taxonomicName>
; ethology). -
<bibRefCitation id="EF544B07B123FF80D7705090619DC64F" author="Richards" box="[1064,1279,510,537]" firstAuthor="Richards" page="50" pageId="12" pageNumber="463" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 50</bibRefCitation>
, 53, 61 (key), 65 (larva in key), 83, Fig. 26 (in subgenus
<taxonomicName id="4CC54D75B123FF80D014534C66C4C66B" authorityName="Dalla Torre" authorityYear="1904" box="[844,934,547,573]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D014534C66C4C66B" box="[844,934,547,573]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; redescription; nest; ethology). -
<bibRefCitation id="EF544B07B123FF80D670534C64EDC637" author="Carpenter" firstAuthor="Carpenter" pageId="12" pageNumber="463" pagination="11 - 38" refId="ref28609" refString="Carpenter, J. M. (1981 [&quot; 1982 &quot;] The phylogenetic relationships and natural classification of the Vespoidea (Hymenoptera). Systematic Entomology, 7, 11 - 38. https: // doi. org / 10.1111 / j. 1365 - 3113.1982. tb 00124. x" type="journal article" year="1981">Carpenter, 1981</bibRefCitation>
[1982]: 34 (phylogenetic relationships). -
<bibRefCitation id="EF544B07B123FF80D02A53286136C637" author="Kojima" box="[882,1108,583,609]" firstAuthor="Kojima" page="272" pageId="12" pageNumber="463" pagination="272 - 280" refId="ref39051" refString="Kojima, J. (1982 a) Nest architecture of three Ropalidia species (Hymenoptera: Vespidae) on Leyte Island, the Philippines. Biotropica, 14, 272 - 280. https: // doi. org / 10.2307 / 2388085" type="journal article" year="1982" yearSuffix="a">Kojima, 1982a: 272</bibRefCitation>
(in subgenus
<taxonomicName id="4CC54D75B123FF80D7AC5328602CC637" authority="Dalla Torre, 1904" box="[1268,1358,583,609]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D7AC5328602CC637" box="[1268,1358,583,609]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; nest); 1982b: 17 (ethology); 1984a: 52, Fig. 2 (larva); 1984b: 525 (key), 528, Figs 6, 34, 39, 44, 49 (in subgenus
<taxonomicName id="4CC54D75B123FF80D29153E06741C6FF" authority="Dalla Torre, 1904" box="[457,547,655,681]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D29153E06741C6FF" box="[457,547,655,681]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; male genitalia;
<collectingCountry id="F3D27666B123FF80D1B053E06609C6FF" box="[744,875,655,681]" name="Philippines" pageId="12" pageNumber="463">Philippines</collectingCountry>
). -
<bibRefCitation id="EF544B07B123FF80D0CB53E061F5C6FF" author="Spradbery" box="[915,1175,655,681]" firstAuthor="Spradbery" page="349" pageId="12" pageNumber="463" pagination="337 - 388" refId="ref50590" refString="Spradbery, J. P. (1991 b) Evolution of queen number and queen control. In: Ross, K. G. &amp; Matthews, R. W. (Eds.), The Social Biology of Wasps. Cornell University Press, Ithaca, New York, pp. 337 - 388." type="book chapter" year="1991" yearSuffix="b">Spradbery, 1991b: 349</bibRefCitation>
(colony population). -
<bibRefCitation id="EF544B07B123FF80D20F53DC674EC69B" author="Wenzel" box="[343,556,691,717]" firstAuthor="Wenzel" page="487" pageId="12" pageNumber="463" pagination="480 - 519" refId="ref52565" refString="Wenzel, J. W. (1991) Evolution of nest architecture. In: Ross, K. G. &amp; Matthews, R. W. (Eds.), The Social Biology of Wasps. Cornell University Press, Ithaca, New York, pp. 480 - 519. https: // doi. org / 10.7591 / 9781501718670 - 017" type="book chapter" year="1991">Wenzel, 1991: 487</bibRefCitation>
(nest). -
<bibRefCitation id="EF544B07B123FF80D1CE53DC6648C69B" author="Ito" box="[662,810,691,717]" firstAuthor="Ito" page="87" pageId="12" pageNumber="463" refId="ref37643" refString="Ito, Y. (1993) Behaviour and Social Evolution of Wasps. The Communal Aggregation Hypothesis. Oxford University Press, Oxford, 159 pp." type="book" year="1993">Itô, 1993: 87</bibRefCitation>
(in subgenus
<taxonomicName id="4CC54D75B123FF80D09353DC6147C69B" authority="Dalla Torre, 1904" box="[971,1061,691,717]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D09353DC6147C69B" box="[971,1061,691,717]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; ethology). -
<bibRefCitation id="EF544B07B123FF80D79A53DC6018C69B" author="Naumann" box="[1218,1402,691,717]" firstAuthor="Naumann" pageId="12" pageNumber="463" pagination="173 - 203" refId="ref44932" refString="Naumann, I. D., Edwards, E. D., Weir, T. A. &amp; Rentz, D. C. F. (1993) Insects of the Heathlands Area, Cape York Peninsula, Queensland. Cape York Peninsula Scientific Expedition Wet Season 1992 Report, 2, 173 - 203." type="journal article" year="1993">Naumann, 1993</bibRefCitation>
,
<emphasis id="B9B1EAE4B123FF80D6DE53DC60FEC69A" box="[1414,1436,691,716]" italics="true" pageId="12" pageNumber="463">in</emphasis>
Naumann
<emphasis id="B9B1EAE4B123FF80D29353B86761C6A7" box="[459,515,727,753]" italics="true" pageId="12" pageNumber="463">et al.</emphasis>
: 180 (distr.). - Macalintal &amp; Starr: tables 12 (in subgenus
<taxonomicName id="4CC54D75B123FF80D7C253B86196C6A7" authority="Dalla Torre, 1904" box="[1178,1268,727,753]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D7C253B86196C6A7" box="[1178,1268,727,753]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; morphology). -
<bibRefCitation id="EF544B07B123FF80D23F5394674EC743" author="Wenzel" box="[359,556,763,789]" firstAuthor="Wenzel" page="26" pageId="12" pageNumber="463" pagination="1 - 39" refId="ref52632" refString="Wenzel, J. W. (1998) A generic key to the nests of hornets, yellowjackets, and paper wasps worldwide (Vespidae: Vespinae, Polistinae). American Museum Novitates, 3224, 1 - 39." type="journal article" year="1998">Wenzel, 1998: 26</bibRefCitation>
(nest examined). -
<bibRefCitation id="EF544B07B123FF80D05E53946689C743" author="Kojima" box="[774,1003,763,789]" firstAuthor="Kojima" page="367" pageId="12" pageNumber="463" pagination="367 - 377" refId="ref39867" refString="Kojima, J. (1999 b) Taxonomic notes on Australian Ropalidia Guerin-Meneville, 1831 (Hymenoptera: Vespidae, Polistinae). Entomological Science, 2, 367 - 377." type="journal article" year="1999" yearSuffix="b">Kojima, 1999b: 367</bibRefCitation>
, 371 (syn.:
<taxonomicName id="4CC54D75B123FF80D736539460FAC743" ID-CoL="3P8LM" authority="(Cameron)" baseAuthorityName="Cameron" baseAuthorityYear="1906" box="[1134,1432,763,789]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="spilocephala">
<emphasis id="B9B1EAE4B123FF80D73653946076C743" box="[1134,1300,763,789]" italics="true" pageId="12" pageNumber="463">I. spilocephala</emphasis>
(Cameron)
</taxonomicName>
, erroneously marked “Syn. nov.”; W. A., N. T., Qld., N. S. W., Vic.); 1999c: 381 (key), 386 (distr.), Figs 17, 21, 25, 36.
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF520864A6C047" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF52086483C7D6" ID-CoL="4TBXP" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[247,481,871,897]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B123FF80D3AF52086483C7D6" box="[247,481,871,897]" italics="true" pageId="12" pageNumber="463">Ropalidia impetuosa</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF544B07B123FF80D2AA520867ACC7D7" author="Bequaert" box="[498,718,870,897]" firstAuthor="Bequaert" page="51" pageId="12" pageNumber="463" pagination="47 - 51" refId="ref26133" refString="Bequaert, J. (1932) Aculeata (Vespidae), genre Ropalidia. Resultats Scientifiques du Voyage aux Indes Orientales Neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique, 4 (5), 47 - 51." type="journal article" year="1932">Bequaert, 1932: 51</bibRefCitation>
. -
<bibRefCitation id="EF544B07B123FF80D1B5520866A7C7D7" author="Richards" box="[749,965,870,897]" firstAuthor="Richards" page="58" pageId="12" pageNumber="463" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 58</bibRefCitation>
(in subgenus
<taxonomicName id="4CC54D75B123FF80D73E520861A2C7D7" authorityName="Dalla Torre" authorityYear="1904" box="[1126,1216,871,897]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D73E520861A2C7D7" box="[1126,1216,871,897]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; list), 86 (probable syn.:
<taxonomicName id="4CC54D75B123FF80D2C052E466FCC7F3" ID-CoL="5KXX3" authority="(Cameron))" baseAuthorityName="Cameron" box="[408,926,907,933]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="spilocephala">
<emphasis id="B9B1EAE4B123FF80D2C052E4666EC7F3" box="[408,780,907,933]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria spilocephala</emphasis>
(Cameron))
</taxonomicName>
. -
<bibRefCitation id="EF544B07B123FF80D09852E461E9C7F3" author="Kojima" box="[960,1163,907,933]" firstAuthor="Kojima" page="9" pageId="12" pageNumber="463" pagination="1 - 22" refId="ref40107" refString="Kojima, J. (2001 a) Taxonomic notes on types of the social wasp tribe Ropalidiini (Insecta: Hymenoptera; Vespidae, Polistinae) housed mainly in the Museum d'Histoire Naturelle, Geneve. Natural History Bulletin of Ibaraki University, 5, 1 - 22." type="journal article" year="2001" yearSuffix="a">Kojima, 2001a: 9</bibRefCitation>
(syns.:
<taxonomicName id="4CC54D75B123FF80D7B352E460F7C7F3" ID-CoL="3P8LM" baseAuthorityName="Cameron" baseAuthorityYear="1906" box="[1259,1429,907,933]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="spilocephala">
<emphasis id="B9B1EAE4B123FF80D7B352E460F7C7F3" box="[1259,1429,907,933]" italics="true" pageId="12" pageNumber="463">I. spilocephala</emphasis>
</taxonomicName>
,
<taxonomicName id="4CC54D75B123FF80D20F52C064CDC79F" ID-CoL="3P8KL" box="[343,431,943,969]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cohni">
<emphasis id="B9B1EAE4B123FF80D20F52C064CDC79F" box="[343,431,943,969]" italics="true" pageId="12" pageNumber="463">I. cohni</emphasis>
</taxonomicName>
,
<taxonomicName id="4CC54D75B123FF80D29852C067CAC79E" ID-CoL="4TBXT" box="[448,680,943,968]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="inquieta">
<emphasis id="B9B1EAE4B123FF80D29852C067CAC79E" box="[448,680,943,968]" italics="true" pageId="12" pageNumber="463">R. gregaria inquieta</emphasis>
</taxonomicName>
,
<taxonomicName id="4CC54D75B123FF80D1E052C066B1C79F" ID-CoL="4TC3B" box="[696,979,943,969]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="variety" species="gregaria" variety="tolerans">
<emphasis id="B9B1EAE4B123FF80D1E052C06659C79E" box="[696,827,943,968]" italics="true" pageId="12" pageNumber="463">R. gregaria</emphasis>
var.
<emphasis id="B9B1EAE4B123FF80D02F52C066B1C79F" box="[887,979,943,969]" italics="true" pageId="12" pageNumber="463">tolerans</emphasis>
</taxonomicName>
); 2001b: 32, 56 (syn.; distr.). -
<bibRefCitation id="EF544B07B123FF80D61B52C06713C7BB" author="Saito &amp; Kojima" firstAuthor="Saito" page="273 - 274" pageId="12" pageNumber="463" pagination="179 - 188" refId="ref47473" refString="Saito, F. &amp; Kojima, J. (2005 a) Taxonomy and biogeography of Australian species of the Ropalidia stigma group and R. variegata group (Hymenoptera: Vespidae). Entomological Science, 8, 179 - 188. https: // doi. org / 10.1111 / j. 1479 - 8298.2005.00111. x" type="journal article" year="2005" yearSuffix="a">Saito &amp; Kojima, 2005a: 273274</bibRefCitation>
(not established south of latitude
<geoCoordinate id="EEF15031B123FF80D0AD52BC614DC7BB" box="[1013,1071,979,1005]" degrees="30" direction="south" orientation="latitude" pageId="12" pageNumber="463" precision="55555" value="-30.0">30°S</geoCoordinate>
). -
<bibRefCitation id="EF544B07B123FF80D70E52BC6020C7BB" author="Nugroho" box="[1110,1346,979,1005]" etAl="et al." firstAuthor="Nugroho" pageId="12" pageNumber="463" pagination="71 - 186" refId="ref45315" refString="Nugroho, H., Kojima, J. &amp; Carpenter, J. M. (2012 [&quot; 2011 &quot;]) Checklist of vespid species (Insecta: Hymenoptera: Vespidae) occurring in Indonesian Archipelago. Treubia, 38, 71 - 186." type="journal article" year="2012">
Nugroho
<emphasis id="B9B1EAE4B123FF80D79A52BC6199C7BB" box="[1218,1275,979,1005]" italics="true" pageId="12" pageNumber="463">et al.</emphasis>
, 2012
</bibRefCitation>
[2011]: 152 (list).
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF557467F4C063" blockId="12.[247,1437,151,2013]" box="[247,662,1050,1077]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="ABD511D8B123FF80D3AF557467F4C063" box="[247,662,1050,1077]" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF557464C9C063" ID-CoL="3P8KL" authorityName="Sonan" authorityYear="1938" box="[247,427,1051,1077]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cohni">
<emphasis id="B9B1EAE4B123FF80D3AF557464C9C063" box="[247,427,1051,1077]" italics="true" pageId="12" pageNumber="463">Ropalidia cohni</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A6410E7B123FF80D2E2557467F0C063" author="Bequaert, J." box="[442,658,1050,1077]" page="51" pageId="12" pageNumber="463" year="1932">
<bibRefCitation id="EF544B07B123FF80D2E2557467F0C063" author="Bequaert" box="[442,658,1050,1077]" firstAuthor="Bequaert" page="51" pageId="12" pageNumber="463" pagination="47 - 51" refId="ref26133" refString="Bequaert, J. (1932) Aculeata (Vespidae), genre Ropalidia. Resultats Scientifiques du Voyage aux Indes Orientales Neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique, 4 (5), 47 - 51." type="journal article" year="1932">Bequaert, 1932: 51</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF555067D2C02B" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="ABD511D8B123FF80D3AF555067D2C02B" pageId="12" pageNumber="463">
?
<taxonomicName id="4CC54D75B123FF80D25255506769C00F" ID-CoL="78WC8" baseAuthorityName="Fabricius" baseAuthorityYear="1804" box="[266,523,1087,1113]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cyathiformis">
<emphasis id="B9B1EAE4B123FF80D25255506769C00F" box="[266,523,1087,1113]" italics="true" pageId="12" pageNumber="463">Ropalidia cyathiformis</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A6410E7B123FF80D1405550678FC00F" author="Bequaert, J." box="[536,749,1086,1113]" page="51" pageId="12" pageNumber="463" year="1932">
<bibRefCitation id="EF544B07B123FF80D1405550678FC00F" author="Bequaert" box="[536,749,1086,1113]" firstAuthor="Bequaert" page="51" pageId="12" pageNumber="463" pagination="47 - 51" refId="ref26133" refString="Bequaert, J. (1932) Aculeata (Vespidae), genre Ropalidia. Resultats Scientifiques du Voyage aux Indes Orientales Neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique, 4 (5), 47 - 51." type="journal article" year="1932">Bequaert, 1932: 51</bibRefCitation>
</treatmentCitation>
(recorded from New
<collectingCountry id="F3D27666B123FF80D08455506152C00F" box="[988,1072,1087,1113]" name="Guinea-Bissau" pageId="12" pageNumber="463">Guinea</collectingCountry>
; syns.:
<taxonomicName id="4CC54D75B123FF80D7DA5550605BC00F" ID-CoL="4TC3M" authority="Smith" authorityName="Smith" box="[1154,1337,1087,1113]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B9B1EAE4B123FF80D7DA5550618FC00E" box="[1154,1261,1087,1112]" italics="true" pageId="12" pageNumber="463">variegata</emphasis>
Smith
</taxonomicName>
,
<taxonomicName id="4CC54D75B123FF80D61C555064DEC02B" ID-CoL="6N4C8" authority="de Saussure" authorityName="de Saussure" authorityYear="1853" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B9B1EAE4B123FF80D61C5550601EC00E" box="[1348,1404,1087,1112]" italics="true" pageId="12" pageNumber="463">picta</emphasis>
de Saussure
</taxonomicName>
) [misidentification?].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF55E86785C0F7" blockId="12.[247,1437,151,2013]" box="[247,743,1158,1185]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="ABD511D8B123FF80D3AF55E86785C0F7" box="[247,743,1158,1185]" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF55E8649EC0F7" ID-CoL="3P8LM" box="[247,508,1159,1185]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="spilocephala">
<emphasis id="B9B1EAE4B123FF80D3AF55E8649EC0F7" box="[247,508,1159,1185]" italics="true" pageId="12" pageNumber="463">Ropalidia spilocephala</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A6410E7B123FF80D15255E86781C0F7" author="Bequaert, J." box="[522,739,1158,1185]" page="51" pageId="12" pageNumber="463" year="1932">
<bibRefCitation id="EF544B07B123FF80D15255E86781C0F7" author="Bequaert" box="[522,739,1158,1185]" firstAuthor="Bequaert" page="51" pageId="12" pageNumber="463" pagination="47 - 51" refId="ref26133" refString="Bequaert, J. (1932) Aculeata (Vespidae), genre Ropalidia. Resultats Scientifiques du Voyage aux Indes Orientales Neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique, 4 (5), 47 - 51." type="journal article" year="1932">Bequaert, 1932: 51</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF55C46717C167" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="ABD511D8B123FF80D3AF55C46717C167" pageId="12" pageNumber="463">
<treatmentCitation id="0A6410E7B123FF80D3AF55C46670C093" author="Cheesman, L. E." box="[247,786,1194,1221]" page="3" pageId="12" pageNumber="463" year="1952">
<taxonomicName id="4CC54D75B123FF80D3AF55C46670C093" ID-CoL="4TBXT" authority="Cheesman, 1952: 3" authorityName="Cheesman" authorityPageNumber="3" authorityYear="1952" box="[247,786,1194,1221]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="inquieta">
<emphasis id="B9B1EAE4B123FF80D3AF55C46753C092" box="[247,561,1195,1221]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria inquieta</emphasis>
<bibRefCitation id="EF544B07B123FF80D16155C46670C093" author="Cheesman" box="[569,786,1194,1221]" firstAuthor="Cheesman" page="3" pageId="12" pageNumber="463" pagination="1 - 26" refId="ref29995" refString="Cheesman, L. E. (1952) Ropalidia of Papuasia. Annals and Magazine of Natural History, Series 12, 5, 1 - 26, https: // doi. org / 10.1080 / 00222935208654259" type="journal article" year="1952">Cheesman, 1952: 3</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 20, female, male - “Papua,...Mafulu,
<quantity id="4C3D9B13B123FF80D79955C46078C093" box="[1217,1306,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="12" pageNumber="463" unit="ft" value="4000.0">4000 ft.</quantity>
” (
<typeStatus id="547E8854B123FF80D66C55C460FFC093" box="[1332,1437,1195,1221]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
male London); also from three other localities in New
<collectingCountry id="F3D27666B123FF80D09A55A06175C0BF" box="[962,1047,1231,1257]" name="Guinea-Bissau" pageId="12" pageNumber="463">Guinea</collectingCountry>
. -
<treatmentCitation id="0A6410E7B123FF80D76C55A06068C0BF" author="Richards, O. W." box="[1076,1290,1231,1257]" page="86" pageId="12" pageNumber="463" year="1978">
<bibRefCitation id="EF544B07B123FF80D76C55A06068C0BF" author="Richards" box="[1076,1290,1231,1257]" firstAuthor="Richards" page="86" pageId="12" pageNumber="463" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 86</bibRefCitation>
</treatmentCitation>
(designation of
<typeStatus id="547E8854B123FF80D22D559C64BEC15B" box="[373,476,1267,1293]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
[error in locality: Mafalu]; syn. of
<taxonomicName id="4CC54D75B123FF80D039559C6114C15B" ID-CoL="5KXX3" box="[865,1142,1267,1293]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="spilocephala">
<emphasis id="B9B1EAE4B123FF80D039559C6114C15B" box="[865,1142,1267,1293]" italics="true" pageId="12" pageNumber="463">R. gregaria spilocephala</emphasis>
</taxonomicName>
). -
<treatmentCitation id="0A6410E7B123FF80D7C5559C603DC15B" author="Kojima, J." box="[1181,1375,1267,1293]" page="9" pageId="12" pageNumber="463" year="2001">
<bibRefCitation id="EF544B07B123FF80D7C5559C603DC15B" author="Kojima" box="[1181,1375,1267,1293]" firstAuthor="Kojima" page="9" pageId="12" pageNumber="463" pagination="1 - 22" refId="ref40107" refString="Kojima, J. (2001 a) Taxonomic notes on types of the social wasp tribe Ropalidiini (Insecta: Hymenoptera; Vespidae, Polistinae) housed mainly in the Museum d'Histoire Naturelle, Geneve. Natural History Bulletin of Ibaraki University, 5, 1 - 22." type="journal article" year="2001" yearSuffix="a">Kojima, 2001a: 9</bibRefCitation>
</treatmentCitation>
(syn. of
<taxonomicName id="4CC54D75B123FF80D22E54786713C167" ID-CoL="4TBXP" authority="(Smith))" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[374,625,1303,1329]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B123FF80D22E5478676AC166" box="[374,520,1303,1328]" italics="true" pageId="12" pageNumber="463">R. impetuosa</emphasis>
(Smith))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF54546156C1CB" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="ABD511D8B123FF80D3AF54546156C1CB" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF5454664BC103" ID-CoL="4TC3B" authority="Cheesman, 1952" authorityName="Cheesman" authorityYear="1952" box="[247,809,1338,1365]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="variety" species="gregaria" variety="tolerans">
<emphasis id="B9B1EAE4B123FF80D3AF545464ACC102" box="[247,462,1339,1365]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria</emphasis>
var.
<emphasis id="B9B1EAE4B123FF80D15E54546700C103" box="[518,610,1339,1365]" italics="true" pageId="12" pageNumber="463">tolerans</emphasis>
<bibRefCitation id="EF544B07B123FF80D1315454664BC103" author="Cheesman" box="[617,809,1338,1365]" firstAuthor="Cheesman" pageId="12" pageNumber="463" pagination="1 - 26" refId="ref29995" refString="Cheesman, L. E. (1952) Ropalidia of Papuasia. Annals and Magazine of Natural History, Series 12, 5, 1 - 26, https: // doi. org / 10.1080 / 00222935208654259" type="journal article" year="1952">Cheesman, 1952</bibRefCitation>
</taxonomicName>
): 3 (as subspecies), 21 (as var.), female - “Cyclops Mts. Sabron” (
<typeStatus id="547E8854B123FF80D29B5430674EC12F" box="[451,556,1375,1401]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
London). -
<treatmentCitation id="0A6410E7B123FF80D1EF543066EDC12F" author="Richards, O. W." box="[695,911,1375,1401]" page="86" pageId="12" pageNumber="463" year="1978">
<bibRefCitation id="EF544B07B123FF80D1EF543066EDC12F" author="Richards" box="[695,911,1375,1401]" firstAuthor="Richards" page="86" pageId="12" pageNumber="463" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 86</bibRefCitation>
</treatmentCitation>
(designation of
<typeStatus id="547E8854B123FF80D714543061D4C12F" box="[1100,1206,1375,1401]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
; syn. of
<taxonomicName id="4CC54D75B123FF80D64254306484C1CB" ID-CoL="5KXX3" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="spilocephala">
<emphasis id="B9B1EAE4B123FF80D64254306484C1CB" italics="true" pageId="12" pageNumber="463">R. gregaria spilocephala</emphasis>
</taxonomicName>
). -
<treatmentCitation id="0A6410E7B123FF80D15654EC67B3C1CB" author="Kojima, J." box="[526,721,1411,1437]" page="9" pageId="12" pageNumber="463" year="2001">
<bibRefCitation id="EF544B07B123FF80D15654EC67B3C1CB" author="Kojima" box="[526,721,1411,1437]" firstAuthor="Kojima" page="9" pageId="12" pageNumber="463" pagination="1 - 22" refId="ref40107" refString="Kojima, J. (2001 a) Taxonomic notes on types of the social wasp tribe Ropalidiini (Insecta: Hymenoptera; Vespidae, Polistinae) housed mainly in the Museum d'Histoire Naturelle, Geneve. Natural History Bulletin of Ibaraki University, 5, 1 - 22." type="journal article" year="2001" yearSuffix="a">Kojima, 2001a: 9</bibRefCitation>
</treatmentCitation>
(syn. of
<taxonomicName id="4CC54D75B123FF80D06D54EC6152C1CB" ID-CoL="4TBXP" authority="(Smith))" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[821,1072,1411,1437]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B123FF80D06D54EC66A5C1CA" box="[821,967,1411,1436]" italics="true" pageId="12" pageNumber="463">R. impetuosa</emphasis>
(Smith))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF54C86701C2CF" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="ABD511D8B123FF80D3AF54C86701C2CF" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF54C86752C196" ID-CoL="4TBXH" box="[247,560,1447,1473]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="gregaria">
<emphasis id="B9B1EAE4B123FF80D3AF54C86752C196" box="[247,560,1447,1473]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria gregaria</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A6410E7B123FF80D16454C86669C197" author="Richards, O. W." box="[572,779,1446,1473]" page="53" pageId="12" pageNumber="463" year="1978">
<bibRefCitation id="EF544B07B123FF80D16454C86669C197" author="Richards" box="[572,779,1446,1473]" firstAuthor="Richards" page="53" pageId="12" pageNumber="463" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 53</bibRefCitation>
</treatmentCitation>
(nest), 62 (key), 86 (in subgenus
<taxonomicName id="4CC54D75B123FF80D72954C861A9C197" authority="Dalla Torre, 1904" box="[1137,1227,1447,1473]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D72954C861A9C197" box="[1137,1227,1447,1473]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; “two [subspecies] can be recognized in the Australia-New
<collectingCountry id="F3D27666B123FF80D04054A4660BC1B3" box="[792,873,1483,1509]" name="Guinea-Bissau" pageId="12" pageNumber="463">Guinea</collectingCountry>
region. The forms of Malaya will require separate study.”). -
<treatmentCitation id="0A6410E7B123FF80D296548067C6C25F" author="Cardale, J. C." box="[462,676,1518,1545]" page="214" pageId="12" pageNumber="463" year="1985">
<bibRefCitation id="EF544B07B123FF80D296548067C6C25F" author="Cardale" box="[462,676,1518,1545]" firstAuthor="Cardale" page="214" pageId="12" pageNumber="463" pagination="150 - 303" refId="ref28558" refString="Cardale, J. C. (1985) Vespoidea and Sphecoidea. Zoological Catalogue of Australia, 2, 150 - 303." type="journal article" year="1985">Cardale, 1985: 214</bibRefCitation>
</treatmentCitation>
(in subgenus
<taxonomicName id="4CC54D75B123FF80D019548066F9C25F" authority="Dalla Torre, 1904" box="[833,923,1519,1545]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D019548066F9C25F" box="[833,923,1519,1545]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; cat.). -
<treatmentCitation id="0A6410E7B123FF80D0A2548061C5C25F" author="Ito, Y." box="[1018,1191,1519,1545]" page="313" pageId="12" pageNumber="463" year="1986">
<bibRefCitation id="EF544B07B123FF80D0A2548061C5C25F" author="Ito" box="[1018,1191,1519,1545]" firstAuthor="Ito" page="313" pageId="12" pageNumber="463" pagination="309 - 314" refId="ref37464" refString="Ito, Y. (1986 b) Observations on the social behaviour of three polistine wasps (Hymenoptera: Vespidae). Journal of the Australian Entomological Society, 25, 309 - 314. https: // doi. org / 10.1111 / j. 1440 - 6055.1986. tb 01121. x" type="journal article" year="1986" yearSuffix="b">Itô, 1986b: 313</bibRefCitation>
</treatmentCitation>
(ethology). -
<treatmentCitation id="0A6410E7B123FF80D619548064BBC27B" author="Kojima, J." page="145" pageId="12" pageNumber="463" year="1987">
<bibRefCitation id="EF544B07B123FF80D619548064BBC27B" author="Kojima" firstAuthor="Kojima" page="145" pageId="12" pageNumber="463" pagination="141 - 148" refId="ref39198" refString="Kojima, J. (1987 a) Descriptions of mature larvae of eight Australian Polistinae (Hymenoptera: Vespidae). Journal of the Australian Entomological Society, 26, 141 - 148. https: // doi. org / 10.1111 / j. 1440 - 6055.1987. tb 00274. x" type="journal article" year="1987" yearSuffix="a">Kojima, 1987a: 145</bibRefCitation>
</treatmentCitation>
, Figs 5258 (larva). -
<treatmentCitation id="0A6410E7B123FF80D1B9577C6687C27B" author="Spradbery, J. P." box="[737,997,1554,1581]" page="351" pageId="12" pageNumber="463" year="1991">
<bibRefCitation id="EF544B07B123FF80D1B9577C6687C27B" author="Spradbery" box="[737,997,1554,1581]" firstAuthor="Spradbery" page="351" pageId="12" pageNumber="463" pagination="337 - 388" refId="ref50590" refString="Spradbery, J. P. (1991 b) Evolution of queen number and queen control. In: Ross, K. G. &amp; Matthews, R. W. (Eds.), The Social Biology of Wasps. Cornell University Press, Ithaca, New York, pp. 337 - 388." type="book chapter" year="1991" yearSuffix="b">Spradbery, 1991b: 351</bibRefCitation>
</treatmentCitation>
(colony population). -
<treatmentCitation id="0A6410E7B123FF80D7AB577C6484C207" author="Ito, Y. &amp; Yamane, S." page="63" pageId="12" pageNumber="463" year="1992">
<bibRefCitation id="EF544B07B123FF80D7AB577C6484C207" author="Ito &amp; Yamane" firstAuthor="Ito" page="63 - 74" pageId="12" pageNumber="463" pagination="63 - 74" refId="ref37826" refString="Ito, Y. &amp; Yamane, S. (1992) Social behaviour of two primitively eusocial wasps, Ropalidia sp. nr. variegata and R. gregaria gregaria (Hymenoptera: Vespidae) in the Northern Territory, Australia, with special reference to task specialization and mating inhibition. Journal of Ethology, 10, 63 - 74. https: // doi. org / 10.1007 / BF 02350188" type="journal article" year="1992">Itô &amp; Yamane, 1992: 6374</bibRefCitation>
</treatmentCitation>
(ethology). -
<treatmentCitation id="0A6410E7B123FF80D1DF5758667CC207" author="Ito, Y." box="[647,798,1591,1617]" page="27" pageId="12" pageNumber="463" year="1993">
<bibRefCitation id="EF544B07B123FF80D1DF5758667CC207" author="Ito" box="[647,798,1591,1617]" firstAuthor="Ito" page="27" pageId="12" pageNumber="463" refId="ref37643" refString="Ito, Y. (1993) Behaviour and Social Evolution of Wasps. The Communal Aggregation Hypothesis. Oxford University Press, Oxford, 159 pp." type="book" year="1993">Itô, 1993: 27</bibRefCitation>
</treatmentCitation>
, 46, 66, 68 (in subgenus
<taxonomicName id="4CC54D75B123FF80D71C575861FCC207" authority="Dalla Torre, 1904" box="[1092,1182,1591,1617]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D71C575861FCC207" box="[1092,1182,1591,1617]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; ethology). -
<bibRefCitation id="EF544B07B123FF80D666575864EDC223" author="Borsato" firstAuthor="Borsato" pageId="12" pageNumber="463" pagination="245 - 251" refId="ref26912" refString="Borsato, W. (1994 [&quot; 1993 &quot;]) Contributo alla conoscenza degli Eumenidi e dei Vespidi australiani (Hymenoptera Eumenidae et Vespidae). Bollettino della Societa Entomologica Italiana, 125 (3), 245 - 251." type="journal article" year="1994">Borsato, 1994</bibRefCitation>
[1993],: 250 (in subgenus
<taxonomicName id="4CC54D75B123FF80D1E65734667AC223" authority="Dalla Torre, 1904" box="[702,792,1627,1653]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D1E65734667AC223" box="[702,792,1627,1653]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; N. T.). -
<treatmentCitation id="0A6410E7B123FF80D0DE5734618BC223" author="Kojima, J. &amp; Carpenter, J. M." box="[902,1257,1627,1653]" page="27" pageId="12" pageNumber="463" year="1997">
<bibRefCitation id="EF544B07B123FF80D0DE5734618BC223" author="Kojima &amp; Carpenter" box="[902,1257,1627,1653]" firstAuthor="Kojima" page="27" pageId="12" pageNumber="463" pagination="1 - 96" refId="ref40310" refString="Kojima, J. &amp; Carpenter, J. M. (1997 a) Catalog of species in the polistine tribe Ropalidiini (Hymenoptera: Vespidae). American Museum Novitates, 3199, 1 - 96." type="journal article" year="1997" yearSuffix="a">Kojima &amp; Carpenter, 1997a: 27</bibRefCitation>
</treatmentCitation>
(cat.). -
<treatmentCitation id="0A6410E7B123FF80D611573464D8C2CF" author="Brown, G. R." page="30" pageId="12" pageNumber="463" year="2009">
<bibRefCitation id="EF544B07B123FF80D611573464D8C2CF" author="Brown" firstAuthor="Brown" page="30" pageId="12" pageNumber="463" refId="ref27501" refString="Brown, G. R. (2009) Northern Territory Insects: A Comprehensive Guide. University of Queensland, Centre for Biological Information Technology, Brisbane. [CD-Rom]" type="book" year="2009">Brown, 2009: 30</bibRefCitation>
</treatmentCitation>
(North Coast).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B123FF80D3AF57CC67C5C3C3" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
<taxonomicName id="4CC54D75B123FF80D3AF57CC670BC2EB" ID-CoL="5KXX3" box="[247,617,1699,1725]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="spilocephala">
<emphasis id="B9B1EAE4B123FF80D3AF57CC670BC2EB" box="[247,617,1699,1725]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria spilocephala</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF544B07B123FF80D12357CC663AC2EB" author="Richards" box="[635,856,1698,1725]" firstAuthor="Richards" page="53" pageId="12" pageNumber="463" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 53</bibRefCitation>
, 62 (key), 86 (syns.:
<taxonomicName id="4CC54D75B123FF80D70A57CC61BDC2EB" ID-CoL="3P8L2" authorityName="Buysson" authorityYear="1909" box="[1106,1247,1699,1725]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="kohni">
<emphasis id="B9B1EAE4B123FF80D70A57CC61BDC2EB" box="[1106,1247,1699,1725]" italics="true" pageId="12" pageNumber="463">Icaria kohni</emphasis>
</taxonomicName>
[!] du Buysson,
<taxonomicName id="4CC54D75B123FF80D20F57A8667CC2B7" ID-CoL="4TBXT" authority="Cheesman" authorityName="Cheesman" box="[343,798,1735,1761]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subSpecies" species="gregaria" subSpecies="inquieta">
<emphasis id="B9B1EAE4B123FF80D20F57A867F8C2B6" box="[343,666,1735,1761]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria inquieta</emphasis>
Cheesman
</taxonomicName>
,
<taxonomicName id="4CC54D75B123FF80D07557A8604BC2B7" ID-CoL="4TC3B" authority="Cheesman" authorityName="Cheesman" box="[813,1321,1735,1761]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="variety" species="gregaria" variety="tolerans">
<emphasis id="B9B1EAE4B123FF80D07557A8616AC2B6" box="[813,1032,1735,1761]" italics="true" pageId="12" pageNumber="463">Ropalidia gregaria</emphasis>
var.
<emphasis id="B9B1EAE4B123FF80D71157A861C7C2B7" box="[1097,1189,1735,1761]" italics="true" pageId="12" pageNumber="463">tolerans</emphasis>
Cheesman
</taxonomicName>
; probably syn. of
<taxonomicName id="4CC54D75B123FF80D2E8578467A3C353" authority="Smith" authorityName="Smith" authorityYear="1860" baseAuthorityName="Smith" box="[432,705,1771,1797]" class="Insecta" family="Vespidae" genus="Icaria" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="impetuosa">
<emphasis id="B9B1EAE4B123FF80D2E857846713C352" box="[432,625,1771,1796]" italics="true" pageId="12" pageNumber="463">Icaria impetuosa</emphasis>
Smith
</taxonomicName>
; “I examined the
<typeStatus id="547E8854B123FF80D0CA57846696C353" box="[914,1012,1771,1797]" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
male [!] in the Zoological Museum, Amsterdam.”; redescription; nest). -
<bibRefCitation id="EF544B07B123FF80D1AE566066ACC37F" author="Cardale" box="[758,974,1806,1833]" firstAuthor="Cardale" page="215" pageId="12" pageNumber="463" pagination="150 - 303" refId="ref28558" refString="Cardale, J. C. (1985) Vespoidea and Sphecoidea. Zoological Catalogue of Australia, 2, 150 - 303." type="journal article" year="1985">Cardale, 1985: 215</bibRefCitation>
(in subgenus
<taxonomicName id="4CC54D75B123FF80D736566061AAC37F" authorityName="Dalla Torre" authorityYear="1904" box="[1134,1224,1807,1833]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D736566061AAC37F" box="[1134,1224,1807,1833]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; cat.). -
<bibRefCitation id="EF544B07B123FF80D6715660672FC31B" author="Spradbery &amp; Kojima" firstAuthor="Spradbery" page="636" pageId="12" pageNumber="463" pagination="632 - 653" refId="ref50704" refString="Spradbery, J. P. &amp; Kojima, J. (1989) Nest descriptions and colony populations of eleven species of Ropalidia (Hymenoptera, Vespidae) in New Guinea. Japanese Journal of Entomology, 57 (3), 632 - 653." type="journal article" year="1989">Spradbery &amp; Kojima, 1989: 636</bibRefCitation>
(nest; colony population). -
<bibRefCitation id="EF544B07B123FF80D0CF565C61F8C31B" author="Spradbery" box="[919,1178,1842,1869]" firstAuthor="Spradbery" page="351" pageId="12" pageNumber="463" pagination="337 - 388" refId="ref50590" refString="Spradbery, J. P. (1991 b) Evolution of queen number and queen control. In: Ross, K. G. &amp; Matthews, R. W. (Eds.), The Social Biology of Wasps. Cornell University Press, Ithaca, New York, pp. 337 - 388." type="book chapter" year="1991" yearSuffix="b">Spradbery, 1991b: 351</bibRefCitation>
(colony population). -
<bibRefCitation id="EF544B07B123FF80D20F56386496C327" author="Borsato" box="[343,500,1879,1905]" firstAuthor="Borsato" pageId="12" pageNumber="463" pagination="245 - 251" refId="ref26912" refString="Borsato, W. (1994 [&quot; 1993 &quot;]) Contributo alla conoscenza degli Eumenidi e dei Vespidi australiani (Hymenoptera Eumenidae et Vespidae). Bollettino della Societa Entomologica Italiana, 125 (3), 245 - 251." type="journal article" year="1994">Borsato, 1994</bibRefCitation>
[1993]: 250 (in subgenus
<taxonomicName id="4CC54D75B123FF80D04F56386613C327" authority="Dalla Torre, 1904" box="[791,881,1879,1905]" class="Insecta" family="Vespidae" genus="Ropalidia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="subGenus" subGenus="Icariola">
<emphasis id="B9B1EAE4B123FF80D04F56386613C327" box="[791,881,1879,1905]" italics="true" pageId="12" pageNumber="463">Icariola</emphasis>
</taxonomicName>
; Qld.). -
<bibRefCitation id="EF544B07B123FF80D08E563861D2C327" author="Ito" box="[982,1200,1879,1905]" etAl="et al." firstAuthor="Ito" page="231" pageId="12" pageNumber="463" pagination="231 - 233" refId="ref37769" refString="Ito, Y., Katada, S., Tsuchida, K. &amp; Rowe, R. J. (1996) Social structure of the primitively eusocial paper wasp Ropalidia gregaria spilocephala (Cameron) (Hymenoptera: Vespidae) in Queensland. Australian Journal of Entomology, 35, 231 - 233." type="journal article" year="1996">
Itô
<emphasis id="B9B1EAE4B123FF80D0A45638614EC327" box="[1020,1068,1879,1905]" italics="true" pageId="12" pageNumber="463">et al</emphasis>
., 1996: 231
</bibRefCitation>
(ethology). -
<bibRefCitation id="EF544B07B123FF80D6105638673CC3C3" author="Kojima &amp; Carpenter" firstAuthor="Kojima" page="27" pageId="12" pageNumber="463" pagination="1 - 96" refId="ref40310" refString="Kojima, J. &amp; Carpenter, J. M. (1997 a) Catalog of species in the polistine tribe Ropalidiini (Hymenoptera: Vespidae). American Museum Novitates, 3199, 1 - 96." type="journal article" year="1997" yearSuffix="a">Kojima &amp; Carpenter, 1997a: 27</bibRefCitation>
(cat.).
</paragraph>
</subSubSection>
<subSubSection id="C3DF657DB123FF80D3AF56F0614EC38B" pageId="12" pageNumber="463" type="distribution">
<paragraph id="8B7A36F6B123FF80D3AF56F0614EC38B" blockId="12.[247,1437,151,2013]" pageId="12" pageNumber="463">
Distribution: NE coastal, Murray-Darling basin; N. T., Qld., N. S. W.; also New
<collectingCountry id="F3D27666B123FF80D7FB56F06195C3EF" box="[1187,1271,1951,1977]" name="Guinea-Bissau" pageId="12" pageNumber="463">Guinea</collectingCountry>
; New Ireland; Solomon Is.;
<collectingCountry id="F3D27666B123FF80D2B556AC677AC38B" box="[493,536,1987,2013]" name="Fiji" pageId="12" pageNumber="463">Fiji</collectingCountry>
;
<collectingRegion id="4901F814B123FF80D17B56AC67EEC38B" box="[547,652,1987,2013]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="463">Sulawesi</collectingRegion>
; Ceram; Bacan;
<collectingCountry id="F3D27666B123FF80D01B56AC66F9C38B" box="[835,923,1987,2013]" name="Indonesia" pageId="12" pageNumber="463">Ambon</collectingCountry>
;
<collectingCountry id="F3D27666B123FF80D0FE56AC614AC38B" box="[934,1064,1987,2013]" name="Philippines" pageId="12" pageNumber="463">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>